A Classificatory System and Geographical Distribution of the Genus Gynostemma BL. (Cucurbitaceae)

The genus Gynostemma consists of 16 species and 2 varieties, which fall into two subgenera and two sections, i. e. , Subgen. Trirostellum including two new sections, Sect. Pentagynae and Sect. Trirostellae, and Subgen. Gynostemma. The modern distribution centre of the genus ranges from drainage areas of the Yangtze River to Yunnan, southwest China. Northwards, it is distributed to the south slope of the Qinling Range and the south branch of the Huaihe River, eastwards through central-eastern China to Korea Peninsula and northern Japan; in the south, it extends to Indo-China, Malaysia, Philippines, Indonesia and Papua New Guinea, with the west limit of this genus found in northwestern India.The distribution pattern of the genus indicates a nature of Tropical Asia Type. Based on the above mentioned distribution pattern, the centre of diversification and the ecological adaptation of both primitive and advanced groups, also by means of palaeogeological, palaeogeographical and palaeoclimatical evidence and the origin of its out group, the genus Gynostemma, together with its closely allied genera Hemsleya, Gomphogyne, etc. is considered to have originated on the Kham-Dian Oldland during the Early Tertiary.     

绞股蓝属植物的分类系统和分布

绞股蓝属全世界有16种2变种,隶属于2亚属2组,即喙果藤亚属(包括五柱绞股蓝组和喙果藤组)及绞股蓝亚属。其现代分布中心或多样化中心为我国长江流域至西南的云南,由此向北达秦岭南坡和淮河流域以南,向东北经华中、华东至朝鲜半岛和日本北部,向南经中南半岛、马来西亚达菲律宾、印度尼西亚诸岛和巴布亚新几内亚,向西达印度西北部,为热带亚洲分布类型。根据该属的原始类群和进化类型的现代分布和多样化中心及它们的生态适应性,与古地质、古地理和古气候的变迁以及外类群起源地等推测,绞股蓝属植物可能与其近缘属——雪胆属和锥形果属共同起源于康滇古陆,起源时间可能为早第三纪。     

A preliminary study on the classification,distribution and ecological nature of genus Stipa L. of China

24 species and 4 varietes of genus Stipa L. of China have been studied and described in this paper. It has been noted that these different species have various geographical distributions, depending on the changes of climatic and edaphic factors of their environments. Based on the study of floral morphology together with ecological and distributional factors, the genus have been divided into 5 sections: 1. Sect. Regelia Tzvel. 2. Sect. Leiostipa Dum. 3. Sect. Barbatae Junge 4. Sect. Stipa 5. Sect. Smirnovia Tzvel. It should be pointed out that the section Regelia as well as two members of section Barbatae, S. purpurea and S. roborowskyi, belong to frigori-xerophilous ecotype, distributing over Qinghai-Xizang Plateau above the forest line. The section Leiostipa belongs to euxerophilous and mesoxerophilous ecotype, distributing widely in northwest, southwest, northeast and east and extends to the forest-steppe vegetational Zone of China. The section Smirnovia and other two members of section Barbatae, S. breviflora and S. orientalis, belong to euxerophilous ecotype, with the latitudinal distribution as far as Nei-Mongo and the yellow soil plateau, with the altitudinal distribution as far as the desert steppe of Sinkang and Qinghai-Xizang plateau. The section Stipa belongs to euxerophilous ecotype, only distributes to the mountain steppe of north Xingkang and the last, section Barbatae is an artificial group having plumes overof the awn only, and its four species have already been mentioned above.     

A Conspectus of the Genus Bergenia Moench

In this paper the classification of the genus Bergenia Moench is provided, its geographic distribution analysed, and the phylogeny also traced. Based on an analysis of morphological characters such as leaves, ocreas, branches of inflorescences, Pedicels, hypanthium, sepals, and glandular indumentum, thi genus is divided into 3 sections: 1. Sect. Scopulosae J. T. Pan, sect. nov., 2. Sect. Bergnia, 3. Sect. Ciliatae (A. Boriss.) J. T. Pan, stat. nov. The Sect. Scopulosae J. T. Pan may be considered as the primitive one, while Sect. Ciliatae (A. Boriss.) J. T. Pan may be regarded as the advanced one, with Sect. Bergenia in between. So far, the genus Bergenia Moench comprises 9 species in the total. Southeast Asia and North Asia (south and east Siberia, USSR) each have only 1 species, West Asia (Afghanistan) has 2, Central Asia (Kirghizia-Tajikistan-Uzbekstan area, USSR) 3, South Asia 4 (Nepal has 4, India, Pakistan and Kashmir area each has 3, Bhutan and Sikkim each has 2), East Asia 6. In East Asia, Mongolia and Korea each have only 1 species, but China has 6 (including endemic species 2 and new species 1). Sichuan Province and Xizang Autonomous Region each have 3, Yunnan Province 2, Shaanxi Province (Qinling Mountains) and Uygur Autonomous Region of Xinjiang each have only 1. Thus the distribution centre of this genus should be in the region covering Sichuan, Yunnan and Xizang. Moreover, it is noteworthy that Bergenia scopulosa T. P. Wang in Sect. Scopulosae seems to have retained primitive characters, for example, non-ciliate leaves and ocreas, glabrous pedicels, hypanthium and sepals, and this primitive species is found in Qinling Mountains and Sichuan. According to the distribution of the primitive species, the author suggests that the centre of origin of this genus be in the region covering Qinling Mountains and Sichuan.     

安蕨属的分类与分布

1849年,捷克人Presl根据菲律宾产的一种蕨类植物Anisocampium cumingia-num Presl,建立了安蕨属Anisocapium Presl,几十年来未被各国植物学家(如Baker、Beddom、Makino、C.Christensen、Christ、Nakai、秦仁昌、Devol、Ohwi和Tagawa)所承认。1940年,Tard.-Blot et C.Chr.编写《印度支那植物通志》(Fl.Gen.Indo-Chine)时接受了安蕨属的概念。1947年,美国人Copeland研究东南亚蕨类植物时,也肯定了安蕨属是一个自然分类群。以后,     

A Revision of Genus Eurya in China

The genus Eurya was established by Thunberg in 1783 on a Japanese species E. japonica. Several decades later, many authors had made more or less intensive study on this genus. But in the earlier period, the different autherities had no clear conception of the genus. Thus Szyszylowicz (1893), Engler (1897) and Melchior (1925) entertained a much broader conception of this genus, including Freziera and Cleyera, both of which are now considered as distinct genera. A. Gray (1855), Vesque (1895), Urban (1896), and Kobuski (1935-37), however, concentrated their study only on the now accepted genus Eurya. Recently, a more detailed study of Chinese Eurya was made by Chang Hung-da (Acta Phytotax. Sin. 3 (1954) 1-59). But all of these authors did not subdivide this genus into more than subgenera except Vesque, who used the number of stamens and the septation of the anther cells to further subdivide this genus into 4 sections. In the present treatment, we base on the following characteristics to make subdivisions: the number of stamens, the septation of the anther cells, the hairness of the ovary, the length of the style, the shape and texture of the sepals, and the shape and hairness of the young branchlets. We assume that the morphology of the flowers has the following evolutionary tendencies: stamens from large (28) to small (5) numbers; anther cells from septate to nonseptate; ovary from being hairy to glabrous, styles from 5-4 free to 3 and more or less connate, from 2-6 mm to 0.5-1 mm long, and ovules from many (60) to few (2-4) in each cell. The genus Eurya has now about 140 species, being divided into 2 subgenera: Subg. Ternstroemiopsis Urban, with 2 species endemic in Sandwich Islands and Subg. Eurya with 138 species, in Asiatic tropic and subtropic regions and southwestern Pacific Islands. According to the characteristics mentioned above, we divide the second subgenus into 2 sections and 8 series, as follows: (I) Sect. Meristotheca Vesque (II) Sect. Eurya Ser. Ciliatae Hu et L. K. Ling Ser. Longistylae Hu et L. K. Ling Ser. Trichocarpae Hu et L. K. Ling Ser. Rigidisepalae Hu et L. K. Ling Ser. Tetragonocladae Hu et L. K. Ling Ser. Nitidae Hu et L. K. Ling Ser. Muricatae Hu et L. K. Ling Ser. Brevistylae Hu et L. K. Ling In China, the subgenus Eurya distributes east from Taiwan, west to the western part of Szechuan and Yunnan, and south from Hainan, north to the southern slope of Tsin-ling Range. The region north from southern part of Nan-ling Range, southwest to southeast Yunnan, south to northern part of the Peoples Republic of Vietnam, seems to be the centre of both maximum variety and frequency of this subgenus, for the number of species and the representatives of more primitive taxa in this region are much richer than in any other regions of the world. From this centre going northeast to Japan and Korea, west to eastern part of India, south to Java and Sumatra, north to the southern slope of Tsinling Range, the number of species and types gradually decreases, and especially the primitive series and species rapidly disappear. In addition, many species are also found in the Island of Irian, which we incline to consider as another young centre of development for this subgenus. Furthermore, according to the distribution of quite a large number of the species in China, we can recognize several boundary lines which are in agreement with the limits of the floristic and geobotanic provinces of China. In this article we have enumerated 80 species, and 11 varieties of the genus Eurya of China, among which are published for the first time 11 new species and 1 new variety, one species, E. persicaefolia Gagnepain, is first recorded from China, a number of specific names have been restored and a number reduced to synonyms. They are as follows: Restored species: Eurya acuminatissima Merrill & Chun E. patentipila Chun E. henryi Hemsley Reduced to synonyms: Eurya parastrigillosa Hsu ( E. patentipila Chun) E. changii Hsu (E. fangii var. megaphylla Hsu) E. chienii Hsu (E. persicaefolia Gagnepain) E. hwangshanensis Hsu (E. saxicola Chang) E. fangii Rehd. var. glaberrima Hsu (E. cavinervis Vesque) E. pseudopolyneura Chang (E. impressinervis Kobuski) E. longistyla Chang (E. stenophylla Merrill) E. huiana Kobuski f. glaberrima Chang (E. muricata Dunn) New combination: Eurya muricata Dunn var. huiana (Kob.) Hu et L. K. Ling E. aurea (Levl.) Hu et L. K. Ling     

A Preliminary Study on the Taxonomy of the Family Magnoliaceae

A new system of classification of Magnoliaceae proposed. This paper deals mainly with taxonomy and phytogeography of the family Magnoliaceae on the basis of external morphology, wood anatomy and palynology. Different authors have had different ideas about the delimitation of genera of this family, their controversy being carried on through more than one hundred years (Table I). Since I have been engaged in the work of the Flora Reipublicae Popularis Sinicae, I have accumulated a considerable amount of information and material and have investigated the living plants at their natural localities, which enable me to find out the evolutionary tendencies and primitive morphological characters of various genera of the family. According to the evolutionary tendencies of the characters and the geographical distribution of this family I propose a new system by dividing it into two subfamilies, Magnolioideae and Liriodendroideae Law (1979), two tribes, Magnolieae and Michelieae Law, four subtribes, Manglietiinae Law, Magnoliinae, Elmerrilliinae Law and Micheliinae, and fifteen genera (Fig. 1 ), a system which is different from those by J. D. Dandy (1964-1974) and the other authors. The recent distribution and possible survival centre of Magnoliaceae. The members of Magnoliaceae are distributed chiefly in temperate and tropical zones of the Northern Hemisphere, ——Southeast Asia and southeast North America, but a few genera and species also occur in the Malay Archipelago and Brazil of the Southern Hemisphere. Forty species of 4 genera occur in America, among which one genus (Dugendiodendron) is endemic to the continent, while about 200 species of 14 genera occur in Southeast Asia, of which 12 genera are endemic. In China there are about 110 species of 11 genera which mostly occur in Guangxi, Guangdong and Yunnan; 58 species and more than 9 genera occur in the mountainous districts of Yunnan. Moreover, one genus (Manglietiastrum Law, 1979) and 19 species are endemic to this region. The family in discussion is much limited to or interruptedly distributed in the mountainous regions of Guangxi, Guangdong and Yunnan. The regions are found to have a great abundance of species, and the members of the relatively primitive taxa are also much more there than in the other regions of the world. The major genera, Manglietia, Magnolia and Michelia, possess 160 out of a total of 240 species in the whole family. Talauma has 40 species, while the other eleven genera each contain only 2 to 7 species, even with one monotypic genus. These three major genera are sufficient for indicating the evolutionary tendency and geographical distribution of Magnoliaceae. It is worthwhile discussing their morphological characters and distributional patterns as follows: The members of Manglietia are all evergreen trees, with flowers terminal, anthers dehiscing introrsely, filaments very short and flat, ovules 4 or more per carpel. This is considered as the most primitive genus in subtribe Manglietiinae. Eighteen out of a total of 35 species of the genus are distributed in the western, southwest to southeast Yunnan. Very primitive species, such as Manglietia hookeri, M. insignis and M. megaphylla, M. grandis, also occur in this region. They are distributed from Yunnan eastwards to Zhejiang and Fujian through central China, south China, with only one species (Manglietia microtricha) of the genus westwards to Xizang. There are several species distributing southwards from northeast India to the Malay Archipelago (Fig. 7). The members of Magnolia are evergreen and deciduous trees or shrubs, with flowers terminal, anthers dehiscing introrsely or laterally, ovules 2 per carpel, stipule adnate to the petiole. The genus Magnolia is the most primitive in the subtribe Magnoliinae and is the largest genus of the family Magnoliaceae. Its deciduous species are distributed from Yunnan north-eastwards to Korea and Japan (Kurile N. 46’) through Central China, North China and westwards to Burma, the eastern Himalayas and northeast India. The evergreen species are distributed from northeast Yunnan (China) to the Malay Archipelago. In China there are 23 species, of which 15 seem to be very primitive, e.g. Magnolia henryi, M. delavayi, M. officinalis and M. rostrata, which occur in Guangxi, Guangdong and Yunnan. The members of Michelia are evergreen trees or shrubs, with flowers axillary, anthers dehiscing laterally or sublaterally, gynoecium stipitate, carpels numerous or few. Michelia is considered to be the most primitive in the subtribe Micheliinae, and is to the second largest genus of the family. About 23 out of a total of 50 species of this genus are very primitive, e.g. Michelia sphaerantha, M. lacei, M. champaca, and M. flavidiflora, which occur in Guangdong, Guangxi and Yunnan (the distributional center of the family under discussion) and extend eastwards to Taiwan of China, southern Japan through central China, southwards to the Malay Archipelago through Indo-China. westwards to Xizang of China, and south-westwards to India and Sri Lanka (Fig. 7). The members of Magnoliaceae are concentrated in Guangxi, Guangdong and Yunnan and radiate from there. The farther away from the centre, the less members we are able to find, but the more advanced they are in morphology. In this old geographical centre there are more primitive species, more endemics and more monotypic genera. Thus it is reasonable to assume that the region of Guangxi, Guangdong and Yunnan, China, is not only the centre of recent distribution, but also the chief survival centreof Magnoliaceae in the world.     

A Study on the Genus Chrysosplenium L. from China (Cont.)

This paper deals with the taxonomy and geographic distribution of the genus Chrysosplenium L. in China. Based on the characters and evolution of the seed, capsule, disk, ovary and leaf, the species of this genus can be grouped into 2 subgenera, 5 sections and 16 series. There are 2 subgenera, 5 sections and 11 series in China. They are as follows: I. Subgen. Gamosplenium Maxim. emend. J. T. Pan Leaves alternate. Lectotype: Chrysosplenium carnosum Hook. f. et Thoms. 1. Sect. Alternifolia Franch. emend. J. T. Pan Seeds smooth and glabrous. Type: Chrysosplenium alternifolium L. (1) Ser. Nudicaulia Maxim. emend. J. T. Pan Disk obscure or absent; ovary nearly half-inferior, sometimes mostly inferior; capsule generally subtruncate and emarginate at top and bilobed with equal and horizontally divaricate or suberect lobes; seeds smooth and glabrous. Type: Chrysosplenium nudicaule Maxim. (2) Ser. Alternifolia Maxim. emend. J. T. Pan Disk 8-lobed; ovary nearly half-inferior; capsule generally subtruncate and emarginate at top, and bilobed with equal and horizontally divaricate lobes; seeds smooth and glabrous. Type: Chrysosplenium alternifolia L. 2. Sect. Nephrophylloides Turcz. Seeds minutely papillose or pilose. Type: Chrysosplenium sedakowii Turcz. (1) Ser. Macrophylla Franch. emend. J. T. Pan Disk obscure or absent; ovary nearly half-inferior; capsule nearly truncate and emarginate at top, and bilobed with equal lobes; seeds minutely papillose. Type: Chrysosplenium macrophyllum Oliv. (2) Ser. Ovalifolia Maxim. emend. J. T. Pan Disk generally 8-, rarely 4-, lobed, papillae absent around disk; ovary mostly inferior; capsule subtruncate and emarginate at top; seeds minutely papillose or pilose. Type: Chrysosplenium ovalifolium M. Bieb. ex Bunge (3) Ser. Lanuginosa Hara, emend. J. T. Pan Papillae numerous, brown around reduced disk; ovary mostly inferior; capsule nearly truncate and emarginate at top; seeds minutely papillose. Type: Chrysosplenium lanuginosum Hook. f. et Thoms. II. Subgen. Chrysosplenium Leaves opposite. Type: Chrysosplenium oppositifolium L. 1. Sect. Trichosperma J. T. Pan, sect. nov. Capsule not truncate at top, and bilobed with subequal, suberect or divergent lobes. Type: Chrysosplenium trichospermum Edgew. ex Hook. f. et Thoms. This section is divided into 4 series in the world, with only 1 in China. (1) Ser. Nepalensia Maxim. emend. J. T. Pan Disk obscure or absent; ovary generally mostly inferior; cassule not truncate at top, and bilobed with subequal and suberect or divergent lobes; seeds smooth and glabrous. Type: Chrysosplenium nepalense D. Don 2. Sect. Grayana J. T. Pan, sect. nov. Capsule bilobed with distinctly unequal and ascending lobes. Type: Chrysosplenium grayanum Maxim. This section consists of 4 series in the world, with 3 series in China. (1) Ser. Sinica Maxim. emend. J. T. Pan Disk obscure or absent; ovary nearly half-superior; capsule bilobed with distinctly unequal and ascending lobes; seeds minutely papillose. Type: Chrysosplenium sinicum Maxim. (2) Ser. Esulcata Franch. emend. J. T. Pan Disk (4)-8-lobed; ovary generally half-inferior; capsule bilobed with unequal and ascending lobes; seeds minutely papillose or pilose. Lectotype: Chrysosplenium dubium J. Gayex DC. (3) Ser. pilosa maxim. emend. J. T. Pan Disk obscure or absent; ovary nearly half-inferior; capsule bilobed with distinctly unequal and ascending lobes; seeds distinctly longitudinally ll-18-costate and minutely papillose or tuberculate on the ridge. Type: Chrysosplenium pilosum Maxim. 3. Sect. Chrysosplenium Capsule nearly truncate and emarginate at top, and bilobed with equal and horizontally divaricate lobes. Type: Chrysosplenium oppositifolium L. (1) Ser. Romosa J. T. Pan, ser. nov. Disk distinctly 8-lobed, papillae sparse, brown around disk; ovary mostly inferior; capsule nearly truncate and emarginate at top, and bilobed with equal and horizontally divaricate lobes; seeds smooth and glabrous. Type: Chrysosplenium ramosum Maxim. This series is monospecific one, also occurring in China, namely C. ramosum Maxim. (2) Ser. Delavayi Hara Disk distinctly 8-lobed, Papillae sparse, brown around the disk; ovary mostly inferior; capsule nearly truncate and emarginate at top, and bilobed with equal and horizontally divaricate lobes; seeds distinctly longitudinally 10-16-costate and transversely striate on the ridge. Type: Chrysosplenium delavayi Franch. This series can be considered as the most advanced one in the Chrysaspleninm L. So far, the Chrysosplenium L. comprises 64 species in the world, among which 1 species is found in North Africa, 2 in South America, 4 in Europe, 5 in North America, 56 in Asia, of which 3 occur in Sikkim, 5 Bhutan, 5 Mongolia, 6 north Burma, 6 Korea, 7 north India, 8 Nepal, 12 Japan, 17 U.S.S.R. (of which 3 also in Europe), 34 China (including 22 endemic species and 3 new species). In China, Fujian and Guangdong Provinces and Zhuang Autonomous Region of Guangxi each has only 1 species, Taiwan, Zhejiang, Shanxi and Hebei Provinces and Uygur Autonomous Region of Xinjiang each has 2, Anhui, Jiangxi and Hunan Provinces each has 3, Qinghai Province 4, Heilongjiang, Liaoning and Guizhou Provinces each has 5, Jilin and Hubei Provinces each has 6, Gausu Province 8, Shaanxi Province and Xizang (Tibet) Autonomous Region each has 10, Yunnan Province has 11, Sichuan Province has 14. Thus the distribution centre of this genus should be in the north temperate zone of Asia, and the region covering Shaanxi Gansu, Sichuan, Yunnan and Xizang may be regarded as an important part of this centre. The 7 species of Ser. Nudicaula Maxim. emend. J. T. Pan can be considered as the most primitive ones in this genus. They are mostly distributed in Shaanxi (Qin Ling), south Gansu, southeast Qinghai, southwest Sichuan and nothwest Yunnan of China. This region may be considered as the centre of the origin (or at least differentiation) of this genus. The new species and the new varieties described in this paper are as follows: C. hydrocotylifolium Levl. et Vant. var. emeiense J. T. Pan, C. taibaishanense J. T. Pan, C. lixianense Jien ex J. T. Pan, C. qinlingense Jien ex J. T. Pan.     

论世界蒿属植物区系

本文从多学科, 包括形态、地理分布、孢粉、古植物、谱系分支分析及化学成分等学科的研究, 论述了世界蒿属植物的系统分类及其种属地理、历史地理和区系地理.     

湖北鳞毛蕨属植物的地理分布及区系特征

对湖北鳞毛蕨后植物的地理分布和区系特点进行了研究。鳞毛蕨后植物广布于世界各地,该后的分布和多样性中心位于中国西南部和东喜马拉雅山区;另一中心则位于日本,中国东南部和南部。鳞毛蕨后是一个自然的北温带分布属。中国有鳞毛蕨后植物134种(包括7变种),西南地区(云南、四川、贵州等)是国产鳞毛蕨属植物分布最集中的地区。区系分析表明：湖北鳞毛蕨后植物种类比较丰富,有36种,主要分布于鄂西北和鄂西南山区,是构成湖北森林植物区系林下草本植物的主要成分之一;地理成分比较复杂,种的分析显示出以中国一日本分布和中国持有分布为主的特点;与相邻省鳞毛蕨属植物区系的关系比较密切;区系过渡性明显。     

中国桂樱属植物的分类研究

桂樱属(Laurocerasus Tourn.ex Duh.)是蔷薇科李亚科的一个属。它是De Tournefort(Institutiones rei Herbariae 627.t.403.1700)以Prunus laurocerasus L.和P.lusitanica L.两种奠定基础,而至1755年由Duhmal代为正式发表而建立。自该属建立以后的二百多年来,各国植物学家对其分类位置各持不同观点。     

论世界芨芨草属(禾本科)的地理分布

本文详细讨论了世界芨芨草属的地理分布等问题。1．全世界芨芨草属共有23种1变种,分为5个组。本文对它们进行了系统介绍。2．属的地理分布,最北为北纬62°(羽茅、毛颖芨芨草),最南为北纬26°(林阴芨芨草)。就海拔而论,分布最低的海拔记录为120m(雀麦芨芨草),分布最高的海拔记录为4600m(干生芨芨草和藏芨芨草)。3．本文讨论了芨芨草属5个组(芨芨草组,钝基草组,直芒草组,新芨芨草组,拟芨芨草组)的系统位置,和每个组包括的种类及5个组的分布格局。4．根据塔赫他间世界植物区系区划,统计了每个区的种数,明显看出伊朗—土兰区种类(18／24)是第一位,东亚区(14／24)居第二位。中国有17种,横断山脉地区、华北地区和唐古特地区种数最丰富(10种和9种)。5．研究结果表明：(A)从种的分布格局分析可见,横断山脉地区北部、唐古特地区东部和华北地区西部的交汇地是芨芨草属分布中心。(B)根据芨芨草属形态特征演化趋势分析和地史学资料推测横断山脉地区北部是芨芨草属的起源地。(C)有三条路线向外散布:a)从横断山脉地区向西沿喜马拉雅山脉,经克什米尔地区抵达地中海和中欧;b)从横断山脉向西北经祁连山、天山、塔里木盆地西侧山地,抵吉尔吉斯斯坦伊塞克湖; c)由横断山脉向东北经甘肃、宁夏、陕西、山西、河北和东北,抵达西伯利亚,东达堪察加半岛,西至鄂毕河上游,并经白令海峡陆桥分布到美国内华达山脉和落基山山脉。(D)该属植物集中分布于北半球半湿润、半干旱和干旱地区,以及极端干旱的荒漠区山地。植物的形成、发展和生态适应与气候相联系,并经过长期的适应和进化,塑造了一系列中生、旱中生的形态－生态特征和生活型。     

A Study of the Genus Atraphaxis in China and the System of Atraphaxideae (Polygonaceae)

The genus Atraphaxis is found mainly in gravel steppes, sandhills and stony slopes or desert, with only a few species in meadow or river valleys. In China there are two sections and eleven species (including three varieties), which are distributed mainly in the northwest, with a few in the northeast and the north. Characters of the genus were compared and analyzed in the present work. Outer perianth segments 2, small, reflexed in fruit, inner perianth segments 2 or 3, large, erected and enveloping the nut. The embryo is curved, but those in the other genera of Atraphaxideae are straight. The curved embryo is the primitive character, because most species of the order Caryophyllales are of a circular embryo, which is belived to be ancestral in the Polygonaceae. The Pollen ornamentations of Atraphaxis are striate or striate-reticulate, different from those of the other genera. A new system of the tribe Atraphaxideae in proposed in the present paper. Based on the character analysis of the species in China, the present authors believe that Sect. Tragopyrum is more primitive than Sect. Atraphaxis. According to the distribution, the genus Atraphaxis might originate in Kazakstan, where not only are most species found, but also the most primitive species, like A. muchketovii, as considered by A. N. Krasnov, are found. Central Asia is considered as the distribution centre and origin centre of thegenus Atraphaxis.     

First record of the family Thamnocephalidae (Crustacea: Anostraca) from Southeast Asia and description of a new species of Branchinella

During a two-year investigation of freshwater microcrustaceans in Thailand, an undescribed species of fairy shrimp of the genus Branchinella Sayce, 1903 in the family Thamnocephalidae, was discovered. This is the first record of the genus and the family from Southeast Asia and the second anostracan reported from Thailand. The new species, Branchinella thailandensis n. sp. appears to be morphologically intermediate between B. kugenumaensis (Ishikawa, 1895) from Japan and east China, and B. madurai Raj, 1951 from India and Pakistan. B. thailandensis n. sp. was collected from several temporary ponds in 11 provinces of the northeast and the central Thailand, often co-occurring with Streptocephalus sirindhornae Sanoamuang, Murugan, Weekers & Dumont, 2000.     

鳜亚科SINIPERCINAE鱼类的分类整理和地理分布

鳜亚科鱼类仅产于东亚。通过对中国各地、日本和朝鲜标本的研究,本文把鳜亚科鱼类总结为鳜、长体鳜和少鳞鳜3属11种,还对过去一些种的异名作了讨论,提出了看法。 本亚科共有3属9种分布于中国,另两种少鳞鳜只见于日本和朝鲜,是日本和朝鲜地区的特有种。越南有3种记录(Mai,1978)。其中鳜鱼和斑鳜分布最北,是横跨东洋地区和古北区的两个广布种。其他各种皆集中分布于江、淮流域以南,主要在长江流域以南和红河流域以东,但台湾省无鳜类分布。长江流域以南低纬度的华南区是鳜亚科鱼类的分布中心。     

中国蹄盖蕨属轴果蹄盖蕨系的研究

本文是中国蹄盖蕨属植物研究的系列论文的第三篇,对国产轴果蹄盖蕨系植物进行了分类学订正,详细记载了中国产该系植物11种,首次将20余个名称归入该系的一些种下做为异名处理。轴果蹄盖蕨系植物是蹄盖蕨属中自然的一群,以其铁角蕨类型的孢子囊群和囊群益与其它属下类群相区别。本系植物分布于亚洲热带和亚热带山地,常见于山地常绿阔叶林下,海拔500～1800m。在华中和华东地区其分布区北界不超过长江一线。中国西南地区和台湾及日本为该系的三个分化中心。     

On the Genus Athyrium Ser. Epiraches Ching et Y. T. Hsieh ex Y. T. Hsieh

This paper,third in the series of an account of the genus Athyrium in China, presents a revision of the series Epiraches Ching et Y．T．Hsieh ex Y．T．Hsieh．A total of 11 species are recognized from China,and more than 20 names are reduced to synonyms in the present paper．Ser．Epiraches is anatural group in Athyrium,it is distinguished from all the other groups of the genus by its Asplenioid sori and indusia,The series is entirely Asian tropical,subtropical and primarily montane．Its northern limits in central and eastern China are not beyond the Yangtze River．Southwestern part and Taiwan of China and Japan Islands are centers of diversity for the series,each with more than five species．Some speciesare disjunctly distributed in southwest China-Taiwan-Japan;one species(Athyrium roseum) occurs discontinuously between Yunnan and Taiwan;one endemic to Hainan(A. hainanense)is the only representative of Athyrium on the island．All members of this series grow in evergreen broad-leaf forests,generally from 500 to 1800 m above sea level．A key to the species,and illustrations of the middle pinnae from different specimens are provided．     

The Origin,Dispersal and Formation of the Distribution Pattern of Swertia L. (Gentianaceae)

The genus Swertia is one of the large genera in Gentianaceae, including 154 species, 16 series and 11 sections. It is disjunctly distributed in Europe, Asia, Africa and N. America, but entirely absent from Oceania and S. America. According to Takhtajan’s (1978) regionalization of the world flora, Swertia is found in 14 regions. Eastern Asiatic region with 86 species, of which 58 are local endemics, 13 series and 9 sections, ranks the first among all the regions. The highest concentration of the taxa and endemics in Eastern Asiatic region occurs in SW China-Himalayan area (Sikang-Yunnan P. , W. Sichuan, W. Yunnan-Guichou Plateau of China and NE. Burma, N. Burmense P. , E. Himalayan P. and Khasi-Manipur P. ). In this area there are 74 species (48 endemics), 12 series, and 9 sections; thus about half species of the world total, three quarters of series and 82% of sections occur in this small area. Besides, the taxa at different evolutionary stages in Swertia also survive here. It is an indication that SW. China-Himalayan area is a major distribution centre of the genus Swertia. In addition, Sudan-Zambezian Region in Africa, with 22 species, 4 series and 2 sections, is a second distribution centre. The primitive type of the genus Swertia is Sect. Rugosa which consists of 2 series and 23 species. It is highly centred in the mountains of SW. China (Yunnan, Sichuan, Guizhou and SE. Xizang) where 2 series and 16 species occur. Among them 15 species of Ser. Rugosae were considered as the most primitive groups in this genus. From our study, the outgroup of Swertia is the genus Latouchea Frahch. , which is distributed in Yunnan, Sichuan, Guizhou, Hunan, Guangdong, Guangxi and Fujian. The two groups overlap in distribution in SW. China. According to the principle of common origin, the ancestor of two genera ap peared most probably in this overlapping area. It was inferred that SW. China Was the birth-place of the genus Swertia. Four sections of Swertia have different disjunct distribution patterns: Sect. Ophelia is of Tropic Asia, Africa and Madagascar disjunct distribution; sect. Swertia is of north temperate distribution; sect. Spinosisemina is in Tropical Asia (Trop. India to S. China and Philipines); sect. Platynema also is in Tropical Asia (Java, Sumatra, Himalayas to SW. China). These disjunct patterns indicate that the Swertia floras between the continents or between continent and islands have a connection with each other. From paleogeographical analysis, Swertia plants dispersed to Madagascar before the Late Cretaceous, to SE. Asian Islands in the Pleistocene, to North America in the Miocene. The distribution of Swertia in Madagascar might be later than that in Asia. Therefore the origin time of the genus Swertia was at least not later than the Late Cretaceous, and might be back to the Mid-Cretaceous. The genus Swertia first fully developed and differentiated, forming some taxa at different evolutionary stages (Rugosa, Swertia, Poephila, Ophelia and Platynema etc. ) in the original area, and these taxa quickly dispersed in certain directions during the Late Cretaceous-Middle Tertiary when the global climate was warm and no much change. There seem to be three main dispersal routes from the origin area to different continents; (1) The westward route i. e. from SW. China, along the Himalayas area to Kashmir, Pakistan, Afghanistan and Iran, and then southwestwards into Africa throuth Arabia. Four sections (Poephila, Macranthos, Kingdon-Wardia and Ophelia) took this dispersal route. Most species of sect. Ophelia dispersed along this route, but a few along southern route and north ern route. Sect. Ophelia greatly differentiated in Africa and the African endemic sectionSect. Montana was derived from it. The two sections form there a second distribution center of Swertia. (2) The southward route, i. e. towards S. India through the Himalayas, and towards SE. Asian islands through C. and S. China, Indo-China. Along this dispersal route sect. Platynema, Sect. Spinosisemina and a few species of Sect. Ophelia dispersed; (3) The northward rout, i. e. northwards across N. China, C. Asia to a high latitude of Euasia, and also through E. Asia into N. America. The following groups took this route: sect. Rugosa, sect. Swertia, sect. Frasera, sect. Heteranthos and sect. Ophelia ser. Dichotomae. Therefore, it seems that the genus Swertia originated in SW. China and then dispersed from there to N. and S. Asia, Africa, Europe and North America and formed the moderndistribution pattern of this genus.     

椴树属的地理分布

椴树属Tilia是椴树科一个形态特殊且唯一的北温带分布属,分布于亚洲、欧洲和北美,构成典型的北温带分布格局,三个分离的分布区之间缺乏共有种。本文对各分布区的种类进行重新评价,确认全属25种。其中东亚17种,占68％,包含了现存种类各个演化阶段的类群,是现代分布中心;欧洲-西西伯利亚6种,属于木果组及壳果组;北美2种,均为木果组成员。化石分布与现代地理分布格局基本相似,但分布纬度较现代分布偏北,达到北纬80°附近,且还出现于现今无椴树分布的亚洲大陆腹地,北美西部椴树至第三纪末完全绝迹,而东部到第四纪才有化石记录。根据现代地理分布,结合化石证据、地质历史、气候变迁及形态演化推测,椴树属可能在白垩纪晚期起源于中国东部亚热带山地,至少到始新世之前已散布至欧洲和北美西部。渐新世之后的全球降温和更新世大冰期对椴树属现代地理分布格局的形成起着至关重要的作用。     

A Study on the Tribe Astilbeae Miq. (Saxifragaceae)

This paper deals with the phylogeny and geographic distribution of the Tribe Astilbeae. Based on the theory of evolution, the general evolutionary trends of the angiospermous characters and outgroup comparison Penthorum, the polarity of the characters of Trib. Astilbeae is determined. The basic chromosome number (x=8) of Penthorum may be considered as the primitive one; x = 7 (8-1 ) of Astilbe and x = 17 (8 + 9) of Astilboides have evidently evolved from Penthorum, whereas x = 15 (7 + 8) of Rodgersia is derived from x =7 and x = 8 through hybridization. The striate reticulate ornamentation of pollen is more advanced than the striate. The Axile placentation is more primitive than the parietal. Carpels, stamens, petals and sepals have evolved from more to fewer, then to absent in number.Veins of sepals have evolved from uninerviate and pinnate together (or uninerviate) to pinnate and arcuat together, then to arcuat. Leaves have evolved from simple to simple and compound together, then to compound. A schema showing phylogenetic tree of Trib. Astilbeae and its outgroup Penthorum is given according to a cladistic analysis using the method of maximal same steps by Xu (1989). In the schema, Rodgersia and Astilbaides is a monophyletic group having a sister group Astilbe, Rodgersia is more advanced than Astilbe; Astilboides is intermediate; while Penthorum is the most primitive and had a common ancestor with its sister group, The Trib. Astilbeae consist of three genera. Astilbe, Astilboides and Rodgersia. Based on the present authort s revision, this tribe comprises 24 species and 13 varieties (excl. typical ones) in the total. Takhtajan's (1986) Eastern Asiatic Region has 32 species and varieties (all of them are endemic) in three genera, ranking the first; the Malesian Region has three species (endemic) in one genus; the North Amerian Atlantic Region has two species (endemic) in one genus; the Irano-Turanian Region has only one species, ranking the last. In the Eastern Asiatic Region, Japan, Korea and eastern Jilin-Liaoning of China (The main part of this area is Takhtajan's Japanese Korean Provine) has 17 species and varieties in three genera, which constitute 45.9% of the total in Trib. Astilbeae and include the groups at different evolutionary stages and 15 endemic species and varieties, of which Astilbe platyphylla, A. simplicifolia and Rodgersia podophylla may be considered as the primitive ones. However, the Hengduan Mountains region (west Sichuan and northwest Yunnan) has 11 species and varieties (10 occur in west Sichuan, 9 in northwest Yunnan) in two genera, which constitute 29.7% of the total number of species and varieties in Trib. Astilbeae and include 5 endemic species and varieties. From the above mentioned the author suggests that the centre of origin, present distribution and differentiation of Trib. Astilbeae should be in Japan, Korea and eastern Jilin-Liaoning, while the Hengduan Mountains region should be another centre of present distribution. The more advanced species, i. e. Astilbe biternata, A. crenatilobata, A. philippinensis, A. a 'poensis, A. indica, A. khasiana and Rodgersia nepalensis are found in the area far from the centre of origin. Thus the Trib. Astilbeae plants may have migrated from Japan, Korea and eastern Jilin-Liaoning northwards and then southeastwards to southeastern North America through eastern Siberia and the Bering bridge, southwards to the philippines and Java through South China, and southwestwards to the Himalayas through the Qinling-Daba Mountains and the Hengduan Mountains. At present, in eastern Siberia and most parts of North America there are no plants in the Tribe Astilbeae, Which may be explained by their extinct ness there during the Quaternary glaciation. Both Astilbe and Rodgersia are distributed in the Asian continent and Japan. Japan has been isolated from the Asian continent since the late Tertiary, so that both Astilbe and Rodgersia had undoubtedly occurred before Japan was isolated (Early Tertiary). Therefore, the origin time of the Tribe Astilbeae may be considered as the Early Tertiary or may be traced back to the Late Cretaceous. The pollen morphology of Penthorum sedoides L., Astilbe grandis Stapf ex Wils. andAstilboides tabularis (Hemsl.) Engl. was examined under SEM and is shown in Plate 1. Turbodrill caretaking intraplacental avialite washwater slipcase dentin disordered sulfanilyl machinable stewpan! Netherward pressbodies horror abscissa, keratosis frieze. 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