The burrows of the miocene beaver palaeocastor,Western Nebraska,U.S.A

Daimonelix is a name given to terrestrial lebensspuren of the late Oligocene—early Miocene beaver genus Palaeocastor, and is not a plant or fresh-water sponge as was originally believed by Barbour. Palaeocastor belongs to a lineage of castorids always found in upland habitat, never near evidence of ponded water. Daimonelices are found in high concentrations in the Harrison Formation of western Nebraska and eastern Wyoming, which represents a semiarid, upland paleoenvironment of sandy substrate. Assignment of a large sample of contemporaneous Daimonelix to the species P. fossor is based on a series of incisor width measurements and on mean shaft diameters. The remarkable preservation of daimonelices, which permits detailed analysis of events in beaver burrow construction, is shown to be due to rapid silicification of plant roots invading the burrows. Some aspects of Palaeocastor ecology and ethology are also clarified. The spatial distribution of Daimonelix consists of scattered towns of high burrow density. One such colony is shown in a detailed paleogeographic map. Criteria for time equivalency of such burrow aggregations are established. The entoptychine gopher Gregorymys, the carnivore Zodiolestes, and the larger beaver P. magnus are also shown to be occasional inhabitants of Palaeocastor fossor colonies.     

A new lowermost middle Cambrian (Series 3, Stage 5) faunule from Saxony (Germany) and its bearing on the tectonostratigraphic history of the Saxothuringian domain

The core of borehole 1209/78 west of Doberlug–Kirchhain and south of Herzberg in the Torgau–Doberlug Syncline records an atypical lower part of the Tröbitz Formation with thin limestone horizons. These limestone layers include the remains of a low to moderately diverse fauna with the trilobites Protolenus (Hupeolenus) bergstroemi n. sp., Cambrunicornia saxonica n. sp., Ornamentaspis? aff. todraensis Geyer 1990a, Calodiscus? n. sp., the remains of two undetermined olenelloid? and paradoxidid? species, at least two brachiopods (Trematobolus, undetermined acrotretoid), and one hyolith. The fauna clearly suggests a position in the lower Agdzian stage of the West Gondwana chronostratigraphic scheme and correlation with the lowermost to lower Middle Cambrian strata in regions such as the Moroccan Atlas ranges and northern Spain, so the assemblages represent the oldest Middle Cambrian fauna known from the Saxothuringian domain and reconfirm the palaeogeographic position in the Perigondwanan segment. The lithological differences of the fossiliferous cores from those of the typical Tröbitz Formation and the recorded high-energy conditions indicate high-frequency sea-level changes suggesting that this part of the succession may be a late stage of the subglobally recognizable eustatic sea-level fluctuations at the traditional Lower–Middle Cambrian boundary interval.     

Early Miocene Amphibians (Caudata, Salientia) from the Mokrá-Western Quarry (Czech Republic) with comments on the evolution of Early Miocene amphibian assemblages in Central Europe

The Mokrá-Western Quarry exhibits the rare occurrence of Early Miocene (MN 4) vertebrate fauna within the area of the eastern part of Central Europe. In addition to a rich fauna of reptiles and mammals, two fossiliferous karst joints (Mokrá-Western Quarry, 1/2001 Turtle Joint and Mokrá-Western Quarry, 2/2003 Reptile Joint) yielded a rich fauna of amphibians including 13 amphibian taxa: Salamandridae: Mioproteus sp., Chelotriton sp., type I, Chelotriton sp., type II, Triturus aff. roehrsi, Triturus cf. marmoratus, Triturus sp. (T. cristatus species group), Chioglossa meini, Mertensiella mera, Salamandridae gen. and sp. indet.; Pelobatidae: Pelobates sanchizi; Ranidae: Rana sp. (synklepton Rana esculenta); Bufonidae: Bufo sp. The first records of the West European species Triturus cf. marmoratus and Chioglossa meini are reported from the eastern part of Central Europe indicating the wide distribution of those taxa throughout the whole of Europe as early as MN 4. The oldest known record of Pelobates sanchizi documents the Early Miocene presence of representatives closely related to the extinct Late Oligocene representatives of Pelobates. The slow evolution of amphibian species is documented by the presence of Triturus cf. marmoratus and the oldest known occurrence of the extinct salamander Mertensiella mera.     

Early Silurian (Telychian) rugose coral fauna of Daguan area, northeast Yunnan province, China

Study on rugose coral fauna of the Sifengya Formation (early Telychian) and Daluzhai Formation (mid-late Telychian) in Daguan area, northeast Yunnan Province, China was carried out. Rugose coral fauna of the Sifengya Formation included 18 genera and 34 species, while Daluzhai Formation with nine genera, ten species. We described rugose coral fauna (12 genera, 19 species) including one new genus and five new species, i.e. Protoketophyllum daguanense gen. et sp. nov., Crassilasma huanggexiense sp. nov., Pseudophaulactis heae sp. nov., P. convolutus sp. nov., and Shensiphyllum minor sp. nov. The characteristics and geological significance of rugose coral fauna of Sifengya Formation and Daluzhai Formation were analyzed. Particularly, rugose coral fauna of the Sifengya Formation represent early Telychian rugosan fauna in the Upper Yangtze region and improve the sequences of early Silurian (Llandovery) rugose coral assemblages in Yangtze region. It is therefore very meaningful to further analyze radiation period of rugose coral fauna in such epoch. __________ Translated from Acta Palaeontologica Sinica, 2005, 44(2): 229–246 [译自: 古生物学报, 2005, 44(2): 229–246]     

Taxonomical and palaeoecological investigations of the chondrichthyan and osteichthyan fish remains from the Middle-Late Triassic deposits of the Villány Hills (Southern Hungary)

Two sites in the Villány Hills, Hungary, have yielded rich fish assemblages from Middle to Late Triassic shallow marine deposits. The collected material comes from the Ladinian Templomhegy Dolomite Member and from the Carnian Mészhegy Sandstone Formation. The ichthyofauna is composed of both chondrichthyans (Hybodontidae indet., Palaeobates angustissimus, ‘Polyacrodus’ sp., ?Lissodus sp.) and osteichthyans (Gyrolepis sp., Birgeria sp., and further indeterminate actinopterygians). Despite the large sample size, no remains of neoselachians have been found. The Ladinian Templomhegy Dolomite is dominated by durophagous hybodontiforms (Palaeobates angustissimus, ?Lissodus sp.), but the piscivorous hybodontid and the generalist ‘Polyacrodus’ sp. are missing, while in the fish fauna collected from the Carnian Mészhegy Formation indeterminate piscivorous hybodontids are the most common elements and durophagous forms are much less abundant. The dominance of piscivorous hybodontids in the Carnian Mészhegy Sandstone could be related to the global decrease of diversity of marine fish-eating reptiles (e.g., nothosaurs) or to a change of paleoenvironmental conditions. The present study improves our knowledge on the poorly known Triassic vertebrate faunas of the Tisza Mega-unit, which formed a segment of the passive Neotethys margin of the European Plate and shows an important example of a potential vertebrate faunal shift during the Middle to Late Triassic.     

A New Brachylophosaurin Hadrosaur (Dinosauria: Ornithischia) with an Intermediate Nasal Crest from the Campanian Judith River Formation of Northcentral Montana

Background

Brachylophosaurini is a clade of hadrosaurine dinosaurs currently known from the Campanian (Late Cretaceous) of North America. Its members include: Acristavus gagslarsoni, which lacks a nasal crest; Brachylophosaurus canadensis, which possesses a flat paddle-shaped nasal crest projecting posteriorly over the dorsal skull roof; and Maiasaura peeblesorum, which possesses a dorsally-projecting nasofrontal crest. Acristavus, from the lower Two Medicine Formation of Montana (~81–80 Ma), is hypothesized to be the ancestral member of the clade. Brachylophosaurus specimens are from the middle Oldman Formation of Alberta and equivalent beds in the Judith River Formation of Montana; the upper Oldman Formation is dated 77.8 Ma.

Methodology/Principal Findings

A new brachylophosaurin hadrosaur, Probrachylophosaurus bergei (gen. et sp. nov.) is described and phylogenetically analyzed based on the skull and postcranium of a large individual from the Judith River Formation of northcentral Montana (79.8–79.5 Ma); the horizon is equivalent to the lower Oldman Formation of Alberta. Cranial morphology of Probrachylophosaurus, most notably the nasal crest, is intermediate between Acristavus and Brachylophosaurus. In Brachylophosaurus, the nasal crest lengthens and flattens ontogenetically, covering the supratemporal fenestrae in large adults. The smaller nasal crest of Probrachylophosaurus is strongly triangular in cross section and only minimally overhangs the supratemporal fenestrae, similar to an ontogenetically earlier stage of Brachylophosaurus. Sutural fusion and tibial osteohistology reveal that the holotype of Probrachylophosaurus was relatively more mature than a similarly large Brachylophosaurus specimen; thus, Probrachylophosaurus is not simply an immature Brachylophosaurus.

Conclusions/Significance

The small triangular posteriorly oriented nasal crest of Probrachylophosaurus is proposed to represent a transitional nasal morphology between that of a non-crested ancestor such as Acristavus and the large flat posteriorly oriented nasal crest of adult Brachylophosaurus. Because Probrachylophosaurus is stratigraphically and morphologically intermediate between these taxa, Probrachylophosaurus is hypothesized to be an intermediate member of the Acristavus-Brachylophosaurus evolutionary lineage.     

The enamel microstructure ofAnchitheriomys (Rodentia, Mammalia) in comparison with that of other beavers and of porcupines

In the enamel microstructure of the incisors, porcupines (Hystricidae) are characterized by multiserial Hunter-Schreger bands (HSB) while beavers (Castoridae) basically have uniserial HSB. In addition, two groups can be differentiated among beavers based on enamel microstructure.Palaeocastor, Steneofiber andCastor show rather typical uniserial HSB. OnlyCastor shows a slight tendency of fusion of the bands. In contrast,Trogontherium andCastoroides are characterized by the regular occurrence of fused bands.Anchitheriomys, having uniserial HSB and a high frequency of fused bands fits well with theTrogontherium group, proving the castorid nature ofAnchitheriomys. Furthermore, the enamel differentiation observed among the Castoridae thus provides an important character for further systematic subdivision of the beavers. In the enamel of the cheek teeth, porcupines can be told apart from beavers as well.Anchitheriomys shows a two-layered schmelzmuster with inner radial enamel like other beavers. Porcupines have thick HSB throughout the enamel layer. This character in the cheek teeth facilitates the separation of isolated beaver teeth from porcupine teeth.     

A late early to early middle Eocene mammal assemblage from Bayan Ulan (Inner Mongolia,China): Implication for the reassessment of the Arshantan Asian Land Mammal Age

Paleogene mammal localities of North China are particularly well represented in the Erlian Basin, Inner Mongolia. Among them, the locality of Bayan Ulan is most famous for its late Paleocene Gashatan fauna. However, the younger Arshantan fauna of the same site is not well known, since no extensive study has been done so far. Here, we present a small mammal assemblage based on dental and tarsal material from a new Arshantan collection retrieved from the red beds of the late early to early middle Eocene Arshanto Formation at Bayan Ulan. It consists of at least six different taxa: the basal lagomorph Dawsonolagus antiquus, the large pantodont Pantolambdodon sp., the tapiroid Schlosseria magister, and the rhinocerotoids Hyrachyus crista and Rhodopagus guoi nov. sp. The assemblage is dominated by perissodactyls, especially Lophialetidae and Hyracodontidae. For the first time, p4-m1 of Dawsonolagus antiquus, tarsal material from Pantolambdodon sp., and lower dentition and tarsals of Hyrachyus crista are described and illustrated. Unlike other described Arshantan faunas, the Bayan Ulan Arshantan mammal assemblage has been collected exclusively from a single locality, which contributes to the reassessment of the misunderstood Arshantan Asian Land Mammal Age.     

Eine Ammoniten-Fauna aus der Badamu-Formation (Unterbajocium,sauzei- Zone) westlich von Kerman (Zentraliran)

A fauna of small-sized ammonites is described from the Upper Badamu Formation west of Kerman, including 15 species. Following species are new: “Pelekodites”kermanensis n. sp., “Pelekodites”nodosus n. sp.,Lissoceras (Microlissoceras) badamui n. sp.,Etropolia sapunovi n. sp. The fauna is closely related to the west and northwest European realm. It corresponds to thesauzei zone. Kurzfassung: Aus der oberen Badamu-Formation wird eine insgesamt kleinwüchsige Ammoniten-Fauna mit 15 Arten beschrieben, darunter die neuen Arten: »Pelekodites«kermanensis n. sp., »Pelekodites«nodosus n. sp.,Lissoceras (Microlissoceras) badamui n. sp.,Etropolia sapunovi n. sp. Die Fauna hat eindeutig einen west- bis nordwesteuropäischen Charakter und gehört dersauzei- Zone an.     

The Basal Nodosaurid Ankylosaur Europelta carbonensis n. gen., n. sp. from the Lower Cretaceous (Lower Albian) Escucha Formation of Northeastern Spain

Nodosaurids are poorly known from the Lower Cretaceous of Europe. Two associated ankylosaur skeletons excavated from the lower Albian carbonaceous member of the Escucha Formation near Ariño in northeastern Teruel, Spain reveal nearly all the diagnostic recognized character that define nodosaurid ankylosaurs. These new specimens comprise a new genus and species of nodosaurid ankylosaur and represent the single most complete taxon of ankylosaur from the Cretaceous of Europe. These two specimens were examined and compared to all other known ankylosaurs. Comparisons of these specimens document that Europelta carbonensis n. gen., n. sp. is a nodosaur and is the sister taxon to the Late Cretaceous nodosaurids Anoplosaurus, Hungarosaurus, and Struthiosaurus, defining a monophyletic clade of European nodosaurids– the Struthiosaurinae.     

新疆吐鲁番盆地桃树园晚二叠世非海相腹足类

提要新疆吐鲁番盆地桃树园地区晚二叠世地层称作下仓房沟群,自下而上分为泉子街组、梧桐沟组和锅底坑组,为河–湖相碎屑沉积,剖面连续,层序清楚,动、植物化石丰富。文中研究的腹足类标本产于桃东沟剖面梧桐沟组下部和中部的介壳灰岩层和灰岩透镜体中,见有2层,计有2科4属6种:Xinjiangospira rotundata Yu et Zhu,Xinjiangospira habita sp.nov.、Hydrobia turpanensis Wei、Hydrobia orientalis sp.nov.、Pseudamnicola taodonggouensis sp.nov.和Valvata complanusa sp.nov.;与腹足类共生的有双壳类、叶肢介、介形类,以及植物和脊椎动物化石等,其中双壳类主要是Palaeanodonta,Palaeomutela和Anthraconauta等属。该腹足动物群由Hydrobiidae和Valvatidae的属种组成,标本数量多,壳体小,保存完好,是迄今已知属种最丰富的古生代淡水腹足类动物群。当前梧桐沟组腹足动物群面貌与准噶尔盆地大龙口剖面小龙口组...     

Tremadoc (earliest Ordovician) brachiopods from Purmamarca and the Sierra de Mojotoro, Cordillera Oriental of northwestern Argentina

The Late Tremadoc storm-dominated shoreface to inner platform deposits exposed west of the Purmamarca village (Coquena Formation) contain a considerably more diverse brachiopod fauna than previously reported. Coquinite horizons from the lower heterolitic succession have yielded monospecific associations of Nanorthis purmamarcaensis nov. sp. (formerly assigned to N. christianiae KJERULF), which is also reported from the Late Tremadoc rocks of the Cerro San Bernardo area. The fine-grained Upper Member of the Coquena Formation contains a more diverse fauna composed by Nanorthis brachymyaria nov. sp., Astraborthis quebradensis nov. sp. and the new plectorthid genus Lipanorthis (type species L. andinus nov. sp.). A different species of Lipanorthis (L. santalaurae nov. sp.) from the Mid Tremadoc Floresta Formation of the Sierra de Mojotoro is also described.     

Brachypotherium (Perissodactyla,Rhinocerotidae) from the late Miocene of Samburu Hills,Kenya

Several isolated cheek teeth and mandibular specimens of Rhinocerotidae (Mammalia, Perissodactyla) from the upper Miocene Namurungule Formation in Samburu Hills, Kenya, are redescribed. Previously, these specimens had been identified as Chilotheridium pattersoni, Chilotheridium sp., Paradiceros mukirii, and Paradiceros sp. They are reidentified here as documenting the genus Brachypotherium based on their bucco-lingually broad molariform upper premolars with short crochet and flattened buccal walls on both upper and lower molars, the latter having a shallow external groove. Comparisons with other Brachypotherium species suggest that the present specimens belong to Brachypotherium sp. cf. B. minor. The presence of Brachypotherium in the Samburu Hills, at ca. 9.5 Ma, is concordant with the paleoenvironment (presence of lacustrine and river environments) known for this locality during the early late Miocene.     

Eocene siliceous sponges from the Ebro Basin (Catalonia, Spain)

The Eocene (Bartonian) marls of the La Guixa Member and Gurb Member, Vic Marls Formation (Ebro Basin, Catalonia, Spain), contain a very rich and diversified siliceous sponge fauna. The fauna is dominated by hexactinellids; lithistids and other demosponges are rare. It consists of 16 species representing 16 genera. Eleven new species and two new genera are proposed for these sponges: Reguantella cavernosa nov. gen. nov. sp., Regadrella concinna nov. sp. (both Hexactinellida, Lyssacinosa), Eurete clava nov. sp., Pleuroguettardia iberica nov. sp., Aphrocallistes almeriae nov. sp., Hexactinella informis nov. sp. (all Hexactinellida, Hexactinosa), Brachiolites munterensis nov. sp., Centrosia viquensis nov. sp., Callicylix eocenicus nov. sp., Rhizocheton robustus nov. sp. (all Hexactinellida, Lychniscosa), Propetrosia pristina nov. gen. nov. sp. (Demospongia, Haplosclerida). Some genera of sponges in this fauna are still extant, but, in general, the predominant ones are very close in morphology, and, without doubt, closely related to the Late Cretaceous sponges. This fauna also differs considerably, in terms of composition, from most other described faunas of Tertiary sponges from the Mediterranean region, which are dominated by lithistid sponges. Lithistid sponges are rare in this investigated assemblage, which seems most similar to an as yet undescribed Eocene fauna from Italy. There is ecological differentiation in the proportions of particular sponges in various outcrops and/or stratigraphical levels that is clearly associated with water-depth-related controlling factor(s): Munter, Tona and Sta. Cecilia represent the deepest facies, Gurb is intermediate, and St. Roc and Vespella are the most shallow. The exact bathymetric position of the sponge fauna is difficult to estimate, but it seems that 100 m (but probably 200 m and more in the case of the deepest parts) of water depth may be inferred for this facies.     

Middle Ordovician (Llanvirn) ungulate brachiopods and conodonts from the Malyi Karatau Range,Kazakhstan

A new middle Ordovician fauna including conodonts and brachiopods is described from the early Llanvirn Zhyrykaus Formation, Malyi Karatau Range, southern Kazakhstan. The fauna includes the lingulate brachiopodsHyperobolus fragilis n. sp. andTalasotreta apollonovi n. gen. n. sp., as well as the conodontsCordylodus? cf.horridus, Drepanoistodus? sp.,Erraticodon sp.,Parapaltodus simplicissimus, Prioniodus (Baltoniodus) sp.,Protopanderodus cf.varicostatus, andWalliserodus sp. s. f. It represents the first record of Llanvirn conodonts and Ungulate brachiopods from southern Kazakhstan.     

A new diverse shark fauna from the Wordian (Middle Permian) Khuff Formation in the interior Haushi‐Huqf area,Sultanate of Oman

Abstract: Chondrichthyans are newly reported from the autochthonous Wordian Khuff Formation (middle Permian), cropping out in well‐exposed, low‐palaeolatitude sections in the interior Haushi‐Huqf area of Oman. The shark remains comprise isolated teeth, dermal denticles and fin spines and have been recovered by processing limestone in buffered acetic acid from bulk rock samples. The fauna consists of mainly ctenacanthiform and hybodontiform taxa, identified as Glikmanius cf. myachkovensis, Glikmanius culmenis sp. nov., Omanoselache hendersoni gen. et sp. nov., Omanoselache angiolinii gen. et sp. nov., cf. Omanoselache sp., Reesodus underwoodi gen et sp. nov., Teresodus amplexus gen. et sp. nov., Gunnellodus bellistriatus, Khuffia lenis gen. et sp. nov., Khuffia prolixa gen. et sp. nov. and Euselachii sp. indet. Additional specimens include rare teeth of the lonchidiid cf. ‘Palaeozoic Genus 1’ sp., of the neoselachian Cooleyella cf. fordi and a further indeterminate neoselachian, of an indeterminate petalodont and of the holocephalan Deltodus aff. mercurei and Solenodus cf. crenulatus. Fin spines add a further two taxa, Nemacanthus sp. and Amelacanthus cf. sulcatus, which have neoselachian affinities and therefore an unclear relationship to the recovered teeth. The occurrence of Nemacanthus within this Wordian fauna represents the oldest record of this taxon and its only known occurrence in the Palaeozoic. Of the remaining genera, Glikmanius has previously been recorded from the Wordian, whereas for all the others, this study represents their youngest known stratigraphic occurrence and first occurrence in Guadalupian (middle Permian) strata. This adds significantly to our knowledge of the global diversity of chondrichthyans preceding the end‐Guadalupian biotic crisis. Palaeogeographically, for all taxa, this study represents the first record from the western fringe of the marine Neotethyan basin, and only Cooleyella was previously known from the southern (Gondwanan) part of the Pangaean continental margin.     

Early Jurassic (latest Toarcian) brachiopods from the northeastern margin of the Western Tethys (Central Iran) and their paleobiogeographical significance

Brachiopod fauna from central Iran, recorded in the upper part of the Shemshak Group and attributed to the upper Toarcian (Pseudoradiosa-Aalensis zones), are reported for the first time in Iran. The assemblage recognized includes six different taxa: Homoeorhynchia sepahanensis nov. sp., formally described in this paper, Globirhynchia subobsoleta, Pseudogibbirhynchia sp., Tetrarhynchiidae sp. indet., Monsardithyris? aff. haresfieldensis, and Zeilleria cf. leckenbyi. Analysis of faunal affinities with other paleobiogeographical regions shows a free connection of the central Iranian brachiopod fauna and wide areas of the northern shelf margin of the Tethys Ocean. This is due to an apparent disruption of bioprovinciality inferred for the late Toarcian-earliest Aalenian, congruent with a connection through the northern seaway across the peri-Laurasian epicontinental platforms.