When the pond turtle followed the reindeer: effect of the last extreme global warming event on the timing of faunal change in Northern Europe

Faunal communities have been shaped in different ways by past climatic change. The impact of the termination of the last Glacial and the onset of the present (Holocene) Interglacial on large‐scale faunal shifts, extinction dynamics and gene pools of species are of special interest in natural sciences. A general pattern of climate‐triggered range expansion and local extinction of vertebrate species is known for Europe, and shows that in the modern temperate zone the main faunal change took place mainly during the Late Glacial (14 700–11 700 years ago) and Early Holocene (11 700–9 100 years ago). Based on large datasets of new radiocarbon data, we present precise temporal dynamics of climate‐driven disappearance and appearance of reindeer and pond turtle in southern Sweden. These two species are significant climate indicators in Late Quaternary biostratigraphy. Our data reveal that the reindeer disappeared from southern Sweden ca. 10 300 years ago, whereas the pond turtle colonized the area ca. 9 860 years ago, with a 450‐year gap between each species. This provides evidence for a sudden environmental turnover, causing the replacement of an arctic faunal element by a thermophilic species. The postglacial range dynamics of pond turtle and reindeer are a unique model case, allowing insights into the faunal turnover of other vertebrates during the last dramatic natural global warming event at the Pleistocene–Holocene transition.     

Cladocera response to Late Glacial to Early Holocene climate change in a South Carpathian mountain lake

This study explores changes in cladoceran composition in a high mountain lake of the Retezat (Lake Brazi), the South Carpathian Mountains of Romania, during the Late Glacial–Early Holocene (14,500–11,600 cal. yr. bp) transition using a paleolimnological approach. The lake had a species poor cladoceran community throughout this period. Daphnia longispina, Chydorus sphaericus and Alona affinis were the most common, showing marked fluctuations in their relative abundances through time. Distinct faunal response to warming at the Younger Dryas (YD)/Preboreal transition was recorded by increasing fossil densities and distinct community composition change: Alona affinis became dominant while numbers of Chydorus sphaericus dramatically decreased. In the Early Holocene, the productivity of Lake Brazi seem to have increased as reflected by higher numbers of Cladocera due to appearance of new species (Alona rectangula, A. quadrangularis and A. guttata) which are common in productive waters. Significant negative correlation was found between average dorsal length of daphnid ephippia and the NGRIP ^δ18O isotope values. Given the absence of fish predation, changes in Daphnia ephippia size were taken to indicate climatic change: larger ephippium size inferred cold conditions during the Late Glacial, while smaller size reflected climate warming during the Early Holocene. We conclude that Cladocera fossils are good indicators of climatic change that happened during the transition from the Late Glacial to the Holocene. We found that climatic conditions can be tracked either by size distribution of Daphnia ephippia (larger ephippium size under colder climate) and/or by community change of cladocerans.     

根据孢粉分析论青藏高原西部和北部全新世环境变化

通过对青藏高原西部班公湖钻孔和北部中昆仑山３个湖相剖面的孢粉研究,揭示青藏高原西部和北部地区全新世１万年期间植被的演替和气候变化,西部在全新世早期９９００－７８００ｙｒ．Ｂ．Ｐ．植被由荒漠转为草原,气候好转;中期７８００－３５００ｙｒ．Ｂ．Ｐ．,草原发展,气候较适宜,以７２００－６３００ｙｒ．Ｂ．Ｐ．为高温湿期,５５００ｙｒ．Ｂ．Ｐ．和３５００ｙｒ．Ｂ．Ｐ．出现干旱,晚期从３５００ｙｒ．Ｂ．Ｐ．至今植被为荒漠,气候干旱,其中７００ｙｒ．Ｂ．Ｐ．气候恶化。北部地区全新世时期为荒漠植被,当气候温湿时,蒿和禾本科,莎草科成份增加,藜科减少,气候干旱时则相反,北部全新世的气候分期和干湿波动与西部相近,两地在晚期气候朝干旱化发展。     

Testate amoebae record from the Laptev Sea coast and its implication for the reconstruction of Late Pleistocene and Holocene environments in the Arctic Siberia

Our study aims to look for testate amobae (TA) in the surface and fossil sediments from the Cape Mamontov Klyk (73°60′–73°63′ N, 116°88′–117°18′ E), southern coast of the Laptev Sea, ca 150 km west of the Lena Delta and to discuss the potential of TA analysis for Glacial/Interglacial environmental reconstructions in Arctic Siberia. The radiocarbon age determination suggests that the studied sediments accumulated between ca 45,000 ¹⁴C yr BP and present. A total of 98 TA taxa were identified in the 10 recent surface and 59 fossil samples. Results of taxonomical identification and ecological analysis of TA in the modern and fossil samples suggest that major changes in the soil moisture conditions took place. Our results show that soil-living taxa dominated the TA assemblages at the study sites during the past 45,000 years. The environmental conditions of the study area were most favourable (relatively warm and humid) during the Kargin Interstadial (ca 45,000–25,000 ¹⁴C yr BP). An opposite situation is reconstructed for the Sartan Stadial (ca. 25,000–15,000 ¹⁴C yr BP). During the Kargin Interstadial, optimum conditions occurred between ca 44,000 and 40,000 ¹⁴C yr BP characterised by highest TA abundances and taxa diversity. This initial optimal phase was followed by the interval with drier and colder conditions about 40,000–30,000 ¹⁴C yr BP. The sediments dated between ca. 24,000 and 18,000 ¹⁴C yr BP show low TA abundances and diversity, in agreement with the much colder and drier environments during the maximum phase of the Last Glacial. The onset of the Holocene is indicated by a broad representation of obligate hydrophilic taxa, especially from genus Difflugia, suggesting wet and relatively warm conditions. By comparison with other environmental proxies used in the studied sections as well as from the neighbouring arctic regions our results suggest that TA analysis can provide valuable information, contributing to the better understanding of the Late Quaternary climate and environments in Arctic Siberia.     

Early Holocene environmental change in the Kreekrak area (Zeeland, SW-Netherlands): A multi-proxy analysis

Detailed botanical (microfossil and macroremain), zoological and geochemical analyses (major and trace elements including C, Al, S, Ca, Fe, P, As, Zn, U, Ba and Rare Earth Elements) of organic deposits provide new insights into Early Holocene environmental change in the Kreekrak area (southwestern Netherlands). The age assessment of the record is based on high resolution AMS ¹⁴C wiggle-match dating (WMD). For the first time an AMS ¹⁴C WMD based chronology covering the Late Glacial/Holocene transition and early Preboreal is introduced for a site in The Netherlands.The Kreekrak botanical record reflects the end of the Younger Dryas to early Boreal and can be well correlated with pollen records from other sites in The Netherlands and Belgium. The palaeo-topography showed that the Kreekrak deposits formed in an abandoned channel of the River Schelde. Around ca. 11,490 cal BP, at the end of the Late Glacial/Holocene transition, infilling of the lake started with predominantly organic deposits in slowly running water. As a result of the warmer climate the area became forested with birch and poplar during the Friesland Phase (ca. 11,490-11,365 cal BP). Biological productivity of the lake and its surroundings increased. Aquatic vegetation developed in the lake, while shrubs of willow, reed swamps and grasslands fringed the shores. Precipitation increased, which caused a rise in the lake water table and an increase in the supply of oxic surface (= river) water into the Kreekrak lake. During this period, the Kreekrak lake was fed by inflowing river water, run-off, precipitation and seepage of Fe-rich groundwater. Around ca. 11,435 cal BP the water became stagnant probably as result of a total cut-off of the river channel. Inflow of river water ceased, while the supply of reduced Fe-rich groundwater became dominant. During the Rammelbeek Phase (ca. 11,365-11,250 cal BP), the climate was more continental and the abundance of grasslands and open herbaceous vegetation increased. Biological productivity remained high. In the lake, the supply of Fe-rich groundwater continued, the water level slightly decreased but aquatic vegetation remained present. At the end of the Rammelbeek Phase a sudden reduction in the supply of Fe-rich reduced groundwater caused a lowering of the groundwater level in the area, resulting in the development of a hiatus. Due to this hiatus, the Late Preboreal (11,250-10,710 cal BP) is absent from the record. During the early Boreal (10,710-10,000 cal BP) the landscape became densely forested and accumulation of peat in the former lake resumed due to a slowly rising groundwater level. The Boreal was a relatively stable period with low sedimentation rates.The combination of palaeobotanical and geochemical analyses in the Kreekrak record shows a close interrelation between landscape development and geochemistry. It appears that the environmental development of this area during the Late Glacial/Holocene transition and Early Holocene was largely influenced (directly or indirectly) by major climatic changes that occurred during this period, which determined local phenomena such as the composition and density of the vegetation, occurrence of seepage and river activity. Further research of this type has the potential to develop the application of major- and trace element geochemistry in palaeoenvironmental reconstructions.     

Contribution to the lithostratigraphy and history of Lake Ladoga

New geological data from Lake Ladoga are presented and an interpretation of the Late Pleistocene and Holocene history of the lake is given. The deglaciation of the southern part of the lake and an origin of the first ice margin Lake Ladoga took place at ca. 14,000 yr B.P.     

Evidence for Late Glacial and Holocene environmental changes from subfossil assemblages in sediments of Lake Neuchâtel, Switzerland

Ostracod and diatom assemblages, mollusc remains, plus pollen and sediment stratigraphy from three cores taken at sites 40, 80 and 135 m deep in Lake Neuchâtel, Switzerland, show distinct changes since deglaciation. Shifts are interpreted in terms both of climatic change and major changes in the catchment input caused by the Aar River entering or bypassing Lake Neuchâtel. Three palaeoenvironmental phases are recognized: (1) the Late Glacial, with the inflow of the river before 10,000 yr B.P.; (2) the early to mid-Holocene, with temporary bypassing of the river between 10,000 and 4850 yr B.P.; and (3) the mid- to late Holocene, without river input after approx. 4850 yr B.P. During periods with river input, rhythmites were deposited. Ostracods indicate a well oxygenated hypolimnion, suggesting deep ventilation enhanced by the river inflow. The diatom assemblage indicates eutrophic to mesotrophic conditions and is dominated by generally well preserved periphytic species. The high number of unidentifiable pollen suggests oxidation during transport in the river. During periods without river input, a massive calcareous silt with abundant authigenic calcite was deposited, diatoms indicate a change to mesotrophic and oligotrophic conditions, and decreased sedimentation rates and higher alkalinity enhanced the dissolution of diatoms. Terrestrial and littoral molluscs as well as periphytic diatoms indicate erosion of the shore when the lake level rose or fell with onset or offset of the river inflow. Before 12,600 yr B.P., the patchy vegetation cover led to intensive soil erosion and high sedimentation rates. The lake was still frozen during the winter months. After approx. 12,600 yr B.P. the sediment changed from clastic rhythmites to thinly bedded, non-glacial rhythmites, the vegetation cover became denser, the abundance of ostracods increased, and molluscs started to colonize the profundal zone of the lake. The Younger Dryas (approx. 10,700–10,000 yr B.P.) displays maximum abundance of ostracods, indicating deep ventilation. The transition to the Holocene is characterized by a change in the ostracod assemblage and the river bypassed the lake between approximately 10,000 and 8900 yr B.P. When the river reentered the lake after approximately 8900 yr B.P., a new ostracod species appeared. After two short periods of river bypassing between approximately 7200 and 6950 yr B.P., diatoms indicate a higher trophic level about 6000–5000 yr B.P. associated with higher erosion rates and increased nutrient input due to Neolithic agricultural activities. After approximately 4850 yr B.P. the river bypassed the lake again and the trophic level decreased. In contrast to the early to mid-Holocene periods of decreased oxygen supply, the mid- to late Holocene is characterized by deep ventilation. This is attributed to the northward retreat of the polarfront and enhanced westerlies consequent to decreased insolation after 6000 yr B.P. The change of the river flow pattern correlates with glacier oscillations and may suggest a climatic control. The final cessation of the inflow is attributed to diversion of the river consequent to isostatic uplift and late Neolithic clearance activity.     

Late Glacial and Holocene Ostracoda of the Gulf of Gdańsk,the Baltic Sea,Poland

To trace environmental changes in water hydrology and salinity in the Late Glacial to Holocene of the Gulf of Gdańsk, a south‐eastern bay of the Baltic Sea within the maritime zone of Poland, the distribution of ostracod valves was studied in 20 sediment cores collected from both the shallow‐ and deep‐water zones (depth 10.9–67.5 m). The studied sediment sequences yielded ca. 3000 valves of 21 ostracod species, of which only five are known to live today in the Gulf, which has a present maximum depth of 118 m and water salinity up to 7–8‰. The majority of the studied sediment layers that contained ostracod valves corresponded to the period of the Late Glacial to Mid‐Holocene and was dominated by non‐marine species, of which the most common were Candona neglecta (present in 17 cores), Cytherissa lacustris (15 cores) and Candona candida (14 cores). By clustering classification five major ostracod assemblage types were recognised in the studied cores. Initial assemblage types dominated mostly by inhabitants of the profundal/sub‐littoral zones of modern oligo‐mesotrophic lakes (C. lacustris and C. neglecta) in some sediment sequences were replaced in stratigraphical order by the assemblages dominated by brackish‐water species (Cyprideis torosa or Cytheromorpha fuscata). The structure and species composition of the distinguished ostracod assemblage types as well as their successional transitions indicate that the studied sediments were deposited initially in the Late Glacial in freshwater lacustrine conditions, and subsequently, during the Holocene marine transgression, covered by marine sands. The present results confirm and consolidate inferences based on previously published data on ostracods from the western part of the Gulf of Gdańsk as well as on other biotic (molluscs, diatoms) and abiotic (seismoacoustic) indices from this area (© 2012 WILEY‐VCH Verlag GmbH & Co. KGaA, Weinheim)     

Holocene persistence of wooded steppe in the Great Hungarian Plain

Aim We used a combination of new and previously published palaeoecological data to test three hypotheses: (1) that wooded steppe persisted in the Great Hungarian Plain throughout the Holocene; (2) that wooded steppe and steppe were most extensive between c. 9900 and 8300 cal. yr bp (the ‘Boreal steppe’ period); and (3) that Southern Continental, Pontic and Eastern Sub‐Mediterranean steppe species reached the region during the early Holocene via the ‘Lower Danube Corridor’. Location Sarló‐hát oxbow lake, Hungary and the Eastern European wooded steppe zone. Methods Holocene sediments deposited in the Sarló‐hát oxbow lake were subjected to pollen and microcharcoal analyses. Twelve radiocarbon age estimates were obtained to determine sediment chronology. In addition, previously published palaeoecological data from the Great Hungarian Plain were compiled, analysed and compared with previous studies in other regions of steppe and wooded steppe in eastern Europe. Results Palynological data from two sediment cores extending to c. 11,400 cal. yr bp indicate the persistent dominance of the landscape by temperate deciduous wooded steppe throughout the Holocene, although with varying canopy composition. Warm‐continental steppe grasslands and saline tall‐grass meadows developed on edaphically constrained areas, which remained steppe‐dominated throughout the Holocene. The extent of steppe grasslands did not increase between 9900 and 8300 cal. yr bp . After c. 3100 cal. yr bp , anthropogenic activities led to the development of cultural steppe. Thermophilous steppe species of the Southern Continental, Pontic and Sub‐Mediterranean floristic elements probably reached the Great Hungarian Plain principally via the Lower Danube Corridor during the late glacial interstadial and Holocene. Eurythermic members of these elements, however, probably survived the Last Glacial Maximum in favourable microsites, extending their ranges during the Holocene from these local sources. Main conclusions Our results confirm the Holocene persistence of wooded steppe in the Great Hungarian Plain, disprove the ‘Boreal steppe’ theory, and suggest an Early Holocene period of greater vegetation openness between 11,400 and 9900 cal. yr bp . Evidence for the post‐glacial immigration of south‐eastern steppe elements into the Carpathian Basin is equivocal: the last glacial/interglacial presence of several southern steppe species suggests that the Hungarian Plain hosted suitable habitats for them during warm and cold phases alike.     

Vegetation development and human occupation in the Damascus region of southwestern Syria from the Late Pleistocene to Holocene

Rockshelter Baaz in the Damascus region of Syria provided a variety of botanical remains from the Late Pleistocene and Early Holocene period. These remains provide new information about the vegetation evolution in this region. The earliest occupational levels correspond with a moisture peak during the Late Pleistocene, between ca. 34–32 kyr b.p., when pine expanded. The next occupations took place during extreme arid conditions, ca. 23–21 kyr b.p., and probably during the Last Glacial Maximum when a steppe vegetation was established. The occupation level of the Younger Dryas, represented by Natufian remains, suggests that the area had been covered by almond-pistachio steppe, similar to later periods of the Early Holocene, and was probably located just outside the range of dense wild cereal stands. There is no drastic impact of the Younger Dryas visible on the vegetation in the botanical remains. The lack of fruits and seeds at Baaz indicates that the site was more likely to have been a temporary hunting post rather than a plant processing site for much of its history. It is ideally suited to this purpose because of its location over the Jaba′deen Pass and the associated springs. However, archaeological remains from the Natufian period, suggest that the site was more permanently occupied during this time. A. Dodonov deceased.     

青藏高原东南部18ka以来气候与水文变化

青藏高原东南部众多封闭小湖的湖相沉积记录着重要的地理信息,是恢复当地晚更民以来古环境演变的理想材料。西藏海登湖和仁同分辨率的花粉组合,ＡＭＳ＾１４Ｃ测年及磁化率和有机质含量测定,揭示了研究区与鳊季风密切相关的古植被,古气候和古水文变化历史。高分辨率的花粉记录反映,在１６ｋａＢｍ,Ｐ,以前花粉组合以黎科（Ｃｈｅｎｏｐｏｄｉａｃｅａｅ）和蒿属（Ａｒｔｅｍｉｓｉａ）为主,花粉浓度很低,应为荒漠草原植被。     

Mitochondrial DNA analysis shows a Near Eastern Neolithic origin for domestic cattle and no indication of domestication of European aurochs

The extinct aurochs (Bos primigenius primigenius) was a large type of cattle that ranged over almost the whole Eurasian continent. The aurochs is the wild progenitor of modern cattle, but it is unclear whether European aurochs contributed to this process. To provide new insights into the demographic history of aurochs and domestic cattle, we have generated high-confidence mitochondrial DNA sequences from 59 archaeological skeletal finds, which were attributed to wild European cattle populations based on their chronological date and/or morphology. All pre-Neolithic aurochs belonged to the previously designated P haplogroup, indicating that this represents the Late Glacial Central European signature. We also report one new and highly divergent haplotype in a Neolithic aurochs sample from Germany, which points to greater variability during the Pleistocene. Furthermore, the Neolithic and Bronze Age samples that were classified with confidence as European aurochs using morphological criteria all carry P haplotype mitochondrial DNA, suggesting continuity of Late Glacial and Early Holocene aurochs populations in Europe. Bayesian analysis indicates that recent population growth gives a significantly better fit to our data than a constant-sized population, an observation consistent with a postglacial expansion scenario, possibly from a single European refugial population. Previous work has shown that most ancient and modern European domestic cattle carry haplotypes previously designated T. This, in combination with our new finding of a T haplotype in a very Early Neolithic site in Syria, lends persuasive support to a scenario whereby gracile Near Eastern domestic populations, carrying predominantly T haplotypes, replaced P haplotype-carrying robust autochthonous aurochs populations in Europe, from the Early Neolithic onward. During the period of coexistence, it appears that domestic cattle were kept separate from wild aurochs and introgression was extremely rare.     

Late quaternary pollen records from two Ugandan mires: evidence for environmental changes in the Rukiga Highlands of southwest Uganda

High resolution pollen data from two mires in the Rukiga Highlands, southwest Uganda (Ahakagyezi Swamp and Muchoya Swamp) provide detailed evidence of the environmetal history of the region since ca. 42 000 yr B.P. According to these data, there a was period pre ca. 42 000 yr B.P.>. when climatic conditions were similar to the present. Conditions considerably colder and drier than the present occurred during two periods in the past (pre ca. 42 000 yr B.P. to ca. 32 000 yr B.P., and ca. 21 000 yr B.P. to ca. 12 000 yr B.P.). The most recent period of climatic improvement began ca. 14 000 yr B.P. Moist lower montane forest colonised the Muchoya catchment by ca. 11 100 yr B.P. and the Ahakagyezi catchment by ca. 10 600 yr B.P. indicating the establishment there of climatic conditions broadly similar to the present.

Pollen data from both mires reveal forest disturbances beginning ca. 2200 yr B.P. This disturbances is likely to have been human-induced.     

Evidence of past climatic change in the Namib Desert

A preliminary survey of some areas of the Central and Northern Namib Desert was undertaken in order to identify geomorphic and sedimentary features that could potentially provide information on past climatic change in the region. Wherever possible, samples were collected for radiocarbon dating. The visits to the Skeleton Coast area were especially revealing: practically every river valley in this tract shows a sequence of terraces and deposits that would warrant a detailed study. In all, about 120 new radiocarbon dates were obtained for the region. These have supplied a wealth of individual results and, collectively, have enabled the construction of a tentative chronological sequence of the changes in climate for the past 40,000 yr. Three periods of increased aridity in the hinterland, but with decreasing intensity are postulated: during the Last Glacial Maximum, the early Holocene, and between A.D. 1200 and A.D. 1600. The Inter-Pleniglacial up to 23,000 yr B.P., the Late Glacial, and the period between about 4000 yr B.P and 1200 yr B.P., on the other hand, were more humid than today, also with decreasing intensity. Other indicators of past climate were also identified, but since they lie beyond the range of radiocarbon dating, they cannot as yet be placed in a chronological context. Of significance is the fact that convincing evidence was obtained for climatic change within the last millennium and, especially, the rainfall is found to have been increasing since the 16th century.     

Upper quaternary vegetational and climatic sequence of the fuquene area (Eastern Cordillera,Colombia)

In this paper the results of a pollen-analytical study of three bore holes of lake deposits in the Fuquene-Valle de Ubaté area, Colombian Cordillera Oriental (5° N; elev. 2,580 m) are given. By means of ¹⁴C dates and comparison with other pollen diagrams and dates from the Eastern Cordillera, it was possible to correlate the local pollen zones and chronostratigraphical units with the European chronostratigraphical sequence. The longest diagram (Fuquene II) represents some 30, 000 years, including part of the Middle Pleniglacial, the Upper Pleniglacial, the Late Glacial and the Holocene.During the Middle Pleniglacial, Polylepis wood was an important constituent of the vegetation. During climatic extremes of the Upper Pleniglacial an open páramo vegetation existed.During the main part of the Upper Pleniglacial (from ca. 21,000 to ca. 13,000 B.P.) the lake level was low and the climate dry. The lake level rose again in the beginning of the Late Glacial, and the area around the lake became forested. At the beginning of the El Abra Stadial (Late Dryas Stadial) there was a marked cooling and the lake became lower once more; this had a great impact on the vegetation which became partly open again. The beginning of the Holocene is marked by a gradual increase of forest elements, especially of Quercus (oak.) Oak forests dominated the area during the greater part of the Holocene. During the “hypithermal”, elements growing today at a lower level, such as Cecropia, Acalypha (and possibly Alchornea), must have been growing in the area, intermixed with the oak forest. At the same time there was a considerable increase of Urticaceae in the undergrowth of these forests. The vegetation zones were probably situated several hundreds of metres higher than they are at the present time.The influence of man on vegetation had started already before the beginning of our era, but there is a sudden increase of this influence (e.g., increase of Dodonaea, decrease of Quercus and Urticaceae) somewhere around 2,000 B.P. Later on, the diagram shows the rapid ultimate decline of the forest elements and an increase of Gramineae. Today nothing is left of the original oak forests and their place is taken by arable land or by secondary plant communities with low brushwood.In the lake sediments, thin layers of volcanic ash were found dated approximately 9,000 B.P., 11,000 B.P., and 21,000 B.P., respectively; a series of three ash-layers was deposited between approximately 22,000 B.P. and 26,000 B.P. and one possibly at approximately 30,000 B.P. These layers have been found in several other sections in the Eastern Cordillera and are the basis for a tephrochronology.     

When east met west: the sub-fossil footprints of the west European hedgehog and the northern white-breasted hedgehog during the Late Quaternary in Europe

The distinct distribution of the west European hedgehog Erinaceus europaeus and the northern white-breasted hedgehog Erinaceus roumanicus and their separate refugial origins after the Pleistocene is a well-known example in the zoogeography of the Holarctic. Among the Late Quaternary faunal assemblages, the west European hedgehog is recorded at 269 sites whereas the northern white-breasted hedgehog is recorded only at 52 sites in Europe. The distribution patterns of the temporal and spatial Glacial records of the west European hedgehog show a general trend: a strong restriction to glacial refugia (the Iberian and Italian Peninsulas) during the Weichselian Glacial until the end of the Last Glacial Maximum, and a colonization of southern France during the early Late Glacial between 14 000 and 125 00 ¹⁴C years BP (15 000–12 800 cal. BC). Whereas the British Isles could have already been colonised by the end of the Pre-Boreal, in the rest of Central Europe E. europaeus was clearly distributed there in the Boreal for the first time. The west European hedgehog is an absolute Holocene faunal element in Central Europe. It appears in most parts of Central Europe during the Early Holocene, when the west European hedgehog met its eastern relative, which probably was similarly sensitive. After meeting each other, the distribution limit of both Erinaceus species in Central Europe seems to have been relatively constant in its geographic extent. Because of the clear climatic correlation, E. europaeus should be considered as an indicator species for temperate climatic conditions of the Holocene fauna. This should be considered during the reconstruction of climatic conditions with the help of the analysis of quaternary faunal material.     

Earliest Holocene vegetation history and island biogeography of the Isle of Man, British Isles

Aim To present radiocarbon dated early Holocene pollen analytical data from two sites on the northern plain of the Isle of Man and to discuss the implications of the vegetation history in relation to severance of the island from the British Isles and to identify further evidence for divergent biogeographical development previously exemplified by the survival and apparent dwarfism of late glacial Megaloceros giganteus (Giant Deer). Location The Isle of Man, British Isles. Methods Pollen analysis and AMS radiocarbon dating of late glacial to early Holocene lake sequences at Pollies and Curragh‐y‐Cowle on the northern plain of the Isle of Man. Results The pollen data indicate a prolonged period of pre‐woodland vegetation after the late Glacial/Holocene transition, which lasted for most of the first post‐glacial millennium. This persistence of pre‐forest environments meant that the expansion of Betula woodland occurred later in this part of the Isle of Man than in adjacent areas of Britain and Ireland. Conclusions The Isle of Man, in the northern Irish Sea, was isolated from Britain during the late Glacial period perhaps explaining the delayed arrival of tree species. Delayed rise of the Holocene forest compared with surrounding regions probably reflects severance of the land‐bridge with Cumbria, but also could be a function of climate changes during the early Holocene and local environmental conditions. Late survival and the dwarfism of the Megaloceros giganteus (Giant Deer) fauna is another example of biogeographical divergence during the early Holocene/late Glacial of the Isle of Man. The delayed afforestation and absence of human hunters in the Manx early Holocene offers a permissive environmental context for the as yet unproven survival of Megaloceros into the early Holocene.     

Vegetationsverhältnisse und Naturumwelt des Beckens Jestřebská kotlina am Ende des Spätglazials und im Holozän (Doksy-Gebiet)

The method of pollen analysis was used to study a profile sample taken from the Jest?ebské blato mire, the Doksy District (northern Bohemia). The aim of the palacobotanical analyses was to perform a reconstruction of vegetation and environment for geobotanical and archaeological research purposes. Sediments dating back to the end of the Late Glacial and to both the Early and Middle Holocene were pollenanalytically studied. The results of the pollen analyses recorded the development of vegetation during the end of the Late Glacial, Preboreal, Boreal and Atlantic Periods. Parkline pine stands of the end of the Late Glacial developed to closed pine forests, during the Preboreal. Even during the Boreal, the dominant pine formed forest communities were typical of the area under study, though trees with greater demands on temperature and moisture conditions continued to penetrate the existing communities. During the Lower and the first half of the Upper Atlantic Period, spruce forests and alder woods formed the dominant forest communities in mire. Nevertheless,Pinus was still dominant on extreme sites (both from the edaphic and geomorphological points of view). An early occurrence ofAlnus andPicea is another characteristic feature of the studied area. The conclusions drawn from pollenanalytical results were used for the reconstruction of the past environment of the Mesolithic and Neolithic Ages.     

The Quaternary eolian sequence of Madeira: stratigraphy, chronology, and paleoenvironmental interpretation

A thick (ca. 40 m) sequence of coastal eolian sediments occurs on a narrow peninsula on the eastern end of the island of Madeira, located in the Eastern Atlantic at 33°N latitude. The sediments consist of black volcanic sands (with or without bioclasts) as well as clay units up to 2 m thick. A series of inceptisols (Eutrochrepts) and one alfisol (a Hapludalf) are developed in these sediments. Land snail shells and secondary carbonates, in the form of well-developed rhizoliths, calcretes, fissure-fills, and soil nodules, are present in abundance. The chronology of the sequence was determined by ¹⁴C and U---Th analyses of land snail shells and secondary carbonates and amino acid epimerization analysis of land snail shells. All sediments, including the clay units, are originally of eolian origin, derived from the beach to the south of the deposit, but some have been redeposited by colluviation. Temporal variation in the lithology of the sediments relates to variations in sea-level, with black sands being deposited during lower sea level stands and clays at the lowest. It is suggested that fine marine sediments, exposed during low sea-level stands, may also be the dominant source of silty or clayey units in other coastal eolian deposits in the subtropical Atlantic and Mediterranean.

The sequence spans from 200,000–300,000 years ago up to the 20th century. Sedimentation was discontinuous and often rapid; erosional hiatuses are present. During the Holocene, eolian sands started accumulating at 8200 yr B.P. during a transgressive phase and stopped at 4500 yr B.P. as sea level approached its present height. Colluviation increased dramatically following the first human settlement of the island in the 15th century and continued up to the 20th century, as dated by amino acid epimerization analysis of land snails. Earlier periods of colluviation were identified from the age distribution of land snail shells redeposited in younger colluvium.

Paleoenvironmental reconstruction was based mainly on soil and sediment features (including rhizolith morphology) and land snail faunas but also on stable isotope variations (¹³C, ¹⁸O) in land snails and secondary carbonates, pollen (generally not well preserved), and phytoliths. Most of the portion of the Middle Pleistocene represented in the sequence was characterized by moderately dry conditions, in comparison to the late Pleistocene and Holocene. During the last interglacial, relatively wet conditions occurred, wetter than during the Holocene interglacial. Moderately moist conditions were present during the accumulation of the thick unit dating to ca. 80,000 yr B.P. As sea level fell subsequent to this period, conditions appear to have become drier. Starting ca. 50,000–55,000 yr B.P., conditions were especially wet, but prior to the last glacial maximum, markedly arid conditions ensued. Toward the end of the last glacial, wet conditions returned and produced the best-developed soil preserved in the sequence. Moderately moist conditions occurred during the early to middle Holocene but apparently become slightly drier after 4500 yr B.P. The impact of human settlement can be seen in the loss of woody vegetation and enhanced gullying and colluviation during the last ca. 500 years.     

Terrestrial Biomass-Reconstruction for the Pleniglacial and the Holocene Climatic Optimum Around 7000 cal yr BP in Europe

The aim of this research was to estimate the former below- and aboveground biomass during the Last Pleniglacial (22,000 cal yr BP) and during the Holocene climatic optimum, the Atlantic Period (7,000 cal yr BP). Vegetation distributions and soil conditions during these periods were reconstructed according to published literature. Using the present-day bio-masses of similar (but not identical) vegetation and soil types as analogues, estimates of the terrestrial biomass were made for 22,000 and 7,000 cal yr BP. Biomass was low during the Pleniglacial, representing only 5.0 to 12.5 Gt. This reflects weak soil development under cold climatic conditions, combined with soil erosion due to severe frost and strong wind. The results are compared with other biomass reconstructions for the Pleniglacial. The values for terrestrial biomass of the Pleniglacial presented in this paper are much lower than the results of the various studies for this time slice. These differences are likely to be related to the method applied. Biomass was high during the Holocene climatic optimum around 7,000 cal yr BP, representing 261 to 325 Gt compared to 232 to 273 Gt in the present potential situation and 165 to 197 Gt in the present-day real situation. This reflects intensive soil development under warm and moist conditions during the climatic optimum of the Holocene. These results demonstrate the fluctuation of C-content in terrestrial ecosystems since the Last Glacial Maximum on the European continent.