Siphodinarella costata n. gen., n. sp., a new benthic foraminifer from the Coniacian of the Adriatic Carbonate Platform (Slovenia, Croatia)

A new benthic foraminifer is described as Siphodinarella costata n. gen., n. sp. from Coniacian shallow-water platform-interior carbonates of Slovenia and Croatia. The new foraminifer is found in skeletal wackestone in association with small benthic foraminifera, thaumatoporellaceans, and calcimicrobes (Decastronema, Girvanella-type tubes). The existence of an internal siphon in Siphodinarella n. gen. is interpreted as an entosolenian tube and discussed in terms of its generic and suprageneric importance.     

Green algae from the Lower Cretaceous carbonate of northern Shiraz (Zagros Mountains,SW Iran)

In northern Shiraz (SW Iran), Lower Cretaceous carbonate was studied in detail. In this study, nine species of dasycladacea algae were classified. There are different species of dasycladacea algae which belong to seven different genera: Actinoporella, Cylindroporella, Dissocladela, Heteroporella, Neomeris, Salpingoporella, Trinocladus; one species of udoteaceae belongs to Bouenia; one species of acetabulariaceae belongs to Clypeina and the microproblematicum Coptocampylodon was also seen. Among the green algae, dasycladaceae and acetabulariceae are the most frequent and udoteaceans are rare in Zagros Mountains. The genus of Trinocladus is a new record for Lower Cretaceous (Upper Albian) in SW Iran.     

On Late Jurassic and Early Cretaceous oysters (Bivalvia,Ostreidae) from northern Siberia

The morphology and taxonomic value of morphological features of Late Jurassic and Early Cretaceous oysters from northern Siberia are considered. A new species, Deltoideum exoticum Kosenko, sp. nov. is described.     

Morphological abnormalities in Late Cretaceous and early Paleocene foraminifer tests (northern Patagonia, Argentina)

Morphological abnormalities are common in Late Cretaceous and early Paleocene foraminifer tests at two localities in northern Patagonia, Argentina. Protelphidium sp. in the Auca Mahuevo section (late Campanian–early Maastrichtian) exhibit abnormal size or shape of the later chambers, with the last chamber commonly larger than normal or inflated and variably extending onto one of the lateral sides of the test; modification of the coiling plane; protuberances near the proloculus or on one or more chambers; a double last chamber, and complex forms. Protelphidium hofkeri Haynes in the Cerro Azul section (Danian) exhibit abnormal size or shape of one or more chambers, producing peripheral irregularities. In addition, there are rare multiple tests in planktic species from the Cerro Azul section, probably teratological specimens. The sedimentology of the sections and the character of the accompanying faunas indicate that the abnormalities in the two benthic foraminiferal taxa were most probably caused by hypersalinity and/or fluctuations in salinity. The fossil occurrence of assemblages with abundant deformed specimens suggests that investigators should carefully look at many aspects of the environment before concluding that anthropogenic pollution is the only cause of deformations of living benthic foraminifera.     

A fossil feather from the Upper Cretaceous of Kras (Slovenia)

A fossil feather preserved as a carbonised trace is described from the Tomaj Limestone at Kri , in the Kras region of southwestern Slovenia. The Tomaj Limestone is a platy and laminated limestone with cherts, which occurs within a well-bedded rudist limestone of the Santonian–Campanian Lipica Formation. It was deposited in a lagoon environment and has yielded a diverse fossil assemblage. Whether this feather belonged to a bird or to a dinosaur is unclear, but it is an addition to the scanty record of Late Cretaceous feathers, from a palaeobiogeographically interesting area.     

Chondrichthyans from a Cenomanian (Late Cretaceous) bonebed,Saskatchewan, Canada

Abstract: Acid preparation of samples of a bonebed from the Cenomanian of central Canada yielded several thousand well‐preserved chondrichthyan teeth, in addition to numerous other vertebrate remains. Teeth and other remains of one species of chimaeroid, one species of hybodont shark, three species of Ptychodus, 10 species of neoselachian sharks and two species of batoid were recorded. The family Archaeolamnidae fam. nov., genera Meristodonoides gen. nov. and Telodontaspis gen. nov. and species Ptychodus rhombodus sp. nov., Telodontaspis agassizensis gen et sp. nov., Eostriatolamia paucicorrugata sp. nov., Roulletia canadensis sp. nov., Cretorectolobus robustus sp. nov. and Orectoloboides angulatus sp. nov. are described. Status of the genus Palaeoanacorax and the species Cretoxyrhina denticulata, Squalicorax curvatus and ‘Rhinobatos’incertus are discussed, and reconstructed dentitions of Archaeolamna and Roulletia presented. The fauna is of low diversity and dominated by active hunters, with many species apparently endemic to the northern Western Interior Seaway.     

Reconstruction of Late Cretaceous coastal paleotemperature from halite deposits of the Late Cretaceous Nongbok Formation (Khorat Plateau,Laos)

Cretaceous evaporites of the Maha Sarakhan Formation in Thailand (e.g., the Nongbok Formation, Laos) have been studied for almost a century as the huge potash deposits in the world. The consistently high local paleotemperatures should lead to huge salt deposits during the evaporation process. Primary fluid inclusions in halite can provide surface brine water temperatures directly and quantitatively. Until now, there have been no data published from paleotemperature of primary fluid inclusions of Cretaceous halite. The non-marine halite from the Cretaceous Nongbok Formation (Laos) precipitated from shallow brine waters with temperatures of 17.7–42.3 °C.     

Palaeogeography and geodynamic evolution of the Gosau Group of the Northern Calcareous Alps (Late Cretaceous, Eastern Alps, Austria)

The sedimentation of the Late Cretaceous Gosau Group of the Northern Calcareous Alps records the complex geodynamic evolution of the northern part of the Austroalpine microplate. The tectonically controlled sedimentation and facies distribution were mainly governed by the oblique subduction of the Pennic-Ligurian ocean below the Austroalpine margin. Most of the Gosau successions comprise two distinct sedimentary complexes. The Lower Gosau Subgroup (upper Turonian-Campanian) is characterized by terrestrial to shallow-marine facies associations: alluvial fan and fan delta deposits, shallow-marine sandstones and sandy limestones, and storm-influenced nearshore and shelf deposits. During the early Santonian sea-level highstand, a great part of the Northern Calcareous Alps was covered by a shallow shelf sea. Palaeotransport directions point to source areas situated both to the north and to the south of the Northern Calcareous Alps.

The Upper Gosau Subgroup (upper Santonian-Eocene) comprises deep-water hemipelagic and turbiditic deposits. Rapid subsidence into bathyal to abyssal depths started diachronously from the northwest (Santonian) to the southeast (Maastrichtian) after a short phase of deformation and erosion. Abyssal deposits in the northern parts of the Northern Calcareus Alps and north-directed palaeotransport indicate a tilting of the slope towards the north. This subsidence pulse is attributed to an event of subcrustal tectonic erosion, probably triggered by the subduction of a topographic high of the South Penninic ocean. The relationships of the Gosau Group of the Austroalpine unit to Late Cretaceous deposits of the Carpathians and Southern Alps are discussed in the framework of a plate tectonic model.     

Giant Bathysiphon (Foraminiferida) from Cretaceous turbidites, northern California

Straight-sided, tubular fossils occur together with a diverse assemblage of deep-water trace fossils in linegrained Franciscan turbidites at Point Saint George. Pacific Coast of California. Based on comparison with living abyssal organisms and on skeletal microstructure, these tubes are interpreted as recrystallized siliceous tests of the large abyssohenthic foraminiferid, Bathysiphon. Tubes found preserved in-place show that life orientation of the Franciscan species was vertical with the broader, mature end protruding several millimetres above the muddy seafloor. Positioned in this way the organisms probably were suspension feeders or carnivores. Bathysiphon and trace fossils taken together reflect a diverse deep-water benthic community that was disrupted (or locally eliminated) repeatedly by turbidity currents reaching the outer margins of a submarine fan. ▭ Bathysiphon, deep-water benthos, outer fan/basin floor, trace fossils, Franciscan Complex, flysch, Cretaceous.     

The evolution and phytoplankton composition of mucilaginous aggregates in the northern Adriatic Sea

Recurrent occurrences of visible mucilage “clouds” that cover areas up to several hundred kilometres with vertical dimensions of 20–30 m have been recorded in the stratified water column in the northern Adriatic. In the past this was described as “mare sporco” phenomenon. Past studies of the phenomenon indicated that phytoplankton is an important component of mucilage. Our research revealed the composition of phytoplankton assemblages in different types of mucilaginous aggregates collected during the summers of 1997 and 2000 using pigment biomarkers (HPLC). Phytoplankton biomass in the mucilage samples was very high, ranging from 7.9 μg g⁻¹ to 390.8 μg g⁻¹ of chlorophyll a per unit of dry mass of mucilage. The phytoplankton community in the early, loose stage of mucilaginous aggregates was heterogeneous, as indicated by the diversity of detected pigments. The number of phytoplankton groups decreased as the aggregates aged and diatoms increased in relative biomass (up to 92.7%). Phytoplankton biomass in seawater was similar in years with and without mucilage; however, significantly higher contributions to the total biomass of 19′-hexanoyloxyfucoxanthin-containing phytoplankton (prymnesiophytes) were found in the upper 10 m in spring of the “mucilaginous years” (1997 and 2000) followed by prevalence of diatoms in summer. The F_p pigment index used to assess seawater trophic conditions reached lower values in April–May in mucilaginous years (1997 and 2000) compared to non-mucilaginous years (1998 and 1999). We conclude that the role of prymnesiophytes and other small flagellates is crucial for the initial phases of mucilage appearance. Aggregates represent a favourable environment for the secondary development of opportunistic diatoms that foster mucilage formation.     

Cranial and appendicular ontogeny of Bactrosaurus johnsoni,a hadrosauroid dinosaur from the Late Cretaceous of northern China

Abstract: The juvenile anatomy of various cranial and appendicular elements of the hadrosauroid dinosaur Bactrosaurus johnsoni is described in detail. Growth changes are documented from juvenile to adult stages for each skeletal element available. In the studied skull, ontogenetic trends consist of: development of features on the ventral surface of the frontal; reduction in the slope of the posteromedial process of the premaxilla; a posterior shift of the dorsal process of the maxilla; development of concavities on the medial surface of the prefrontal; increased robustness and development of the ventral flange of the jugal; decreased curvature of the long axis of the quadrate; increased ventral deflection of the dentary; and changes in the length/width proportions and depth of the anterior surface of the predentary. In the appendicular skeleton, the majority of ontogenetic variation from juvenile to adult occurs in the limb bones, including increased robustness of the deltopectoral crest of the humerus; relative shortening of the ulna; increased development of the fourth trochanter and mediolateral widening of the distal end of the femur; increased expansion of the cnemial crest of the tibia; and the increased prominence of articular protuberances and flanges of the metatarsals. A survey of the phylogenetically informative characters present in B. johnsoni indicates that several characters concerning the frontal, maxilla, jugal, quadrate, predentary, dentary, scapula, humerus and ilium are affected by ontogeny. Nevertheless, the majority of phylogenetic characters are not ontogenetically variable, suggesting that a substantial amount of the information provided by juvenile and subadult specimens for phylogenetic inference is reliable in basal hadrosauroids.     

Biostratigraphy and sedimentary evolution of Late Miocene and Pliocene continental deposits of the Granada Basin (southern Spain)

The Late Miocene and Pliocene continental sediments in the Granada Basin (southern Spain) have yielded large amounts of fossil small mammals in 37 localities from 11 sections. The aim of this paper is to integrate faunistic, stratigraphic, and sedimentary criteria to unravel the geological history of the continental infilling of the basin. The palaeontological study has led to a detailed biozonation on the basis of rodents, which helps to correlate in detail the different sedimentary units found in the basin, and to follow the changes of the different sedimentary systems and their palaeogeographical evolution through time. Combination of the proposed biostratigraphy and the reinterpretation of the magnetostratigraphic analyses of the Barranco del Purcal section allows us to assign an absolute age slightly older than 5.23 Ma to the Turolian–Ruscinian boundary (MN13‐MN14).     

Skeletal anatomy of the Late Cretaceous shark,Squalicorax (Neoselachii: Anacoracidae)

The paleobiology of the Cretaceous neoselachian shark,Squalicorax, has largely been based on isolated teeth. We examined partial and nearly complete skeletons of three species ofSqualicorax, S. falcatus (Aoassiz),S. kaupi (Agassiz), andS. pristodontus (Agassiz), that were collected from the U.S.A. These specimens suggest that the total body length (TL) ofS. falcatus typically measured 1.8–2.0 m, and probably did not exceed 3 m. Moderatesized individuals ofS. kaupi andS. pristodontus perhaps measured about 3 m TL. AlthoughS. pristodontus was the largest form among the three species examined, this taxon possessed a set of large jaws (with large but fewer teeth) relative to its body size compared toS. falcatus orS. kaupi. This suggests that tooth size is not an accurate indicator of the TL if one compares oneSqualicorax species to another. Neurocranial features suggest that the vision ofSqualicorax was not as acute as that of a contemporaneous macrophagous lamniform shark,Cretoxyrhina mantelli (Agassiz) , but olfaction ofSqualicorax may have been better thanC. mantelli. The morphology of placoid scales suggests thatSqualicorax was capable of fast swimming. New skeletal data support the view that the feeding dynamics ofSqualicorax was similar to the modern tiger shark (Galeocerdo Müller & Henle). The present data do not allow for exact ordinal placement, but, contrary to some previous interpretations,Squalicorax can be excluded from the Hexanchiformes and Orectolobiformes. The taxon should more appropriately be placed within the Lamniformes or Carcharhiniformes.      

Live colouration,morphology and habitat of Vanneaugobius dollfusi (Gobiidae) in the northern Adriatic Sea

The rare deepwater gobiid fish Vanneaugobius dollfusi, hitherto described only from limited material identified in collections, was encountered and collected during scuba dives at depths of 30–45 m in the northern Adriatic Sea.     

A new aquatic pythonomorph (Reptilia,Squamata) from the Turonian (Late Cretaceous) of France

Disarticulated vertebrae from the Turonian of France display a distinctive suite of characters and probably represent a new pythonomorph. This taxon displays some degree of vertebral pachyostosis s.s., often observed in varanoid squamates from the Cenomanian-Turonian interval of the ‘Mediterranean’ portion of the Tethys. The discovery of this new material highlights the importance of also describing possibly new taxa based on isolated remains.     

Revised radiolarian zonation of the Upper Cretaceous Izumi inter-arc basin (SW Japan)

The radiolarian zonation of the Izumi Group is revised for an integrated biostratigraphy across the Campanian/Maastrichtian boundary, together with the ammonite occurrence, fission track and magnetostratigraphic calibration of the acidic tuff key beds. The significance of direct correlation in the same sections, between macro- and micro-fossils (i.e. between relatively near-shore nektonic and turbiditic planktonic-fauna) is discussed in relation to the litho- and bio-facies of the Izumi group. Late Cretaceous paleoceanographic changes inferred from the studied radiolarian fauna represent a regional marker event for the chronostratigraphic correlation of continental shelf to turbidite deep-sea sequences, which accumulated on a mid latitudinal margin facing towards the NW paleo-Pacific. The proposed Campanian/Maastrichtian boundary is at the limit between Pachydiscus awajiensis and Nostoceras hetonaiense ammonite zones. This boundary corresponds to the radiolarian last occurrence of Amphipyndax pseudoconulus (= A. enesseffi), Archaeodictyomitra sliteri, Dictyomitra formosa, D. koslovae and Rhopalosyringium magnificum at the top of the Myllocercion acineton Assemblage Zone (uppermost Campanian), and the first occurrence of Clathrocyclas gravis and Dorcadospyris sp. A, at the base of the Maastrichtian C. gravis Assemblage Zone. The new data settle partly a chronostratigraphic discrepancy between radiolarian and ammonite zonations in the Izumi Group, which was pointed out previously by Ishida et al. (2010).     

Squatiniformes (Chondrichthyes,Neoselachii) from the Late Cretaceous of southern England and northern France with redescription of the holotype of Squatina cranei Woodward, 1888

Abstract: Bulk sampling of phosphate‐rich horizons within the Late Cretaceous of the Anglo‐Paris Basin yielded numerous teeth of members of the Squatiniformes. Along with isolated tooth remains, two museum specimens comprising partial articulated encoskeletal remains including the holotype of the species Squatina cranei Woodward, 1888a are described, and a new subgenus Cretascyllium is proposed for species of the genus Squatina with high degree of heterodonty and triangular anterior teeth. The species Squatina (Cretascyllium) cranei comb. nov. and Squatina (Cretascyllium) hassei comb. nov. are referred to this subgenus. The genus Parasquatina Herman, 1982 previously erected on a single tooth is valid, and two new species P. justinensis sp. nov. and P. jarvisi sp. nov. are described along with a third taxon Parasquatina sp. An enigmatic tooth referred to ?Neoselachii incertae sedis is also reported. The palaeoecology of these taxa is discussed.     

Late Triassic sedimentary evolution of Slovenian Basin (eastern Southern Alps): description and correlation of the Slatnik Formation

Successions of the Slovenian Basin structurally belong to the easternmost Southern Alps. During the Late Triassic, they were part of the Adriatic continental margin. Norian–Rhaetian successions of the Slovenian Basin are characterized mainly by dolomite of the Bača Dolomite Formation. However, in the northern part of the basin, Late Triassic limestone is preserved above Bača Dolomite Formation and is formalized as the Slatnik Formation. It is composed of hemipelagic limestone alternating with resedimented limestones. The succession documents an upward progradation of the slope environment composed of three high-frequency cycles. Most prominent progradation is referred to the second, i.e., Early Rhaetian cycle. The Slatnik Formation ends with thin-bedded hemipelagic limestone that records the end-Triassic productivity crisis, or rapid sea-level fall. The overlying resedimented limestones of the Early Jurassic Krikov Formation, document the recovery of production and shedding from the adjacent carbonate platform.