Impact of land use changes on mountain vegetation

Abstract. In this study the impact of land use changes on vegetation in the sub‐alpine‐alpine belt is analysed. The study sites (4.7 km²) are located in the Passeier Valley (South Tyrol, Italy), at an elevation of 1500–2300 m a.s.l. The whole study area was used for hay‐making ca. 60 yr ago. Today, part of the meadows are more intensively used, while other parts have been converted to pasture or have been abandoned. We analysed the reasons for these land use changes and the effects on vegetation with a Geographical Information System and geostatistical analysis. The result of these analyses are: (1) Current land use is mainly controlled by the degree of accessibility for vehicles. Accessible areas are being used more and more intensively, while poorly accessible areas are being abandoned or used as pasture. (2) Current vegetation is highly determined by current land use. Particular vegetation units can be assigned to each form of land use. (3) Succession starts immediately after abandonment. Depending on altitude, succession proceeds at different speeds and with different numbers of stages. Hence the type of vegetation indicates the time passed since abandonment. (4) Land use changes lead to characteristic changes in vegetation; they are considered to be the most important driving force for vegetation change. (5) Measures of intensification and abandonment of extensively used areas both lead to a decrease in the number of species.     

The Circumpolar Arctic vegetation map

Abstract. Question: What are the major vegetation units in the Arctic, what is their composition, and how are they distributed among major bioclimate subzones and countries? Location: The Arctic tundra region, north of the tree line. Methods: A photo‐interpretive approach was used to delineate the vegetation onto an Advanced Very High Resolution Radiometer (AVHRR) base image. Mapping experts within nine Arctic regions prepared draft maps using geographic information technology (ArcInfo) of their portion of the Arctic, and these were later synthesized to make the final map. Area analysis of the map was done according to bioclimate subzones, and country. The integrated mapping procedures resulted in other maps of vegetation, topography, soils, landscapes, lake cover, substrate pH, and above‐ground biomass. Results: The final map was published at 1:7 500 000 scale map. Within the Arctic (total area = 7.11 × 106 km²), about 5.05 × 10⁶ km² is vegetated. The remainder is ice covered. The map legend generally portrays the zonal vegetation within each map polygon. About 26% of the vegetated area is erect shrublands, 18% peaty graminoid tundras, 13% mountain complexes, 12% barrens, 11% mineral graminoid tundras, 11% prostrate‐shrub tundras, and 7% wetlands. Canada has by far the most terrain in the High Arctic mostly associated with abundant barren types and prostrate dwarf‐shrub tundra, whereas Russia has the largest area in the Low Arctic, predominantly low‐shrub tundra. Conclusions: The CAVM is the first vegetation map of an entire global biome at a comparable resolution. The consistent treatment of the vegetation across the circumpolar Arctic, abundant ancillary material, and digital database should promote the application to numerous land‐use, and climate‐change applications and will make updating the map relatively easy.     

旅游干扰对云丘山景区内植被的影响

选择云丘山景区为研究区域,以该区域的主要植被为研究对象,采用样方法对旅游干扰对云丘山景区内植被的影响进行了研究,共设置了40个乔木样方,并利用TWINSPAN聚类分析以及旅游干扰程度（TDD）对所取样方进行分析。结果表明：TWINSPAN聚类分类将景区内的植物群落划分为5个群系,其中、群系Ⅱ中伴人植物的优势度明显高于其它。干扰程度分析表明,在景区的40个乔木样方中,只有4个样方基本没有受到干扰,有3个样方受到中度干扰,其余的33个样方均为轻度干扰。TWINSPAN聚类分析科学合理地对旅游活动作用下植被景观的类型特征进行了分析。旅游干扰程度（TDD）直观地反映出各个样方所在地植被被干扰的程度,该研究结果可为旅游管理者提供一定的理论依据。     

Effects of landslides on the mountain vegetation of Flores Island,Azores

Question: What are the consequences of frequently occurring landslides on vegetation dynamics, floristic and structural diversity? Location: 39°27′N; 31°13′W – Morro Alto, Flores Island, Azores, Portugal. Methods: Six comparable landslides were selected. Plots were placed at the top, slope and toe of landslides. Data on floristic composition and biovolume, demography and size structure of the dominant tree species (Juniperus brevifolia) were collected. Hierarchical agglomerative clustering and Principal Component Analysis were used in order to identify succession stages and compare succession pathways and vegetation recovery in different parts of the landslides. Results: Four stages of primary succession on substrates formed by landslides were identified: pioneer (Festuca‐Sphagnum grassland), assembly (Juniperus‐Festuca‐Sphagnum open scrub), building (Juniperus‐Sphagnum scrub) and mature (Juniperus‐Sphagnum woodland). Concerning J. brevifolia populations, the succession pathways are independent of location on the landslide. However, at the floristic level, there are some differences, mainly in the pioneer stage at the toes of landslides. Better abiotic conditions, resulting in a higher succession rate, are probably responsible for a faster vegetation recovery on landslide toes. Conclusion: Landslides trigger succession processes that enable a massive regeneration of the dominant tree species and existence of species not present in mature forests. They are also responsible for the simultaneous occurrence of vegetation of different structures. Overall, landslides increase the floristic and structural diversity of the vegetation, consequently increasing landscape heterogeneity.     

Oligotrophic spring vegetation in Spanish mountain ranges

Montio-Cardaminetea syntaxa occurring in the siliceous mountains of the Iberian Peninsula are reviewed. At the highest altitudes three associations are recognized in Spanish mountain ranges: one in the Pyrenees,Montio-Bryetum schleicheri (alpine-subalpine); another in the Ibero-Atlantic ranges,Stellario alsines-Saxifragetum alpigenae (oromediterranean); and a third in the Sierra Nevada,Sedo melanantheri-Saxifragetum alpigenae (oromediterranean). At slightly lower altitudes two other associations have been identified:Myosotidetum stoloniferae (supramediterranean), located in the western ranges; andSaxifragetum aquaticae (subalpine), located in the central and eastern Pyrenees. Syntaxonomically the Pyrenean associations are classified in theCardamino-Montion alliance, the Ibero-Atlantic associations in theMyosotidion stoloniferae alliance, and the Sierra Nevada association in a hypothetical Nevadan-North African alliance. A western European framework is suggested for the Spanish mountain associations which mainly show an E-W distribution pattern.     

A vegetation map of The Netherlands,based on the relationship between ecotopes and types of potential natural vegetation

Summary The method of mapping the vegetation on scale 1: 200,000 and the starting points in relation to the potential natural vegetation and ecotopes, are discussed.In view of the planological background of this study, some restrictions have been added to the concept of potential natural vegetation, concerning the period of development and the human influence.The relationship between soil, ground water and vegetation was studied, which resulted in the map of the potential natural vegetation.Each type of potential natural vegetation stands for a series of vegetation types on the same site. Seven main series, with a number of sub-series are distinguished. Within each vegetation series the plant communities have been spread over five groups, according to their structure and naturalness.Ecotopes and ecotope complexes are considered as landscape ecological units. A list of ecotopes was obtained by interpreting topographical maps and by inventory data.The actual vegetation was mapped by estimating the size of the ecotopes within the separate areas. It was expressed in a five figure code for the five groups from the vegetation and ecotopes is combined into the vegetation map of The Netherlands.Interpretation problems, some of them specific for The Netherlands, are discussed and some remarks are made on the necessity of further research.Contribution to the Symposium on Plant Species and Plant Communities, held at Nijmegen, 11–12 November 1976, on the occasion of the 60th birthday of Professor Victor Westhoff.Nomenclature follows Heukels-van Ooststroom, Flora van Nederland, 18e druk, 1975, Wolters-Noordhoff, Groningen; nomenclature of syntaxa follows Westhoff & den Held (1969)     

Gradients in field-layer vegetation on an arid misty mountain plateau in the Sudan

Abstract. Earlier studies have described how moist, on-shore winds cause meso-scale vegetation patterns on arid mountains near the sea. However, all protruding objects such as trees, micro-relief, and hill slopes influence the distribution of sea-mist. The influence of the tree-canopy, aspect, and distance to the sea on the field-layer vegetation in montane savanna was investigated on 16 hills in the Red Sea Hills, at 34 - 38 km from the sea. At 32 sites, total field-layer cover, species cover, and species number were estimated in a sub-canopy plot and in a nearby open plot on seaward and leeward slopes. Cover and species number in the understorey are significantly higher than in the open. The difference is highest on seaward slopes. Detrended correspondence analysis reveals short species-axes of ca. 2 SD-units. Differences between plots are mainly in species cover. This fits a principal components ordination model. PCA and its constrained version RDA give concordant results. The explanatory variables, Tree-cover and Relative Radiation Index (aspect), have similar indirect influences on plants, and are significantly correlated with axis 1, which is interpreted as a moisture and temperature gradient. The moist seaward plots show an independent trend in species composition along axis 2, which correlates with distance to the sea. On a presence basis the variables, all representing different spatial separation, correlate on the first axis. Presumably, the species composition, at all spatial scales, is directly or indirectly related to the variation in temperature and moisture.     

Geomorphic predictors of riparian vegetation in small mountain watersheds

Aims Hydrogeomorphic processes operating at watershed, process zone and site scales influence the distribution of riparian vegetation. However, most studies examining the relationships between hydrogeomorphic processes and riparian vegetation are conducted at site scales. We quantified the relative importance of watershed, process zone and site geomorphic characteristics for predicting riparian plant community types and plant species abundances in four small mountain watersheds in central Nevada, USA.Methods We mapped riparian vegetation types and identified process zones (based on dominant geomorphic process and valley fill material) within the watersheds. We sampled sites in each combination of vegetation type and process zone (n = 184 sites) and collected data on watershed scale factors, valley and stream geomorphic characteristics and on plant cover of each geomorphic surface. Plant community types were defined by cluster and indicator species analyses of plant cover data, and related to geomorphic variables using ordination analysis (nonmetric multidimensional scaling). Linear mixed effects models were used to predict abundances of indicator species.Important findings Variables describing position in the watershed (elevation, contributing area) that are related to gradients of temperature, moisture and stream discharge were of primary importance in predicting plant community types. Variables describing local geomorphic setting (valley width, stream gradient, channel sediments, geomorphic surface height) were of secondary importance, but accurately described the geomorphic setting of indicator species. The process zone classification did not include position in the watershed or channel characteristics and only predicted plant community types with unique geomorphic settings. In small mountain watersheds, predicting riparian vegetation distribution requires explicit consideration of scale and geomorphic context within and among watersheds in addition to site variables.     

Impact of climate change on alpine vegetation of mountain summits in Norway

Climate change is affecting the composition and functioning of ecosystems across the globe. Mountain ecosystems are particularly sensitive to climate warming since their biota is generally limited by low temperatures. Cryptogams such as lichens and bryophytes are important for the biodiversity and functioning of these ecosystems, but have not often been incorporated in vegetation resurvey studies. Hence, we lack a good understanding of how vascular plants, lichens and bryophytes respond interactively to climate warming in alpine communities. Here we quantified long-term changes in species richness, cover, composition and thermophilization (i.e. the increasing dominance of warm-adapted species) of vascular plants, lichens and bryophytes on four summits at Dovrefjell, Norway. These summits are situated along an elevational gradient from the low alpine to high alpine zone and were surveyed for all species in 2001, 2008 and 2015. During the 15-year period, a decline in lichen richness and increase in bryophyte richness was detected, whereas no change in vascular plant richness was found. Dwarf-shrub abundance progressively increased at the expense of lichens, and thermophilization was most pronounced for vascular plants, but occurred only on the lowest summits and northern aspects. Lichens showed less thermophilization and, for the bryophytes, no significant thermophilization was found. Although recent climate change may have primarily caused the observed changes in vegetation, combined effects with non-climatic factors (e.g. grazing and trampling) are likely important as well. At a larger scale, alpine vegetation shifts could have a profound impact on biosphere functioning with feedbacks to the global climate.     

A simulated map of the potential natural forest vegetation of Switzerland

Using empirical data (ca. 7500 phytosociological releves), a simple, probabilistic ‘vegetation-site’ model was developed, to simulate geographical distribution of 71 forest community types, representing the potential natural vegetation (PNV) of Switzerland. The model was interfaced to a geographic information system (GIS) and used to generate a numerical vegetation map, on the basis of digital maps of 12 environmental variables including climatic conditions (temperature and precipitation), topography (elevation, slope, aspect), and soil parameters (soil pH and physical soil parameters). The predicted distribution of forest communities was compared with several vegetation maps, prepared for some subregions of Switzerland by means of traditional field methods. Similarity ranged from 50 to 80 %, depending on the community type, level of vegetational hierarchy and the geographical region. The current resolution and accuracy of the simulated vegetation map allows us to study the vegetational patterns on the level of the entire country or its major geographical and climatic regions. The simulated vegetation map is potentially an important tool in ecological risk assessment studies concerning the possible impacts of climate change on the ecological potential of forest sites and biological diversity of forest communities.     

A pilot study on pollen representation of mountain valley vegetation in the central Alps

The relative pollen productivity (RPP) of Pinus cembra, Larix, Picea, Gramineae, and Cyperaceae was estimated for the Upper Engadine area, an inner-alpine sub-continental valley in SE Switzerland lying above 1800 m a.s.l. The influx of pollen originating from lowlands was assumed constant between the investigated sites and could thus be used to establish relative pollen accumulation rates (rPAR) as a basis for further calculations. Wind above the canopy was assumed as the major mode of pollen transport and the effect of different wind speeds was tested. In order to evaluate the results pollen percentage values were simulated and compared with the empirical data. The method is sufficiently accurate for evaluation of spatially-explicit vegetation models, although the low number of sites eludes robust statistical testing.     

Running off the road: roadside non-native plants invading mountain vegetation

Prevention is regarded as a cost-effective management action to avoid unwanted impacts of non-native species. However, targeted prevention can be difficult if little is known about the traits of successfully invading non-native species or habitat characteristics that make native vegetation more resistant to invasion. Here, we surveyed mountain roads in seven regions worldwide, to investigate whether different species traits are beneficial during primary invasion (i.e. spread of non-native species along roadside dispersal corridors) and secondary invasion (i.e. percolation from roadsides into natural adjacent vegetation), and to determine if particular habitat characteristics increase biotic resistance to invasion. We found primary invasion up mountain roads tends to be by longer lived, non-ruderal species without seed dispersal traits. For secondary invasion, we demonstrate that both traits of the non-native species and attributes of the receiving natural vegetation contribute to the extent of invasion. Non-native species that invade natural adjacent vegetation tend to be shade and moisture tolerant. Furthermore, non-native species invasion was greater when the receiving vegetation was similarly rich in native species. Our results show how mountain roads define which non-native species are successful; first by favouring certain traits in mountain roadsides (the key dispersal pathway to the top), and secondly by requiring a different set of traits when species invade the natural adjacent vegetation. While patterns in species traits were observed at a global level, regional abiotic and biotic variables largely generated region-specific levels of response, suggesting that management should be regionally driven.     

Complex vegetation responses to soil disturbances in mountain grassland

We studied vegetation responses to disturbances originated by ants and voles in subalpine grasslands in the Eastern Pyrenees. We compared the effects of these small-scale disturbances with those of a large-scale disturbance caused by ploughing. We wanted to know if these soil disturbances promoted species richness through the existence of a specific guild of plants colonizing these areas, and if this guild was the same for all soil disturbances, independently of their extent. In general, grassland vegetation seemed to recover relatively quickly from soil-displacement disturbances, and the effects could be scaled up in time and space in terms of species richness and composition. Vole mound composition was similar to that in the surrounding grassland, suggesting that mounds were rapidly colonized by the neighbouring vegetation. Vegetation composition differed between the grassland and the ant mounds. Grasses and erect dicots coped well with repeated disturbance, while rosette-forming species and sedges were very sensitive to it. Landscape processes could be important to understanding recolonization. Species from xeric grasslands were found in mesic grasslands when disturbed by ploughing and on the tops of active ant mounds. Furrows in mesic grasslands recovered well, but decades after disturbance showed long persistence of some xeric species and increased species richness compared to terraces, while xeric grasslands showed decreased richness. This suggests that, because of those disturbances, within-habitat diversity was increased, although landscape diversity was not. However, specific disturbances showed idiosyncratic effects, which could enhance the species richness globally. In ant-affected areas, the grassland itself showed the highest plant species richness, partially associated to the presence of some species with elaiosomes not, or only rarely, found in adjacent grasslands without ant mounds. Therefore, soil disturbances occurring at different spatial scales contributed to complexity in vegetation patterns in addition to abiotic factors and grazing. Electronic supplementary material  The online version of this article (doi:) contains supplementary material, which is available to authorized users. Nomenclature of the species follows Tutin et al. (1964–1980) and Bolòs et al. (1993).     

Lateglacial and Holocene vegetation history in the mountain range of central Pyrenees

We present the pollen analysis of a new sedimentary sequence taken at La Pouretère ( 1720 m), in the mountain vegetation zone of the Marcadau valley (central Pyrenees). The Lateglacial and Holocene chronology is supported by six 14C-dating results. The complementary analysis of some vegetal macroremains, stomata, pollen-clusters and the use of pollen influx allows us to elucidate the dynamic of mountain species such as Pinus and specially Abies but also to infer the unusual part played by Betula at the beginning of the Postglacial period.     

Topographic control of vegetation in a mountain big sagebrush steppe

Mountain big sagebrush steppes in Wyoming have strong spatial patterning associated with topography. We describe the spatial variability of vegetation in a sagebrush steppe, and test the relationship between topography and vegetation using canonical correlation. Results of the analysis suggest that the main control over vegetation distribution in this system is wind exposure. Exposed sites are characterized by cushion plant communities and Artemisia nova, and less exposed sites by the taller sagebrush species Artemisia tridentata ssp. vaseyana. Topographic depressions and leeward slopes are characterized by aspen stands and nivation hollows. Measurements of soil microclimate suggest that a major influence of topographic position on vegetation is snow redistribution and its effect on soil moisture and temperature.Abbreviations ARNO Artemisia nova - ARTRW Artemisia tridentata ssp. wyomingensis - ARTRV Artemisia tridentata ssp. vaseyana - PUTR Purshia tridentata - RIP riparian community - POTR Populus tremuloides - NIV nivation hollow community