A new specimen of the thyreophoran dinosaur cf. Scelidosaurus with soft tissue preservation

A new specimen, comprising eight articulated caudal vertebrae in a cut slab of blue/grey carbonate mudstone, is comparable to the early thyreophoran dinosaur Scelidosaurus harrisonii Owen from the English Lower Jurassic. Palynological analysis indicates that the specimen is probably late Hettangian–Sinemurian (Early Jurassic), but the exact horizon and locality remain uncertain. The slab has been cut longitudinally in a parasagittal plane along the vertebral series and exposes several articulated centra, haemal arches, neural arches and osteoderms. An envelope of preserved soft tissue wraps around the vertebrae and includes osteoderms with organic material on both the upper and lower surfaces. Basal thyreophoran dinosaur osteoderms were covered by a horny sheath during life.     

Dinosaurier aus Nordostdeutschland

Remains of dinosaurs in NE Germany The former clay pit of Grimmen (NE Germany) represents a glacially dislocated outcrop of upper Lower Jurassic shallow‐marine strata. Apart from typical marine organisms, the Lower Jurassic of Grimmen has also yielded the remains of the basal thyreophoran dinosaur Emausaurus ernsti. E. ernsti was a comparatively small dinosaur, generally characterized by a dermal armor composed of small isolated scutes. Recently, the remains of a second dinosaur belonging to the sauropod clade have been identified. The specimen represents the second Early Jurassic basal sauropod from Germany ever found. The sauropod was probably washed into the marine realm from nearby coastal land after its death, whereas the compacted pile of skeletal elements of E. ernsti found in a concretion is suspected to be a Speiballen produced by a large‐scaled marine reptile.     

A palaeoequatorial ornithischian and new constraints on early dinosaur diversification

Current characterizations of early dinosaur evolution are incomplete: existing palaeobiological and phylogenetic scenarios are based on a fossil record dominated by saurischians and the implications of the early ornithischian record are often overlooked. Moreover, the timings of deep phylogenetic divergences within Dinosauria are poorly constrained owing to the absence of a rigorous chronostratigraphical framework for key Late Triassic–Early Jurassic localities. A new dinosaur from the earliest Jurassic of the Venezuelan Andes is the first basal ornithischian recovered from terrestrial deposits directly associated with a precise radioisotopic date and the first-named dinosaur from northern South America. It expands the early palaeogeographical range of Ornithischia to palaeoequatorial regions, an area sometimes thought to be devoid of early dinosaur taxa, and offers insights into early dinosaur growth rates, the evolution of sociality and the rapid tempo of the global dinosaur radiation following the end-Triassic mass extinction, helping to underscore the importance of the ornithischian record in broad-scale discussions of early dinosaur history.     

The 'fabrosaurid' ornithischian dinosaurs of the Upper Elliot Formation (Lower Jurassic) of South Africa and Lesotho

The Upper Elliot Formation of South Africa and Lesotho contains the world's most diverse fauna of early Jurassic ornithischian dinosaurs. Nevertheless, despite four decades of work on this fauna there remains significant taxonomic confusion and many important specimens remain undescribed. A review of the non-heterodontosaurid ('fabrosaurid') ornithischians of the Upper Elliot Formation is presented, following re-examination of all known ornithischian material from the Elliot Formation. ' Fabrosaurus australis' is based upon a single undiagnostic dentary, and is here considered a nomen dubium . Lesothosaurus diagnosticus is considered to be valid and is rediagnosed based upon a unique combination of plesiomorphic and derived characteristics. Stormbergia dangershoeki gen. et. sp. nov. is described from three partial skeletons including numerous postcranial material. Stormbergia dangershoeki is significantly larger than previously described Elliot Formation ornithischians, and can be recognized on the basis of a unique combination of characters, the most important of which is the possession of a distinctive tab-shaped obturator process on the ischium. A preliminary systematic analysis is presented, the results of which differ significantly from other recent ornithischian phylogenies.  © 2005 The Linnean Society of London, Zoological Journal of the Linnean Society , 2005, 145 , 175–218.     

The anatomy of the basal ornithischian dinosaur Eocursor parvus from the lower Elliot Formation (Late Triassic) of South Africa

Ornithischia is a morphologically and taxonomically diverse clade of dinosaurs that originated during the Late Triassic and were the dominant large‐bodied herbivores in many Cretaceous ecosystems. The early evolution of ornithischian dinosaurs is poorly understood, as a result in part of a paucity of fossil specimens, particularly during the Triassic. The most complete Triassic ornithischian dinosaur yet discovered is Eocursor parvus from the lower Elliot Formation (Late Triassic: Norian–Rhaetian) of Free State, South Africa, represented by a partial skull and relatively complete postcranial skeleton. Here, the anatomy of Eocursor is described in detail for the first time, and detailed comparisons are provided to other basal ornithischian taxa. Eocursor is a small‐bodied taxon (approximately 1 m in length) that possesses a plesiomorphic dentition consisting of unworn leaf‐shaped crowns, a proportionally large manus with similarities to heterodontosaurids, a pelvis that contains an intriguing mix of plesiomorphic and derived character states, and elongate distal hindlimbs suggesting well‐developed cursorial ability. The ontogenetic status of the holotype material is uncertain. Eocursor may represent the sister taxon to Genasauria, the clade that includes most of ornithischian diversity, although this phylogenetic position is partially dependent upon the uncertain phylogenetic position of the enigmatic and controversial clade Heterodontosauridae. © 2010 The Linnean Society of London, Zoological Journal of the Linnean Society, 2010, 160 , 648–684.     

THE AFFINITIES OF THE ENIGMATIC DINOSAUR ESHANOSAURUS DEGUCHIIANUS FROM THE EARLY JURASSIC OF YUNNAN PROVINCE, PEOPLE'S REPUBLIC OF CHINA

Abstract:  Eshanosaurus deguchiianus is based on a single left dentary from the Lower Lufeng Formation (Lower Jurassic) of Yunnan Province, China. It was originally identified as the earliest known member of Therizinosauroidea (Theropoda: Coelurosauria), a conclusion that results in a significant downward range extension for this clade (>65 million years) and for many other major lineages within Coelurosauria. However, this interpretation has been questioned and several authors have proposed that the anatomical features used to refer Eshanosaurus to Therizinosauroidea are more consistent with attribution to a basal sauropodomorph dinosaur. Detailed consideration of the holotype specimen suggests that several features of the dentary and dentition exclude Eshanosaurus from Sauropodomorpha and support its inclusion within Therizinosauroidea. If accepted as an Early Jurassic coelurosaur, Eshanosaurus has important implications for understanding the timing and tempo of early theropod diversification. Moreover, its provenance also suggests that substantial portions of the coelurosaur fossil record may be missing or unsampled. However, the Early Jurassic age of Eshanosaurus requires confirmation if this taxon is to be fully incorporated into broader evolutionary studies.     

产自中国云南下禄丰组(下侏罗统)的爬行动物不完美硕鳄的再研究(英文)

一般认为植龙类在三叠纪末期灭绝,但有研究显示它们可能在欧洲和亚洲延续到了侏罗纪最早期。来自亚洲的证据是产自中国云南下禄丰组(?赫唐阶-锡内穆阶)的不完美硕鳄(Pachysuchus imperfectus)。重新研究显示不完美硕鳄正型标本和植龙类存在许多不同点,它可能属于一个保存较差的、分类位置无法确定的蜥脚型类恐龙头骨。这一属种的归入标本也无法明确归入某一类群,因此亚洲没有可靠的晚于三叠纪的植龙类化石记录。欧洲的侏罗纪植龙类化石也存在着同样的问题。因而认为,植龙类的化石记录仅限于三叠纪晚期。     

The Lower Jurassic ornithischian dinosaur Heterodontosaurus tucki Crompton & Charig, 1962: cranial anatomy,functional morphology,taxonomy, and relationships

The cranial anatomy of the Lower Jurassic ornithischian dinosaur Heterodontosaurus tucki Crompton & Charig, 1962 is described in detail for the first time on the basis of two principal specimens: the holotype (SAM‐PK‐K337) and referred skull (SAM‐PK‐K1332). In addition several other specimens that have a bearing on the interpretation of the anatomy and biology of Heterodontosaurus are described. The skull and lower jaw of Heterodontosaurus are compact and robust but perhaps most notable for the heterodont dentition that merited the generic name. Details of the cranial anatomy are revealed and show that the skull is unexpectedly specialized in such an early representative of the Ornithischia, including: the closely packed, hypsodont crowns and ‘warping’ of the occlusal surfaces (created by progressive variation in the angulation of wear on successive crowns) seen in the cheek dentition; the unusual sutural relationships between the bones along the dorsal edge of the lower jaw; the very narrow, deeply vaulted palate and associated structures on the side wall of the braincase; and the indications of cranial pneumatism (more commonly seen in basal archosaurs and saurischian dinosaurs). Evidence for tooth replacement (which has long been recognized, despite frequent statements to the contrary) is suggestive of an episodic, rather than continuous, style of tooth replacement that is, yet again, unusual in diapsids generally and particularly so amongst ornithischian dinosaurs. Cranial musculature has been reconstructed and seems to conform to that typically seen in diapsids, with the exception of the encroachment of M. adductor mandibulae externus superficialis across the lateral surface of the temporal region and external surface of the lower jaw. Indications, taken from the unusual shape of the occlusal surfaces of the cheek dentition and jaw musculature, are suggestive of a novel form of jaw action in this dinosaur. The taxonomy of currently known late Karoo‐aged heterodontosaurids from southern Africa is reviewed. Although complicated by the inadequate nature of much of the known material, it is concluded that two taxa may be readily recognized: H. tucki and Abrictosaurus consors. At least one additional taxon is recognized within the taxa presently named Lanasaurus and Lycorhinus; however, both remain taxonomically problematic and their status needs to be further tested and may only be resolved by future discoveries. The only other named taxon, Geranosaurus atavus, represents an invalid name. The recognition of at least four distinct taxa indicates that the heterodontosaurids were speciose within the late Karoo ecosystem. The systematics of Heterodontosaurus and its congeners has been analysed, using a restricted sample of taxa. A basal (nongenasaurian) position within Ornithischia is re‐affirmed. There are at least four competing hypotheses concerning the phylogenetic placement of the Heterodontosauridae, so the evidence in support of the various hypotheses is reviewed in some detail. At present the best‐supported hypothesis is the one which places Heterodontosauridae in a basal (non‐genasaurian) position; however, the evidence is not fully conclusive and further information is still needed in respect of the anatomy of proximate outgroups, as well as more complete anatomical details for other heterodontosaurids. Heterodontosaurids were not such rare components of the late Karoo ecosystem as previously thought; evidence also suggests that from a phylogenetic perspective they occupied a potentially crucial position during the earliest phases of ornithischian dinosaur evolution. © 2011 The Linnean Society of London, Zoological Journal of the Linnean Society, 2011.     

A review of the platanaceous woods from the Eocene paratropical rainforest of south-east England

Small diameter pyritized axes, commonly referred to as 'twigs', of fossil platanaceous wood are described from the Lower Eocene London Clay Formation of south-east England. These twigs are characterized by solitary vessels with scalariform perforation plates, opposite intervessel pits, and tall, multiseriate rays that dilate in the phloem region. The wood anatomy supports close relationship to members of extant Platanaceae and the material is placed in the organ genus Plataninium Unger erected for fossil woods with close anatomical similarity to Platanus L. This material supplements the fossil record of platanaceous type wood from the Eocene London Clay and documents the first record of Plataninium decipiens Brett in the twig flora.  © 2002 The Linnean Society of London, Botanical Journal of the Linnean Society , 2002, 139 , 181–191.     

A Late Jurassic salamander (Amphibia: Caudata) from the Morrison Formation of North America

Despite some remarkable recent discoveries, the Mesozoic fossil record of salamanders remains limited, particularly for the Jurassic. Here we describe the first articulated salamander skeleton from the Jurassic of Euramerica, recovered from Upper Jurassic deposits of the Morrison Formation, Dinosaur National Monument, USA. The specimen was studied using both conventional methods and high-resolution computed tomography. It shows a combination of primitive and derived character states that distinguish it from all known Mesozoic salamanders and which permit the erection of a new genus and species, Iridotriton hechti . The derived states (including the presence of spinal nerve foramina in the tail) suggest a position on the stem of the Salamandroidea. Together with microvertebrate material from Britain, Portugal, and North America, this specimen confirms the presence of both stem- and crown-group salamanders in Euramerica from the Middle Jurassic (Bathonian) onwards, paralleling their evolution in Central and eastern Asia. This, in turn, provides qualified support for the current vicariance model of salamander evolution whereby basal caudates on an undivided Laurasian plate became separated into two populations by the incursion of the Turgai Sea in the Middle Jurassic, yielding Cryptobranchoidea in Asia and Salamandroidea in Euramerica.  © 2005 The Linnean Society of London , Zoological Journal of the Linnean Society , 2005, 143 , 599−616.     

First Thyreophoran Type Tracks from the Middle Jurassic Chuanjie Formation of Yunnan Province,China

Two tridactyl footprints from the Chuanjie Formation (Middle Jurassic) of Yunnan Province, China are morphological characteristics of thyreophoran tracks. They show some similarities to Shenmuichnus, known from the Early Jurassic strata of both Shaanxi and Yunnan provinces, but are somewhat larger, thereby resembling the ichnogenus Stegopodus. Based on their general morphology and size being congruent with this ichnogenus, they are tentatively assigned here to cf. Stegopodus. This is the fourth report of large ornithischian (probably thyreophoran) tracks from the Lower-Middle Jurassic of China that indicates relatively large trackmakers that were likely to be taxonomically distinct from much smaller and gracile Anomoepus trackmakers, also of ornithischian affinity. The larger tracks indicate a hitherto unreported abundance, size range and diversity of track types attributed to this group. The parallel orientation of the two best preserved trackways may indicate gregariousness.     

First record of fossil Mesozoic Ctenopoda (Crustacea, Cladocera)

Pre-Pleistocene representatives of the crustacean order Ctenopoda Sars, 1865 are unknown. Here we describe Mesozoic fossil specimens of ctenopods from two localities in Mongolia: Khotont (Jurassic/Cretaceous boundary, about 145 Mya) and Khutel Khara (Lower Cretaceous, about 129 Mya). All specimens apparently belonged to the tribe Latonini Korovchinsky, 1986 of the subfamily Sidinae Baird, 1850. At the Khotont site, ctenopods were the most numerous microscopic animal fossils. We assigned these cladocerans to a new genus and species Archelatona zherikhini gen. nov., sp. nov. Our findings support a pre-Cretaceous origin for cladocerans.  © 2006 The Linnean Society of London, Zoological Journal of the Linnean Society , 2006, 146 , 269–274.     

Reappraisal of the ill-defined Liassic pteridosperm Dichopteris using an ultrastructural approach

This review of the genus Dichopteris De Zigno is based on the original fossil specimens collected from the Lower Jurassic of the Grey Limestones of Veneto (Northern Italy). The cuticular diagnosis of the genus is emended and only Dichopteris visianica De Zigno is retained within the genus. In addition, transverse and longitudinal sections of cuticles examined using a transmission electron microscope (TEM) indicate close similarities between D. visianica and other pteridosperm cuticles, especially the occurrence of granules arranged in areas of different densities in the epidermal cells and stomatal apparatus. Moreover, the dichotomy and the impressive size of the frond of D. visianica imply a specific architecture with a typical arrangement of the pinnae and pinnules. They preclude an herbaceous habit for the plant. Finally, architectural, microstructural and ultrastructural observations are consistent with palaeoclimatic and palaeoenvironmental data, and most probably support an understorey habit for D. visianica .  © 2005 The Linnean Society of London, Botanical Journal of the Linnean Society , 2005, 149 , 313–332.     

Crocodilus affuvelensis Matheron, 1869 from the Late Cretaceous of southern France: a reassessment

Hitherto unpublished crocodylian remains, including four skull fragments, from the lignite-bearing localities of Valdonne and Fuveau (Santonian–Campanian), south-eastern France, are described. They complement the original material of ' Crocodilus affuvelensis ' described by Matheron (1869) and Repelin (1930). Based on overlapping specimens, available elements of the rostrum, the palate, the skull table, the mandible and the pterygoid can be shown to belong to a single taxon, for which a reconstruction is proposed. Designation of a neotype has been proposed to the ICZN committee. A new genus, Massaliasuchus , is proposed here to designate the crocodylian originally described as Crocodilus affuvelensis . This crocodylian presents affinities with basal alligatoroids and complements the picture of crocodylian diversity during the Late Cretaceous in the European archipelago. Massaliasuchus affuvelensis is compared with other European taxa and with basal alligatoroids from North America. The putative basal position of Massaliasuchus as well as its early geological age suggest that the geographical origin of the Alligatoroidea is still uncertain.  © 2008 The Linnean Society of London, Zoological Journal of the Linnean Society , 2008, 152 , 567–580.     

Two new basal crocodylomorph archosaurs from the Lower Jurassic and the monophyly of the Sphenosuchia

We report on and name two new taxa of basal crocodylomorph archosaurs from the Lower Jurassic, Litargosuchus leptorhynchus gen. et sp. nov. , from the upper Elliot Formation (Stormberg Group) of South Africa, and Kayentasuchus walkeri gen. et sp. nov. , from the Kayenta Formation (Glen Canyon Group) of Arizona, USA. Examination of this material led to a reconsideration of basal crocodylomorph interrelationships. A phylogenetic analysis found no support for the monophyly of Sphenosuchia.  © 2002 The Linnean Society of London. Zoological Journal of the Linnean Society , 2002, 136 , 77–95.     

Late Oligocene mesotheriids (Mammalia, Notoungulata) from Salla and Lacayani (Bolivia): implications for basal mesotheriid phylogeny and distribution

The oldest known mesotheriids are referred to the genus Trachytherus , a taxon traditionally included in its own subfamily 'Trachytheriinae'. The ontogeny, morphological variation and systematics of the genus Trachytherus are still poorly understood and the monophyly of this taxon is questionable. For the first time, a large collection of basal mesotheriids (belonging to the genus Trachytherus ) yielded by the Bolivian Salla beds (Deseadan, late Oligocene) and previously referred to T. spegazzinianus was included in a large-scale analysis. This has led to the recognition of a unique new species in the locality of Salla-Luribay, distinct from T. spegazzinianus : Trachytherus alloxus sp. nov . Skulls of this species are described and cheekteeth variation range throughout ontogeny is discussed. These new data are incorporated in a parsimony analysis of basal mesotheriids. The most probable hypothesis of a clade T. spegazzinianus -mesotheriines is discussed. The distinction of the new Salla species from the Patagonian species T. spegazzinianus confirms the faunal differences observed between these Bolivian and Patagonian Deseadan localities. Moreover, the assignment to Trachytherus spegazzinianus of mesotheriid remains from the Bolivian Lacayani deposits complicates the problem of these faunal differences. Further systematic investigations on the Lacayani fauna and on the age of the Patagonian localities are needed.  © 2008 The Linnean Society of London, Zoological Journal of the Linnean Society , 2008, 152 , 153–200.     

New evolutionary evidence of Grylloblattida from the Middle Jurassic of Inner Mongolia, north-east China (Insecta, Polyneoptera)

The small modern insect order Grylloblattida has an abundant fossil record during the Late Palaeozoic and the Mesozoicirca. The relationships between these fossil taxa and the modern grylloblattids remain unclear because most of them are based on isolated wings or have poorly preserved body features. Modern grylloblattids are wingless insects. The new grylloblattid family Plesioblattogryllidae fam. nov. is erected for the new genus and species Plesioblattogryllus magnificus gen. nov., sp. nov. , from the Middle Jurassic of north-eastern China. The well-preserved specimen provides further evidence that could support its close relationships with the modern grylloblattids: (1) several very similar head structures, e.g. developed laciniae with inner row of setae, maxillary palps segmented into five, labial palps segmented into three, large labrum, and morphology of antenna; (2) paired eoplantulae on tarsomeres 1–4; (3) long ovipositor and large eggs comparable with those of modern taxa. The new genus has strongly developed mandibles with sharp pointed apical teeth and strong marginal teeth, and strong hook-like fore claws with basal teeth, suggesting it was carnivorous. The major differences between the extinct and extant Grylloblattida, such as the lack of wings, the eyes and ocelli either degenerated or absent, and the thorax degenerated in the modern forms, are probably related to their adaptation to their life under rocks and rock-crawler habits.  © 2008 The Linnean Society of London, Zoological Journal of the Linnean Society , 2008, 152 , 17–24.     

Spinosaurid teeth from the Lower Cretaceous of Ko Kut,eastern Thailand

Isolated theropod dinosaur teeth from Ko Kut (Kut Island) in eastern Thailand are referred to an indeterminate spinosaurid on the basis of their morphology and ornamentation. On the basis of other spinosaurid occurrences in Thailand and other parts of Asia, they support the attribution of the fossil-bearing beds on Ko Kut to the Lower Cretaceous rather than the Jurassic. The lack of ornithischian remains in the Ko Kut faunal assemblage suggests that it is coeval with the Sao Khua Formation (Barremian) of NE Thailand.