Environmental change in the Colombian subandean forest belt from 8 pollen records: the last 50 kyr

We present a reconstruction of forest history and climatic change based on 11 pollen records from eight sites, all located in the lower montane forest belt of the northern Andes in Colombia. We compared records from the Popayán area in southern Colobia, Timbio (1750 m), Genagra (1750 m) and Pitalito (1300 m) and the new Piagua (1700 m) record with the records from Lusitania (1500 m), Libano (1820 m), Pedro Palo (2000 m) and Ubaqué (2000 m) from Central Colombia. The changes of the altitudinal position of the lower/upper montane (= subandean/Andean, S/At) forest belt transition were used to estimate temperature change for the last 50 kyr. We infer a Last Glacial Maximum (LGM) temperature drop of 6°–7°C at 1700 m, and a steeper LGM lapse rate of 0.76°C/100 m compared to today (ca. 0.6°C/100 m). Around 50 uncal. kyr B.P. the temperature at 1700 m was ca. 3°C lower than today. Until 20 uncal. kyr B.P. the temperature oscillated and gradually decreased. During the LGM, temperature was down to ca. 6°–7°C lower than today. After the LGM, temperature increased and ca. 14 uncal. kyr B.P. it was 2°–3°C lower than today (S/At at ca. 1800 m, 500 m below present elevation; Susacá interstadial). An unquantified cooling (Ciega stadial) followed. During ca. 12.3–11.7 uncal. kyr B.P. the S/At shifted upslope to 2100 m indicating a temperature of 1°–2°C cooler than today (Guantiva interstadial). From 11.7–10.9 uncal. kyr B.P. the S/At was at 1800 m indicating that the temperature was ca. 3°C lower than today and wet conditions prevailed (partly coinciding with the El Abra stadial). The period 10.9–9 uncal. kyr B.P. was also cool, but drier. During 9–7.5 uncal. kyr B.P. temperature was ca. 1°C warmer relative to today (mid Holocene hypsithermal). During the last 5 kyr the presence of cultivated plants demonstates human colonization of the lower montane zone in Colombia. Received June 14, 2000 / Accepted December 19, 2000     

Vegetation in the mountains of northwest Iberia during the last glacial-interglacial transition

A new core taken from a site in the Sierra de Courel range of mountains in the northwest Iberian peninsula, Spain, enables a palaeoenvironmental reconstruction to be made of the biological events resulting from climatic changes and human activities during the last 17,000 years in the area. The sequence begins with a phase characterised by the dominance of Gramineae (Poaceae) and Pinus. A markedly arid period with prevalence of Artemisia pollen occurred between 17,5000 and 15,500 uncal. B.P. Subsequently, a succession of woodlands with Betula, Pinus and other mesophilous and thermophilous trees was recognised during the period 15,500-13,500 uncal. B.P. Between 13,500 and 10,300 uncal. B.P. herbaceous vegetation formations indicating moister conditions dominated the pollen record, although a maximum of Gramineae has been dated during the period 11,300-10,300 uncal. B.P. Comparison with other pollen data from neighbouring mountains allows an interpretation of the vegetation changes during the glacial/interglacial transition in the mountains of northwest Iberia. The Holocene pollen record from the site does not differ markedly from other records in the area, tree expansion occurring before 10,000 uncal. B.P. and high values of deciduous Quercus mixed with other trees and shrubs persisting until 3500 uncal. B.P., when increased human activities are shown by a fall of the tree pollen percentages. Received July 7, 1999 / Accepted May 22, 2000     

Surface-bottom relationships in the Gulf of Salerno (Tyrrhenian Sea) over the last 34 kyr: Compositional data analysis of palaeontological proxies and geochemical evidence

The palaeontological, geochemical and mineralogical records of core GNS84-C106 were analysed in order to reconstruct palaeohydrological changes and palaeoproductivity patterns in the Gulf of Salerno for the last 34 kyr. This approach, including compositional analysis of planktonic and benthic assemblages, gave an insight into the relationships between continental, sea surface and bottom environmental changes. The main source of variability of planktonic and benthic assemblages is related respectively to sea surface temperature and palaeobathymetry. Interrelated changes in surface salinity, nutrients, density gradient in the water column and organic fluxes at the bottom act as a secondary factor controlling the composition of both planktonic and benthic assemblages. The highest palaeoproductivity rates were reached during an interval spanning from late glacial to Middle Holocene, in conditions of enhanced continental run-off. During the Early and Middle Holocene, reduced surface salinity and density stratification were also coupled with the development of a deep chlorophyll maximum and enhanced flux or organic matter at the bottom. From about 6.5 kyr B.P. onward, a sharp reduction in palaeoproductivity took place, coupled with an increase in surface salinities.     

The long history of Cannabis and its cultivation by the Romans in central Italy, shown by pollen records from Lago Albano and Lago di Nemi

The cores from the Albano and Nemi lakes, near Rome, were studied within the European Union funded PALICLAS project and provided high resolution records of the Late-glacial and Holocene. Pollen evidence of increasing human influence on vegetation was recorded in the Holocene parts of both diagrams, and the Cannabis (hemp) curve was one of the major signs. In this paper we present unambiguous pollen evidence from the Cannabaceae records for the cultivation of hemp in central Italy by the Romans. The oldest records of Cannabis and Humulus (hop) date from to the Late-glacial. Hop pollen values rise during the mid Holocene, while hemp pollen becomes more abundant from ca. 3000 cal B.P. onwards. The highest earliest hemp peak (21%) is dated to the 1st century A.D. This ‘Cannabis phase’, with the abrupt rise of hemp pollen soon after the rise of cultivated trees (Castanea, Juglans and Olea) is associated with the increase in cereals and ruderal plants. This unambiguous proof of cultivation by Romans around 2000 B.P. occurs as well as a long lasting pre-Roman presence of hemp in the area, which is natural and possibly also anthropogenic. Subsequent clear episodes of cultivation in the medieval period were found. Received February 4, 2002 / Accepted September 13, 2002 Correspondence to: Anna Maria Mercuri, e-mail: mercuri.annamaria@unimo.it     

Deep-sea ecosystem variability of the Aegean Sea during the past 22 kyr as revealed by Benthic Foraminifera

Specific responses of the regional deep-sea ecosystems to climatic and oceanographic processes during the last 22 kyr are revealed by benthic foraminiferal faunas from two cores in the northern and southern Aegean Sea. Under glacial boundary conditions, high-diversity benthic foraminiferal faunas and elevated benthic foraminiferal numbers indicate enhanced organic matter availability and well-ventilated deep-water masses in the whole Aegean Sea. The glacial termination is accompanied by significant fluctuations in productivity and deep-water ventilation. In the northern Aegean Sea, meltwater inflow from mountain glaciers during the Bølling/Allerød warm period resulted in a restriction of local deep-water formation, as mirrored by a dominance of bolivinids. During the deposition of sapropel S1, drops in benthic foraminiferal number and diversity are more significant in the southern Aegean Sea when compared to the north. This suggests the persistence of local deep-water formation in the northern Aegean Sea during S1 deposition. In addition, faunal fluctuations within S1 at both sites suggest the repeated influence of short-term cooling events on the re-ventilation and re-colonization of Aegean deep-sea ecosystems. During the middle and late Holocene, benthic foraminiferal faunas document the establishment of oligotrophic and well-ventilated conditions in the southern Aegean Sea. The corresponding faunas from the northern Aegean Sea reflect generally mesotrophic conditions and variable deep-water oxygenation. During the entire Holocene, the deep-sea ecosystems of this region responded very sensitively to short-term changes in humidity and temperature. These abrupt climate changes controlled the inflow of nutrients from rivers and the Black Sea and the formation of local deep-water masses.     

On the Late Pleistocene and Holocene history of vegetation and human impact in the Ücker valley, north-eastern Germany

A high-resolution Late Pleistocene and Middle to Late Holocene pollen profile of regional significance is presented. The coring site is located in a mire between two lakes. Ober- and Unter-ückersee, in Brandenburg, north-eastern Germany. The study was carried out in an archaeological context. It provides information about the history of vegetation, climate and human impact in the ücker river valley and the surrounding hills, the Uckerm?rker Hügelland. Hence, it is an important contribution for the reconstruction of the past vegetation of this area of Brandenburg. Seven AMS ¹⁴C-dates based on pollen concentrates provide a chronology for the middle Holocene part of the profile. Phases of intensive human activity can be shown from the middle Neolithic times until the Roman Iron Age. Received February 1, 2000 / Accepted January 18, 2001     

Mid-Holocene environmental and human dynamics in northeastern China reconstructed from pollen and archaeological data

A pollen record from the Taishizhuang site (40°21.5′N, 115°49.5′E) located in the transitional forest-steppe zone near the present-day limit of the summer monsoon is used to reconstruct vegetation and climate. Quantitative biome reconstruction suggests that between ca. 5700 and 4400 cal. years B.P. temperate deciduous forest dominated the vegetation cover around the Taishizhuang site. After that time the landscape became more open and the scores of the steppe biome were always higher than those of the temperate deciduous forest except for two oscillations dated to ca. 4000 cal. years B.P. and ca. 3500 cal. years B.P. However, ca. 3400-2100 cal. years B.P. the common vegetation became steppe and the landscape was more open in comparison with the previous time interval. The results of the pollen-based precipitation reconstruction suggest that annual precipitation was ca. 550-750 mm (ca. 100-300 mm higher than present) during the mid-Holocene ‘forest phase’, and ca. 450-650 mm during the following ‘forest-steppe phase’. From ca. 3400 cal. years B.P. during the ‘steppe phase’ annual precipitation was similar to modern values (ca. 300-500 mm). Archaeological records from 100 sites prove the habitation of northeastern China during the prehistoric and early historic periods from ca. 8200 cal. years B.P., but do not provide evidence of the use of wood resources intensive enough to influence the regional vegetation development and to leave traces in the pollen assemblages. Both archaeological and palaeoenvironmental data support the conclusion that changes in pollen composition in northeastern China between 5700 and 2100 cal. years B.P. reflect natural variations in precipitation and not major deforestation caused by humans.     

Vegetation history of the SE section of the Zagros Mountains during the last five millennia; a pollen record from the Maharlou Lake, Fars Province, Iran

A pollen diagram was derived from a 150 cm core taken from the shallow hypersaline Lake Maharlou in the south-eastern part of the Zagros Mountains, SW Iran. The pollen record shows that Quercus brantii woodland and Pistacia–Amygdalus scrub dominated the area during the late Holocene. The record starts at around 5700 cal b.p. with a dry period during which both Pistacia–Amygdalus scrub and Quercus brantii woodland were at their minimum extent. This period was followed by the expansion of Pistacia–Amygdalus scrub in the area and the spread of Quercus brantii woodlands at higher altitudes. An important occupation phase, characterized by the appearance of several cultivated tree species such as Juglans, Olea, Vitis and Platanus, started at ca. 4300 cal b.p., coinciding with the onset of the Bronze Age civilization of Jiroft in Central Iran. Human activities become very clear after 3700 cal b.p. Around 2700 cal b.p., extensive stands of Pistacia–Amygdalus scrub became profoundly degraded, presumably under strong human pressure coinciding with the beginning of the Persian Empires. The maximum expansion of the Quercus brantii woodland occurred about 2100 to 1700 cal b.p. This woodland remained relatively stable until the end of the diagram at 400 cal b.p.     

Vegetation history and lake-level changes from the Younger Dryas to the present in Eastern Pyrenees (France): pollen, plant macrofossils and lithostratigraphy from Lake Racou (2000 m a.s.l.)

A multi-proxy palaeoecological investigation including pollen, plant macrofossil, radiocarbon and sedimentological analyses, was performed on a small mountain lake in the Eastern Pyrenees. This has allowed the reconstruction of: (1) the vegetation history of the area based on five pollen diagrams and eight AMS¹⁴C dates and (2) the past lake-level changes, based on plant macrofossil, lithological and pollen analysis of two stratigraphical transects correlated by pollen analysis. The palaeolake may have appeared before the Younger Dryas; the lake-level was low and the vegetation dominated by cold steppic grasslands. The lake-level rose to its highest level during the Holocene in the Middle Atlantic (at ca. 5060±45 b.p.). Postglacial forests (Quercetum mixtum and Abieto-Fagetum) developed progressively in the lower part of the valley, while dense Pinus uncinata forests rapidly invaded the surroundings of the mire and remained the dominant local vegetation until present. The observed lowering of the lake levels during the Late Atlantic and the Subboreal (from 5060 ± B.P. to 3590±40 b.p.) was related to the overgrowth of the mire. The first obvious indications of anthropogenic disturbances of the vegetation are recorded at the Atlantic/Subboreal boundary as a reduction in the forest component, which has accelerated during the last two millennia.     

A high-resolution Late-Glacial – Holocene pollen diagram from Pian di Gembro (Central Alps, Northern Italy)

The local and regional history of vegetation and climate, from the Late Glacial to the present, is represented in a new, high-resolution pollen diagram from Pian di Gembro (1350 m asl), ten ¹⁴C dates providing a reliable time control. An open pioneer vegetation dominated by Artemisia, Gramineae, and and Chenopodiaceae followed the retreat of the glaciers after the Last Glacial Maximum. Shrub vegetation with Juniperus, Alnus viridis, and Salix expanded soon after. Denser Betula-Pinus forests were present in Pian di Gembro around 12,320 B.P. Their extent was greatly reduced by the climatic cooling of the Younger Dryas, when open vegetation spread again. The beginning of the Holocene was marked by a considerable expansion in mixed oak taxa. Corylus immigrated to the site at 9,250 B.P. Picea and Abies expanded at 7,370 B.P., recording an abrupt change in the structure of the vegetational belts. A coeval climatic change is evidenced in the GRIP records and also detectable through oscillations of the timberline. Signs of human impact are present since late Atlantic, becoming more intense around 2,200 B.P. As pasture lands increased, Abies and Fagus slowly disappeared. The introduction of Castanea and Juglans is dated to Roman times, and Secale to the Middle Ages. Received February 4, 2002 / Accepted August 2, 2002     

The role of human disturbance in Late Holocene vegetation changes on Kullaberg, southern Sweden

A pollen record from a small peatland located on Kullaberg in northwestern Sk?ne, southern Sweden, revealed that grazing pressure might have been the major disturbance agent controlling the vegetation, at least from ca. 1500 B. C. until the latter part of the nineteenth century. The pollen data also indicate a step-by-step increase in grazing pressure, expressed as a marked increase in non-arboreal pollen representation at ca. 1500 B. C., A. D. 650 and A. D. 1650. The increase at A. D. 650 probably indicates a more regular and intensive use of the area, resulting in a forest structure that was much more open than earlier, together with an expansion of Fagus, which rapidly replaced Quercus as the local forest dominant. At about this time the first patches of heath vegetation originated, but they probably only covered a small part of Kullaberg. Larger areas with an intensively grazed Calluna heath, as shown by eighteenth century maps, probably evolved around ca. A. D. 1650, when much of Kullaberg seems to have been deliberately deforested. Received February 20, 2001 / Accepted September 12, 2001     

Biostratigraphy and <Superscript>14</Superscript>C dating of a lake sediment sequence on the north-west Estonian carbonaceous plateau,interpreted in terms of human impact in the surroundings

The sediment stratigraphy of a medium-sized mixotrophic lake (Ruila) situated below the highest shoreline of the Baltic Ice Lake in the West-Estonian Lowland is described. The lake is without natural inlets our outlets. The reconstruction of vegetation and land-use history based on pollen data, combined with available archaeological data and detailed ¹⁴C dating allows us to give a provisional reconstruction of the temporal and spatial pattern of natural and human induced environmental changes in north-west Estonia during the Holocene. Both radiocarbon dates derived from terrestrial macrofossil dating by accelerator mass spectrometry (AMS) and conventional dating of bulk lake sediment are discussed. The isolation of the lake basin from the Yoldia Sea took place ca. 9700 cal B. C. The Ancylus Lake transgression at ca. 8400 cal B. C. did not reach the basin, but caused a ground water rise, seen in the sediment stratigraphy of the lake. The first signs of human impact on the pollen record appear ca. 5400 cal B. C. (Late Mesolithic). The history of arable farming has been divided into three periods: 1) introduction of crop cultivation and animal husbandry (1500 cal B. C. – A. D. 500); 2) establishment of animal husbandry A. D. 500–1000) and 3) establishment of crop cultivation and intensive cattle breeding (A. D. 1000–today). Due to unfavourable eda-phic conditions the introduction of arable farming was delayed for more than 1000 years compared with elsewhere on the north coast of Esotnia, and intensity of land-use never reached the same proportion as in these areas. Received August 15, 2001 / Accepted August 5, 2002 Correspondence to: Leili Saarse     

Mangroves of Oman during the late Holocene; climatic implications and impact on human settlements

Pollen and micropalaeontological analyses carried out on mangrove swamp sediments of Suwayh, Oman (22°05.589'N, 50°40.033'E) reveal environmental changes linked both to climate (monsoon) and geomorphological (sea-level) variations during the Late Holocene. A Rhizophora mangrove developed at Suwayh around 6000 years B. P. under climate conditions marked by an increased tropical influence as compared to the modern situation, with dominant summer rains. The later extension of Prosopis cineraria at Suwayh provides evidence for a different rainfall pattern, with a winter rainy season. Pollen and micropalaeontological composition shows three episodes influenced by the sea water at Suwayh. The most important corresponded to the mangrove episode centred ca 6000 B. P. This was followed by two episodes of slight seawater incursion at ca. 5100 and 4500 B. P. responsible for the formation of a brackish laagon. Comparison based on ¹⁴C measurements on shell recovered from both the sedimentary sequence of Suwayh and the nearby archaeological sites demonstrates that close relations existed between man and mangroves during the Neolithic and the Early Bronze Age. Received October 31, 2001 / Accepted January 24, 2002 Correspondence to: Anne-Marie Lézine     

Long-term human impact as registered in an upland pollen profile from the southern Black Forest, south-western Germany

A detailed,¹⁴C-dated, pollen profile from Steerenmoos, a raised bog in the uplands of the southern Black Forest (Schwarzwald) is presented. The Late-glacial and early Holocene vegetation history conforms to the known pattern of forest dynamics for that region. At ca. 6100 cal. B.P.,Abies replaced the mixed oak forest, which is in contrast to adjacent regions whereFagus spread beforeAbies. From the Subboreal onwards,Fagus expanded and slowly largely replacedAbies. The mire developed from a fen to a raised bog. The mountain pine (Pinus mugo ssp.rotundata) on the present-day bog surface is a result of medieval burning. Cereal pollen are first recorded in the Neolithic (7600 cal. B.P.) and there is a closed curve forPlantago Lanceolata — a good indicator of human impact — since the Bronze Age (4000 cal. B.P.). On the basis of the cereal pollen record nine human impact phases (HIP) are described. HIP 1 and 2, which are short, date to ca. 7600 and 6700 cal. B.P., respectively, in a mixed oak forest context and are characterized by declines inCorylus, Tilia, Ulmus and bySalix (but no major deforestation) and peaks in charcoal and loss-on-ignition curves. HIP 3 and 4, which are short and weak, date to ca. 6000 and 5300 cal. B.P., respectively, and occur in the context of anAbies alba forest. The Bronze Age and Iron Age HIPs 5-7 are more intense and of longer duration than the Neolithic phases and result in a decline inAbies and an increase inFagus. The early medieval HIP 8, although rather weak, probably finds expression also in an archaeological artefact, namely a dug-out boat from the near-by Schluchsee. Finally, the late Medieval HIP 9 resulted in a major transformation in the landscape. It is argued that the earlier HIPs are not a reflection of distant events in the lowland valleys of the Rhine, Danube or Neckar but reflect more or less local developments.     

Studies on the Holocene history of vegetation in the region of lake Srebarna (northeast Bulgaria)

The results of pollen analysis of two sediment cores from lake Srebarna (northeast Bulgaria) are presented. On the basis of the palynological data and the radiocarbon dates a reconstruction of the past flora and vegetation is made. For the first time a continuous palaeosuccession is established for the area of the Danubian Dobrudza and the Ludogorie district of northeast Bulgaria: a dominance of mixed xero- and mesophilous oak woods with Carpinus betulus, Ulmus, Tilia, Corylus during the Atlantic period; mixed oak woods with increasing importance of Tilia, Ulmus, Acer and a considerable presence of Carpinus betulus and Fagus during the Sub-boreal period; degradation of the woodland vegetation and transition to secondary plant communities with Carpinus orientalis and herbs during the Sub-atlantic period. Received January 5, 2001 / Accepted March 13, 2001     

Archaeobotany of the Old Prague Town defence system, Czech Republic: archaeology, macro-remains, pollen, and diatoms

The stone fortification of Prague Old Town was constructed shortly after A.D. 1230. Large-scale archaeological and archaeobotanical research has been carried out in this area in recent years. The deep moat and the drainage ditch have been excavated at several places in front of a line of former ramparts. During the initial period of their existence, the moat and the ditch contained only minimal waste. A sudden change in the management of these features occurred at the beginning of the 14th cent., when they started to be filled with large amounts of rubbish. This article concentrates on the evaluation of these rubbish strata at three profiles through the drainage ditch and one profile through the dry moat. The results of plant macro-remains (including wood), pollen, and diatom analyses clearly demonstrate the development of the local (aquatic) environment from being relatively clean to highly polluted. An actualistic (phytosociological) approach was adopted to reconstruct the local environment of the town periphery in detail, and to distinguish a variety of habitats that were situated in the economic hinterland of the city. Some species from the rich assemblages of plant remains are interpreted in terms of their use in High Medieval households. Diatom analysis completes the picture by providing the proxy data on water quality history in the drainage ditch. Received September 15, 2001 / Accepted May 21, 2002     

Paleoceanographic conditions in the western Caribbean Sea for the last 560 kyr as inferred from planktonic foraminifera

Faunal analyses of planktonic foraminifera and upper-water temperature reconstructions with the modern analog technique are studied and compared to the magnetic susceptibility and gamma ray logs of ODP Core 999A (western Caribbean) for the past 560 kyr in order to explore changes in paleoceanographic conditions in the western Caribbean Sea. Long-term trends in the percentage abundance of planktonic foraminifera in ODP Core 999A suggest two hydrographic scenarios: before and after 480 ka. High percentage abundances of Neogloboquadrina pachyderma and Globorotalia inflata, low abundances of Globorotalia menardii and Globorotalia truncatulinoides, low diversity, and sea-surface temperatures (SST) under 24 °C are typical characteristics occurring from 480 to 560 ka. These characteristics suggest a “shallow” well-oxygenated upper thermocline and the influx of nutrients by either seasonal upwelling plumes and/or eddy-mediated entrainment. The second scenario occurred after 480 ka, and it is characterized by high and fluctuating percentage abundances of Neogloboquadrina dutertrei, G. truncatulinoides, G. menardii, Globigerinita glutinata, Globigerinella siphonifera, and Globigerinoides ruber; a declining trend in diversity; and large SSTs. These characteristics suggest a steady change from conditions characterized by a “shallow” thermocline and chlorophyll maximum to conditions characterized by a “deep” thermocline (mainly during glacial stages) and by more oligotrophic conditions. The influence of the subtropical North Atlantic on the upper thermocline was apparently larger during glacial stages, thus favoring a deepening of the thermocline, an increase in sea-surface salinity, and a dramatic reduction of nutrients in the Guajira upwelling system. During interglacial stages, the influx of nutrients from the Magdalena River is stronger, thus resulting in a deep chlorophyll maximum and a fresher upper ocean. The eddy entrainment of nutrients is the probable mechanism responsible of transport from the Guajira upwelling and Magdalena River plumes into ODP 999A site.     

Towards the reconstruction of the Holocene vegetation history of Lower Provence: two new pollen profiles from Marais des Baux

Two new¹⁴C-dated Holocene pollen profiles from Marais des Baux, Bouches-du-Rhône, France, are presented. The record begins in the Younger Dryas, when the vegetation consisted mainly of grasses and mugwort (Artemisia). The Preboreal was marked by a transitory expansion of pine forests and was followed by the establishment of a rich deciduous oak-forest that included hazel (Corylus) and elm (Ulmus). During the Boreal, hazel played a dominant role within the oak-forest. The oak forests, which includedQuercus ilex, achieved a major expansion during the Atlantic period. The Subboreal was characterised by the regional establishment of, firstly, fir (Abies) and then beech (Fagus). The spread and expansion of beech coincides with the first clear evidence for farming. Agricultural activities brought about the decline of deciduous oak-forest. During the Subatlantic, forests in the vicinity of Marais des Baux were cleared for farming. Cereal growing, which included rye cultivation, was of considerable importance. Three noteworthy characteristics that serve to differentiate the Holocene vegetation history of the low-lying Provence region from other French regions are as follows: 1) the early establishment (from the onset of Preboreal) of low altitude mixed forest; 2) the expansion during the Subboreal of fir and beech in low altitude areas with a Mediterranean climate and, 3) the exceptional taxonomic richness of the pollen assemblages (120 identified taxa) and the presence of borealalpine and Euro-Siberian taxa that no longer exist in Lower Provence.     

Vegetation history of Abies alba Mill. (silver fir) in Switzerland – pollen analytical and genetic surveys related to aspects of vegetation history of Picea abies (L.) H. Karsten (Norway spruce)

This contribution deals with some new aspects of the relationship between the vegetation history of Abies alba Mill. (silver fir) and genetic studies of this tree species in Switzerland. The results of the present study confirm the pollen analytical hypothesis that A. alba re-immigrated into Switzerland mainly from glacial refugia located in northern and central Italy. In particular, some distinct immigration routes of silver fir into the Ticino Alps, Valais, the Bernese Oberland (northwestern Alps), and Graubünden (eastern Alps) could be confirmed by genetic studies. Furthermore, the occurrence of other area-specific alleles indicates an additional influence from eastern European refugia on the Swiss gene pool. Moreover, genetic studies on Picea abies (L.) H. Karsten (Norway spruce) confirmed the general immigration routes from the eastern to the western Alps and from Savoie to the Jura Mountains. The combination of tree pollen and macrofossil analyses of Quaternary sediments with genetic studies of the same tree species represents a considerable research potential and is a new approach of floristic and genetic research. Received November 17, 2000 / Accepted April 27, 2001     

Vegetation and climate changes during the last 21 000 years in S.W. Africa based on a marine pollen record

A high resolution marine pollen record from site GeoB1023, west of the northern Namib desert provides data on vegetation and climate change for the last 21 ka at an average resolution of 185 y. Pollen and spores are mainly delivered to the site by the Cunene river and by surface and mid-tropospheric wind systems. The main pollen source areas are located between 13°S and 21°S, which includes the northern Namib desert and semi-desert, the Angola-northern Namibian highland, and the north-western Kalahari. The pollen spectra reflect environmental changes in the region. The last glacial maximum (LGM) was characterised by colder and more arid conditions than at present, when a vegetation with temperate elements such as Asteroideae, Ericaceae, and Restionaceae grew north of 21°S. At 17.5 ka cal. B.P., an amelioration both in temperature and humidity terminated the LGM but, in the northern Kalahari, mean annual rainfall in the interval 17.5-14.4 ka cal. B.P. was probably 100–150 mm lower than at present (400–500 mm/y). The Late-glacial to early Holocene transition includes two arid periods, i.e. 14.4–12.5 and 10.9–9.3 ka cal. B.P. The last part of the former period may be correlated with the Younger Dryas. The warmest and most humid period in the Holocene occurred between 6.3 and 4.8 ka cal. B.P. During the last 2000 years, human impact, as reflected by indications of deforestation, enhanced burning and overgrazing, progressively intensified.