Asymptotic behaviour of the non-autonomous competing two-species Lotka–Volterra models with impulsive effect

In this paper, the nonautonomous competing two-species Lotka–Volterra models with impulsive effect are considered, where all the parameters are time-dependent and asymptotically approach the corresponding periodic functions. Under some conditions, it is shown that the semi-trivial positive solutions of the models asymptotically approach the semi-trivial positive periodic solutions of the corresponding periodic system. It is also shown that the positive solution of the models asymptotically approach the positive periodic solution of the corresponding periodic system.     

非自治阶段结构合作系统的持久性与周期解

本文研究一类非自治阶段结构的合作系统,得到系统的最终有界性,对应周期系统正周期解的存在性,唯一性以及全局渐近稳定性的充分条件。     

具有扩散和放养的时滞竞争系统的正周期解

主要研究缀块环境下具有扩散和放养的时滞Lotka-Volterra竞争系统,得到了系统的周期解存在性,唯一性和全局渐近稳定性的充分条件．     

关于一类非自治阶段结构捕食系统的持久性与周期解

本文研究了一类非自治阶段结构捕食系统的渐近性质,得到在适当的条件下系统的持久性,对应周期系统正周期解的存在性、唯一性以及全局渐近稳定性。     

Pest management through continuous and impulsive control strategies

In this paper, we propose two mathematical models concerning continuous and, respectively, impulsive pest control strategies. In the case in which a continuous control is used, it is shown that the model admits a globally asymptotically stable positive equilibrium under appropriate conditions which involve parameter estimations. As a result, the global asymptotic stability of the unique positive equilibrium is used to establish a procedure to maintain the pests at an acceptably low level in the long term. In the case in which an impulsive control is used, it is observed that there exists a globally asymptotically stable susceptible pest-eradication periodic solution on condition that the amount of infective pests released periodically is larger than some critical value. When the amount of infective pests released is less than this critical value, the system is shown to be permanent, which implies that the trivial susceptible pest-eradication solution loses its stability. Further, the existence of a nontrivial periodic solution is also studied by means of numerical simulation. Finally, the efficiency of continuous and impulsive control policies is compared.     

Convergence in almost periodic Fisher and Kolmogorov models

 We study convergence of positive solutions for almost periodic reaction diffusion equations of Fisher or Kolmogorov type. It is proved that under suitable conditions every positive solution is asymptotically almost periodic. Moreover, all positive almost periodic solutions are harmonic and uniformly stable, and if one of them is spatially homogeneous, then so are others. The existence of an almost periodic global attractor is also discussed. Received: 11 November 1996 / Revised version: 8 January 1998     

Existence and global attractivity of positive periodic solutions of periodic n-species Lotka-Volterra competition systems with several deviating arguments.

In this paper, we study the existence and global attractivity of positive periodic solutions of periodic n-species Lotka-Volterra competition systems. By using the method of coincidence degree and Lyapunov functional, a set of easily verifiable sufficient conditions are derived for the existence of at least one strictly positive (componentwise) periodic solution of periodic n-species Lotka-Volterra competition systems with several deviating arguments and the existence of a unique globally asymptotically stable periodic solution with strictly positive components of periodic n-species Lotka-Volterra competition system with several delays. Some new results are obtained. As an application, we also examine some special cases of the system we considered, which have been studied extensively in the literature. Some known results are improved and generalized.     

具有阶段结构的两种群竞争系统的周期解

本文研究了具有阶段结构的两种群竞争系统的渐近行为.我们得到了系统持续生存的条件.由Brouwer不动点定理和李亚普诺夫函数,我们证明相应的周期系统在满足一定的条件下,存在一个唯一的全局渐近稳定的正周期解.最后我们把没有阶段结构的系统与有阶段结构的系统进行了比较.     

Dynamic reduction with applications to mathematical biology and other areas

In a difference or differential equation one is usually interested in finding solutions having certain properties, either intrinsic properties (e.g. bounded, periodic, almost periodic) or extrinsic properties (e.g. stable, asymptotically stable, globally asymptotically stable). In certain instances it may happen that the dependence of these equations on the state variable is such that one may (1) alter that dependency by replacing part of the state variable by a function from a class having some of the above properties and (2) solve the 'reduced' equation for a solution having the remaining properties and lying in the same class. This then sets up a mapping Τ of the class into itself, thus reducing the original problem to one of finding a fixed point of the mapping. The procedure is applied to obtain a globally asymptotically stable periodic solution for a system of difference equations modeling the interaction of wild and genetically altered mosquitoes in an environment yielding periodic parameters. It is also shown that certain coupled periodic systems of difference equations may be completely decoupled so that the mapping Τ is established by solving a set of scalar equations. Periodic difference equations of extended Ricker type and also rational difference equations with a finite number of delays are also considered by reducing them to equations without delays but with a larger period. Conditions are given guaranteeing the existence and global asymptotic stability of periodic solutions.     

具有阶段结构和功能反应混合模型的持久性和周期解

讨论了一类具有阶段结构和第Ⅱ类功能反应的三种群混合模型,其中捕食种群具有阶段结构．得到在适当的条件下系统的持续生存,对应周期系统正周期解的存在性、唯一性以及全局稳定性．     

三种群非自治系统的周期解的吸引性

本文讨论了三种群非自治竞争系统，此系统中所有的参数是与时间相关的，并且分别渐近接近于周期函数，得到了保证周期解吸引性的条件。     

Periodic metabolic systems: Oscillations in multiple-loop negative feedback biochemical control networks

Summary For a general multiple loop feedback inhibition system in which the end product can inhibit any or all of the intermediate reactions it is shown that biologically significant behaviour is always confined to a bounded region of reaction space containing a unique equilibrium. By explicit construction of a Liapunov function for the general n dimensional differential equation it is shown that some values of reaction parameters cause the concentration vector to approach the equilibrium asymptotically for all physically realizable initial conditions. As the parameter values change, periodic solutions can appear within the bounded region. Some information about these periodic solutions can be obtained from the Hopf bifurcation theorem. Alternatively, if specific parameter values are known a numerical method can be used to find periodic solutions and determine their stability by locating a zero of the displacement map. The single loop Goodwin oscillator is analysed in detail. The methods are then used to treat an oscillator with two feedback loops and it is found that oscillations are possible even if both Hill coefficients are equal to one.     

Transient oscillations induced by delayed growth response in the chemostat

In this paper, in order to try to account for the transient oscillations observed in chemostat experiments, we consider a model of single species growth in a chemostat that involves delayed growth response. The time delay models the lag involved in the nutrient conversion process. Both monotone response functions and nonmonotone response functions are considered. The nonmonotone response function models the inhibitory effects of growth response of certain nutrients when concentrations are too high. By applying local and global Hopf bifurcation theorems, we prove that the model has unstable periodic solutions that bifurcate from unstable nonnegative equilibria as the parameter measuring the delay passes through certain critical values and that these local periodic solutions can persist, even if the delay parameter moves far from the critical (local) bifurcation values.When there are two positive equilibria, then positive periodic solutions can exist. When there is a unique positive equilibrium, the model does not have positive periodic oscillations and the unique positive equilibrium is globally asymptotically stable. However, the model can have periodic solutions that change sign. Although these solutions are not biologically meaningful, provided the initial data starts close enough to the unstable manifold of one of these periodic solutions they may still help to account for the transient oscillations that have been frequently observed in chemostat experiments. Numerical simulations are provided to illustrate that the model has varying degrees of transient oscillatory behaviour that can be controlled by the choice of the initial data.Mathematics Subject Classification: 34D20, 34K20, 92D25Research was partially supported by NSERC of Canada.This work was partly done while this author was a postdoc at McMaster.     

State-dependent impulsive models of integrated pest management (IPM) strategies and their dynamic consequences

A state-dependent impulsive model is proposed for integrated pest management (IPM). IPM involves combining biological, mechanical, and chemical tactics to reduce pest numbers to tolerable levels after a pest population has reached its economic threshold (ET). The complete expression of an orbitally asymptotically stable periodic solution to the model with a maximum value no larger than the given ET is presented, the existence of which implies that pests can be controlled at or below their ET levels. We also prove that there is no periodic solution with order larger than or equal to three, except for one special case, by using the properties of the LambertW function and Poincare map. Moreover, we show that the existence of an order two periodic solution implies the existence of an order one periodic solution. Various positive invariant sets and attractors of this impulsive semi-dynamical system are described and discussed. In particular, several horseshoe-like attractors, whose interiors can simultaneously contain stable order 1 periodic solutions and order 2 periodic solutions, are found and the interior structure of the horseshoe-like attractors is discussed. Finally, the largest invariant set and the sufficient conditions which guarantee the global orbital and asymptotic stability of the order 1 periodic solution in the meaningful domain for the system are given using the Lyapunov function. Our results show that, in theory, a pest can be controlled such that its population size is no larger than its ET by applying effects impulsively once, twice, or at most, a finite number of times, or according to a periodic regime. Moreover, our theoretical work suggests how IPM strategies could be used to alter the levels of the ET in the farmers' favour.     

稀疏效应下周期系数捕食-被捕食系统的全局渐近稳定性

研究一类稀疏效应下周期系数捕食-被捕食系统,得到了该系统存在唯一全局渐近稳定的正周期解的充分条件．     

具有庇护所的三种群捕食者-食饵模型

讨论了一类具有庇护所的自治三种群捕食者一食饵模型,运用Liapunov函数方法,得到了该模型持久性的充分条件．对于该模型的周期系统,在一定的条件下,将产生唯一一个全局渐近稳定的周期正解．对更具普遍意义的概周期现象,也得出了概周期正解唯一存在且全局渐近稳定性的充分条件．     

Almost periodic solution of non-autonomous Lotka-Volterra predator-prey dispersal system with delays

This paper studies a non-autonomous Lotka-Volterra almost periodic predator-prey dispersal system with discrete and continuous time delays which consists of n-patches, the prey species can disperse among n-patches, but the predator species is confined to one patch and cannot disperse. By using comparison theorem and delay differential equation basic theory, we prove the system is uniformly persistent under some appropriate conditions. Further, by constructing suitable Lyapunov functional, we show that the system is globally asymptotically stable under some appropriate conditions. By using almost periodic functional hull theory, we show that the almost periodic system has a unique globally asymptotical stable strictly positive almost periodic solution. The conditions for the permanence, global stability of system and the existence, uniqueness of positive almost periodic solution depend on delays, so, time delays are "profitless". Finally, conclusions and two particular cases are given. These results are basically an extension of the known results for non-autonomous Lotka-Volterra systems.     

具有周期系数和连续时滞的扩散模型的周期解

本文讨论了具有周期系数和连续时滞的竞争扩散模型,得到了保证其存在唯一周期解及全局渐近稳定的充分条件．     

Effects of quarantine in six endemic models for infectious diseases

Thresholds, equilibria, and their stability are found for SIQS and SIQR epidemiology models with three forms of the incidence. For most of these models, the endemic equilibrium is asymptotically stable, but for the SIQR model with the quarantine-adjusted incidence, the endemic equilibrium is an unstable spiral for some parameter values and periodic solutions arise by Hopf bifurcation. The Hopf bifurcation surface and stable periodic solutions are found numerically.     

具性别和阶段结构的非自治二种群捕食者-食饵系统研究

讨论了一类食饵具有性别结构,捕食者具有阶段结构的非自治捕食者．食饵系统,运用Liapunov函数方法,得到了该系统一致持续生存的充分条件．对于该模型的周期系统,在适当条件下,存在唯一、全局渐近稳定的周期解．对更具普遍意义的概周期现象,也得出了概周期正解唯一存在且全局渐近稳定的充分条件．