The Middle Stone Age of the northern Kenyan Rift: age and context of new archaeological sites from the Kapedo Tuffs

Rift Valley sites in southern Ethiopia and northern Kenya preserve the oldest fossil remains attributed to Homo sapiens and the earliest archaeological sites attributed to the Middle Stone Age (MSA). New localities from the Kapedo Tuffs augment the sparse sample of MSA sites from the northern Kenya Rift. Tephrostratigraphic correlation with dated pyroclastic deposits from the adjacent volcano Silali suggests an age range of 135-123 ka for archaeological sites of the Kapedo Tuffs. Comparisons of the Kapedo Tuffs archaeological assemblages with those from the adjacent Turkana and Baringo basins show broad lithic technological similarity but reveal that stone raw material availability is a key factor in explaining typologically defined archaeological variability within this region. Spatially and temporally resolved comparisons such as this provide the best means to link the biological and behavioral variation manifest in the record of early Homo sapiens.     

A re-examination of the human fossil specimen from Bački Petrovac (Serbia)

A fragmented human calotte was discovered during the early 1950s near Ba?ki Petrovac (Serbia), in association with Palaeolithic stone tools. After its initial publication, the fossil specimen remained largely unknown outside of the Serbian academe and no detailed comparative study has ever been carried out. Since the whereabouts of the fossil itself are currently unknown, and given its potential significance for the Pleistocene human evolution, we re-examine the data published by 0255 and 0260. Using the original measurements, mostly taken on the frontal bone, and a wide comparative sample of 68 fossil specimens, the fossil was compared and analyzed by statistical multivariate methods. We also conducted a visual examination of the morphology based on the available photographic material. Our analysis reveals phenetic similarity with Middle Pleistocene archaic Homo from Africa and anatomically modern Homo sapiens. However, the absence of primitive cranial traits in Ba?ki Petrovac indicates a clear modern Homo sapiens designation. Although lost at the moment, there is a chance for the re-discovery of the fossil in the years to come. This would give us an opportunity to acquire absolute dates and to study the specimen in a more detailed manner.     

The Lower Palaeolithic assemblage of Layers 15–18 (Unit G) at Hummal. An exemplary case addressing the problems placing undated,archaic-looking stone tool assemblages in the Early and Lower Palaeolithic record by techno-typological classification

The site of Hummal is situated in the region of El Kowm in central Syria. The site comprises an archaeological sequence covering the entire Pleistocene epoch, and encompasses all major Palaeolithic complexes currently known in the Middle East. At the base of the site, 14 m below today's ground level, several layers with a lithic assemblage attributed to the Lower Palaeolithic have been excavated over the past years. At present, the collection recovered from this lowest succession at Hummal contains more than 700 stone artefacts and more than 3000 bone fragments. The lithic assemblage is characterized by a simple flaking technique and the presence of different pebble tools, such as choppers, hammerstones and sphaeroids. Additionally four handaxes were recovered, which have a symmetric shape, are clearly bifacial and rather flat. The lithic assemblage from the lowermost layers of the Hummal excavation largely resembles an Archaic, Lower Palaeolithic assemblage, belonging to the so-called Oldowan or Mode 1 stage. However, the presence of well-shaped and symmetric handaxes sheds doubt on the validity of this attribution to a Mode 1, Oldowan or the Early Acheulean Stage. It can, therefore, be debated, whether the common classifications of lithic industries are adequate for describing the archaeological record from the period in question in the Middle East.     

A Human Deciduous Tooth and New 40Ar/39Ar Dating Results from the Middle Pleistocene Archaeological Site of Isernia La Pineta,Southern Italy

Isernia La Pineta (south-central Italy, Molise) is one of the most important archaeological localities of the Middle Pleistocene in Western Europe. It is an extensive open-air site with abundant lithic industry and faunal remains distributed across four stratified archaeosurfaces that have been found in two sectors of the excavation (3c, 3a, 3s10 in sect. I; 3a in sect. II). The prehistoric attendance was close to a wet environment, with a series of small waterfalls and lakes associated to calcareous tufa deposits. An isolated human deciduous incisor (labelled IS42) was discovered in 2014 within the archaeological level 3 coll (overlying layer 3a) that, according to new ⁴⁰Ar/³⁹Ar measurements, is dated to about 583–561 ka, i.e. to the end of marine isotope stage (MIS) 15. Thus, the tooth is currently the oldest human fossil specimen in Italy; it is an important addition to the scanty European fossil record of the Middle Pleistocene, being associated with a lithic assemblage of local raw materials (flint and limestone) characterized by the absence of handaxes and reduction strategies primarily aimed at the production of small/medium-sized flakes. The faunal assemblage is dominated by ungulates often bearing cut marks. Combining chronology with the archaeological evidence, Isernia La Pineta exhibits a delay in the appearance of handaxes with respect to other European Palaeolithic sites of the Middle Pleistocene. Interestingly, this observation matches the persistence of archaic morphological features shown by the human calvarium from the Middle Pleistocene site of Ceprano, not far from Isernia (south-central Italy, Latium). In this perspective, our analysis is aimed to evaluate morphological features occurring in IS42.     

Baïraki – un site du Paléolithique inférieur sur le territoire de la Plaine d’Europe orientale

The paper presents the preliminary results of five years’ multidisciplinary researches (2010–2014 years) of the new Lower Palaeolithic Bayraki site, situated in the vicinity of Dubossary town in Moldavia, on the high terrace (VII) of the Dniester River. In the Lower and Middle Pleistocene deposits are discovered six layers including lithic artefacts of Lower Palaeolithic character. Two layers (I and II) relate to deposits dated by the Brunhes period and four layers (III, IV, V and VI) are attributed to the Matuyama period. The highest number of artefacts (884 objects) is issue from the layer V (the Jaramillo period), whose lithic industry has an Oldowan character. In other layers are found few flint archaeological objects and sporadic fragments of indeterminable bones. In the layer III is found a fossil horse mandibular (Equus sussenbornensis). At present, this site presents the oldest Low Pleistocene across all East European Plain.     

Les principales étapes de l’occupation humaine en bordure de l’Himalaya occidentale

Lithic industries attest that Western Himalayas, Hindu Kush and Pamir as well as the sub-Himalayan belt that is the Siwalik range, were inhabited during the prehistoric times. However, in thin transition zone between plains and high mountains, distinction is to be kept between low-middle altitude regions and high altitude regions (above 2000 m). In the former regions, the first human occupation tenuously appears around 2 Ma. In the Middle Pleistocene, the Lower Palaeolithic industries belong to two technological facies, the Acheulian and the Early Soanian. In the North-Western Siwaliks, the Soanian occurs in plenty on the river terraces, which were all built after 0.4 Ma, and the Acheulian appears between 0.5 and 0.7 Ma. The Middle Palaeolithic is still rich in cobble tools and therefore named Late Soanian. The Upper Palaeolithic is missing, perhaps due to climatic factors and the Neolithic is characterised by polished axes/adzes, but actually many of the cobble tools usually considered as Early Soanian may well belong to the cobble tool technical tradition common in South-East Asia during the Late Pleistocene and Early Holocene. In the north of the mountain range, the entire Palaeolithic sequence yielded by the loess in Southern Tajikistan is characterised, right from the late Lower Pleistocene, by lithic industries mainly compose of flakes, sometimes with cobble tools. In the high altitude zones, prehistoric sites are unknown before the Holocene and therefore they are contemporaneous with the Neolithic but some technological facies display particular features. The question is to know whether the originality of the industries, observed at different periods of time, is linked to techno-cultural adaptations to particular environments or to the isolation of the mountain populations.     

The Middle-to-Upper Palaeolithic transition in Cova Gran (Catalunya, Spain) and the extinction of Neanderthals in the Iberian Peninsula

The excavations carried out in Cova Gran de Santa Linya (Southeastern PrePyrenees, Catalunya, Spain) have unearthed a new archaeological sequence attributable to the Middle Palaeoloithic/Upper Palaeolithic (MP/UP) transition. This article presents data on the stratigraphy, archaeology, and ¹⁴C AMS dates of three Early Upper Palaeolithic and four Late Middle Palaeolithic levels excavated in Cova Gran. All these archaeological levels fall within the 34-32 ka time span, the temporal frame in which major events of Neanderthal extinction took place. The earliest Early Upper Palaeolithic (497D) and the latest Middle Palaeolithic (S1B) levels in Cova Gran are separated by a sterile gap and permit pinpointing the time period in which the Mousterian disappeared from Northeastern Spain. Technological differences between the Early Upper Palaeolithic and Late Middle Palaeolithic industries in Cova Gran support a cultural rupture between the two periods. A series of 12 ¹⁴C AMS dates prompts reflections on the validity of reconstructions based on radiocarbon data. Thus, results from excavations in Cova Gran lead us to discuss the scenarios relating the MP/UP transition in the Iberian Peninsula, a region considered a refuge of late Neanderthal populations.     

Land Snails as a Diet Diversification Proxy during the Early Upper Palaeolithic in Europe

Despite the ubiquity of terrestrial gastropods in the Late Pleistocene and Holocene archaeological record, it is still unknown when and how this type of invertebrate resource was incorporated into human diets. In this paper, we report the oldest evidence of land snail exploitation as a food resource in Europe dated to 31.3-26.9 ka yr cal BP from the recently discovered site of Cova de la Barriada (eastern Iberian Peninsula). Mono-specific accumulations of large Iberus alonensis land snails (Ferussac 1821) were found in three different archaeological levels in association with combustion structures, along with lithic and faunal assemblages. Using a new analytical protocol based on taphonomic, microX-Ray Diffractometer (DXR) and biometric analyses, we investigated the patterns of selection, consumption and accumulation of land snails at the site. The results display a strong mono-specific gathering of adult individuals, most of them older than 55 weeks, which were roasted in ambers of pine and juniper under 375°C. This case study uncovers new patterns of invertebrate exploitation during the Gravettian in southwestern Europe without known precedents in the Middle Palaeolithic nor the Aurignacian. In the Mediterranean context, such an early occurrence contrasts with the neighbouring areas of Morocco, France, Italy and the Balkans, where the systematic nutritional use of land snails appears approximately 10,000 years later during the Iberomaurisian and the Late Epigravettian. The appearance of this new subsistence activity in the eastern and southern regions of Spain was coeval to other demographically driven transformations in the archaeological record, suggesting different chronological patterns of resource intensification and diet broadening along the Upper Palaeolithic in the Mediterranean basin.     

Middle Pleistocene ecology and Neanderthal subsistence: Insights from stable isotope analyses in Payre (Ardèche,southeastern France)

The Middle Palaeolithic site of Payre in southeastern France yields abundant archaeological material associated with fossil hominid remains. With its long sequence of Middle Pleistocene deposits, Payre is a key site to study the Middle Palaeolithic chronology of this region. This study is the first to investigate carbon and oxygen isotope contents of Neanderthal tooth enamel bioapatite, together with a wide range of herbivorous and carnivorous species. The aim is to contribute to the understanding of hunting behaviour, resource partitioning, diet and habitat use of animals and Neanderthals through a palaeoecological reconstruction.     

Mandibular molar root morphology in Neanderthals and Late Pleistocene and recent Homo sapiens

Neanderthals have a distinctive suite of dental features, including large anterior crown and root dimensions and molars with enlarged pulp cavities. Yet, there is little known about variation in molar root morphology in Neanderthals and other recent and fossil members of Homo. Here, we provide the first comprehensive metric analysis of permanent mandibular molar root morphology in Middle and Late Pleistocene Homo neanderthalensis, and Late Pleistocene (Aterian) and recent Homo sapiens. We specifically address the question of whether root form can be used to distinguish between these groups and assess whether any variation in root form can be related to differences in tooth function. We apply a microtomographic imaging approach to visualise and quantify the external and internal dental morphologies of both isolated molars and molars embedded in the mandible (n = 127). Univariate and multivariate analyses reveal both similarities (root length and pulp volume) and differences (occurrence of pyramidal roots and dental tissue volume proportion) in molar root morphology among penecontemporaneous Neanderthals and Aterian H. sapiens. In contrast, the molars of recent H. sapiens are markedly smaller than both Pleistocene H. sapiens and Neanderthals, but share with the former the dentine volume reduction and a smaller root-to-crown volume compared with Neanderthals. Furthermore, we found the first molar to have the largest average root surface area in recent H. sapiens and Neanderthals, although in the latter the difference between M₁ and M₂ is small. In contrast, Aterian H. sapiens root surface areas peak at M₂. Since root surface area is linked to masticatory function, this suggests a distinct occlusal loading regime in Neanderthals compared with both recent and Pleistocene H. sapiens.     

Os fossiles humains des grottes Muierii et Cioclovina, Roumanie

This paper presents the cultural and archaeological context of the human fossil bones from Muierii Cave, dated by us to the age of 30 150 ± 800 ¹⁴C years BP (Before Present) or 34 810 ± 927 cal years BP (calibrated years Before Present), and from Cioclovina Cave, dated to the age of 29 000 ± 700 ¹⁴C years BP or 33 540 ± 832 cal years BP, in the Southern Carpathians. These are among the most ancient dated human fossil remains from Central and South-Eastern Europe and are described in conjunction with other sites with Mousterian assemblages of the recent Neanderthal population, and sites with Aurignacian assemblage of early modern humans, from Romanian region, for the interval of time 34,000-26,000, the transitional period from the Middle Paleolithic to the Upper Paleolithic.     

Hominin Dispersal into the Nefud Desert and Middle Palaeolithic Settlement along the Jubbah Palaeolake,Northern Arabia

The Arabian Peninsula is a key region for understanding hominin dispersals and the effect of climate change on prehistoric demography, although little information on these topics is presently available owing to the poor preservation of archaeological sites in this desert environment. Here, we describe the discovery of three stratified and buried archaeological sites in the Nefud Desert, which includes the oldest dated occupation for the region. The stone tool assemblages are identified as a Middle Palaeolithic industry that includes Levallois manufacturing methods and the production of tools on flakes. Hominin occupations correspond with humid periods, particularly Marine Isotope Stages 7 and 5 of the Late Pleistocene. The Middle Palaeolithic occupations were situated along the Jubbah palaeolake-shores, in a grassland setting with some trees. Populations procured different raw materials across the lake region to manufacture stone tools, using the implements to process plants and animals. To reach the Jubbah palaeolake, Middle Palaeolithic populations travelled into the ameliorated Nefud Desert interior, possibly gaining access from multiple directions, either using routes from the north and west (the Levant and the Sinai), the north (the Mesopotamian plains and the Euphrates basin), or the east (the Persian Gulf). The Jubbah stone tool assemblages have their own suite of technological characters, but have types reminiscent of both African Middle Stone Age and Levantine Middle Palaeolithic industries. Comparative inter-regional analysis of core technology indicates morphological similarities with the Levantine Tabun C assemblage, associated with human fossils controversially identified as either Neanderthals or Homo sapiens.     

Pedogenic carbonate stable isotopic evidence for wooded habitat preference of early Pleistocene tool makers in the Turkana Basin

The origin and evolution of early Pleistocene hominin lithic technologies in Africa occurred within the context of savanna grassland ecosystems. The Nachukui Formation of the Turkana Basin in northern Kenya, containing Oldowan and Acheulean tool assemblages and fossil evidence for early members of Homo and Paranthropus, provides an extensive spatial and temporal paleosol record of early Pleistocene savanna flora. Here we present new carbon isotopic (δ¹³C_VPDB) values of pedogenic carbonates (68 nodules, 193 analyses) from the Nachukui Formation in order to characterize past vegetation structure and change through time. We compared three members (Kalochoro, Kaitio, and Natoo) at five locations spanning 2.4–1.4 Ma and sampled in proximity to hominin archaeological and paleontological sites. Our results indicate diverse habitats showing a mosaic pattern of vegetation cover at each location yet demonstrate grassland expansion through time influenced by paleogeography. Kalochoro floodplains occurred adjacent to large river systems, and paleosols show evidence of C₃ woodlands averaging 46–50% woody cover. Kaitio habitats were located along smaller rivers and lake margins. Paleosols yielded evidence for reduced portions of woody vegetation averaging 34–37% woody cover. Natoo environments had the highest percentage of grasslands averaging 21% woody cover near a diminishing Lake Turkana precursor. We also compared paleosol δ¹³C_VPDB values of lithic archaeological sites with paleosol δ¹³C_VPDB values of all environments available to hominins at 2.4–1.4 Ma in the Nachukui and Koobi Fora Formations. Grassy environments became more widespread during this interval; woody canopy cover mean percentages steadily decreased by 12%. However, significantly more wooded savanna habitats were present in the vicinity of lithic archaeological sites and did not mirror the basin-wide trend of grassland spread. Hominin lithic archaeological sites consistently demonstrated woody cover circa 40% throughout our study interval and were 4–12% more woody than coeval basin environs. We propose that Turkana Basin early tool makers may have preferred a more wooded portion of the savanna ecosystem to reduce heat stress and to gain differential access to potable water, raw materials, animal carcasses, and edible plants.     

Palaeoloxodon and Human Interaction: Depositional Setting,Chronology and Archaeology at the Middle Pleistocene Ficoncella Site (Tarquinia,Italy)

The Ficoncella site in northern Latium (Italy) represents a unique opportunity to investigate the modalities of a short occupation in an alluvial setting during the Lower Palaeolithic. The small excavation area yielded a lithic assemblage, a carcass of Palaeoloxodon antiquus, and some other faunal remains. The main objectives of the study are to better characterize the depositional context where the Palaeoloxodon and the lithic assemblage occur, and to evaluate with greater precision the occupation dynamics. A 25 m-long well was drilled just above the top of the terrace of the Ficoncella site and faunal and lithic remains were analyzed with current and innovative techniques. The archaeological site contains floodplain deposits as it is located next to a small incised valley that feeds into a larger valley of the Mignone River. A tephra layer capping the site is ⁴⁰Ar/³⁹Ar dated to 441± 8 ka. Collectively, the geochronologic, tephrochronologic and geologic data, suggest the site was occupied during MIS 13. The new results should prompt further research at Ficoncella in order to improve our understanding of the dynamics of human settlement in Europe during the Early to Middle Pleistocene.     

Rethinking the Hummalian industry

Recent research reveals that the production of elongated blanks using different flaking techniques is an important part of the Early Middle Palaeolithic (EMP) industries in Near Eastern sites dated between 250 and 160 ka ago. The excavation at Hummal located in the arid steppe of Central Syria produced blade industries located in the stratigraphy between the Yabrudian and Levalloiso-Mousterian and this sequence is dated to about 200 ky. This paper presents data on the Hummalian culture from its discovery in 1980 to the systematic excavation of in situ archaeological layers between 1997 and 2010. Today the Hummalian industry is seen as a single, but very complex reduction strategy related to both the Laminar and the Levallois-like system of debitage. It is a unique reduction system containing diverse types of core volume management within which blanks of different morphologies have been produced from a single core using a direct, hammer percussion. Though, the Hummalian still shares many techno-typological similarities with the others laminar, lithic assemblages found on the Early Middle Palaeolithic sites in Levant, the same chronological and stratigraphical position and similar land-use strategies. As the EMP blade industries in Levant are preceded by the Acheulo-Yabrudian techno-complexes, the shift between both these lithic complexes, already seen in their chronological boundary, may also imply a technological discontinuity and possibly differing human populations.     

Techno-Cultural Characterization of the MIS 5 (c. 105 – 90 Ka) Lithic Industries at Blombos Cave,Southern Cape,South Africa

Blombos Cave is well known as an important site for understanding the evolution of symbolically mediated behaviours among Homo sapiens during the Middle Stone Age, and during the Still Bay in particular. The lower part of the archaeological sequence (M3 phase) contains 12 layers dating to MIS 5 with ages ranging from 105 to 90 ka ago (MIS 5c to 5b) that provide new perspectives on the technological behaviour of these early humans. The new data obtained from our extensive technological analysis of the lithic material enriches our currently limited knowledge of this time period in the Cape region. By comparing our results with previously described lithic assemblages from sites south of the Orange River, we draw new insights on the extent of the techno-cultural ties between these sites and the M3 phase at Blombos Cave and highlight the importance of this phase within the Middle Stone Age cultural stratigraphy.     

New evidence for a 67,000-year-old human presence at Callao Cave, Luzon, Philippines

Documentation of early human migrations through Island Southeast Asia and Wallacea en route to Australia has always been problematic due to a lack of well-dated human skeletal remains. The best known modern humans are from Niah Cave in Borneo (40-42 ka), and from Tabon Cave on the island of Palawan, southwest Philippines (47 ± 11 ka). The discovery of Homo floresiensis on the island of Flores in eastern Indonesia has also highlighted the possibilities of identifying new hominin species on islands in the region. Here, we report the discovery of a human third metatarsal from Callao Cave in northern Luzon. Direct dating of the specimen using U-series ablation has provided a minimum age estimate of 66.7 ± 1 ka, making it the oldest known human fossil in the Philippines. Its morphological features, as well as size and shape characteristics, indicate that the Callao metatarsal definitely belongs to the genus Homo. Morphometric analysis of the Callao metatarsal indicates that it has a gracile structure, close to that observed in other small-bodied Homo sapiens. Interestingly, the Callao metatarsal also falls within the morphological and size ranges of Homo habilis and H. floresiensis. Identifying whether the metatarsal represents the earliest record of H. sapiens so far recorded anywhere east of Wallace’s Line requires further archaeological research, but its presence on the isolated island of Luzon over 65,000 years ago further demonstrates the abilities of humans to make open ocean crossings in the Late Pleistocene.     

Aeretesmelanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China

The materials of Aeretesmelanopterus from the Tianyuan Cave, a Late Pleistocene site with human fossils recovered recently near Zhoukoudian, are the richest of this species ever reported in North China. Aeretesmelanopterus is an endemic species of China, its relationship with Petaurista and Trogopterus is still controversal. Studies based on the dental characters show that Aeretesmelanopterus resembles Petaurista more closely than Trogopterus. The earliest fossil record of Aeretesmelanopterus was found in South China in Middle Pleistocene deposits. The earliest records in the Beijing area are from Upper Cave and Tianyuan Cave at Zhoukoudian are of late-Late Pleistocene age, around 30 kyrs BP. The geographical distributions of this species are very limited. In evolution, Aeretesmelanopterus experienced both the increase and the decrease in tooth size; but its crown height of cheek teeth was continuously increasing.     

The fragmented character of Middle Palaeolithic stone tool technology

The importance of the transport of stone artefacts in structuring Neandertal lithic assemblages has often been addressed, but the degree to which this led to fragmentation of lithic reduction over Middle Palaeolithic landscapes has not been explicitly studied thus far. Large-scale excavations of Middle Palaeolithic open-air sites and refitting studies of the retrieved assemblages have yielded new, high-resolution data on the mobile aspects of Neandertal stone tool technology. In this paper, we integrate lithic technology and raw material data from recent studies of Middle Palaeolithic open-air and rock shelter sites in Western Europe. We demonstrate that the results of a variety of typological, technological (especially refitting), and lithological studies have important consequences for our knowledge of the acquisition of raw materials and subsequent production, usage and discard of stone artefacts in the Middle Palaeolithic. Neandertal production and use of stone tools was fragmented in three domains: the spatial, the temporal and the social domain. We show that this versatile segmentation of stone artefact handling strategies is a main determinant of the character of the Neandertal archaeological record. Our data testify to ubiquitous and continuous transport of stone artefacts of a wide variety of forms, picked by Neandertals using selection criteria that were sometimes far removed from what archaeologists have traditionally considered, and to some degree still consider, to be desired end products of knapping activities. The data presented here testify to the variability and versatility of Middle Palaeolithic stone tool technology, whose fragmented character created very heterogeneous archaeological assemblages, usually the product of a wide variety of independent import, use, discard and/or subsequent transport events.