Taxonomic composition and zoogeographic characteristics of the flea fauna (Siphonaptera) of Russia

255 species and 59 subspecies of fleas from 55 genera of 7 families are known from Russia, which is 30% of the Palaearctic fauna. Additionally, over 187 species of 47 genera from 7 families are known from the neighboring territories of Central and Southern Europe, Transcaucasia, Kazakhstan, Middle Asia, Mongolia, Northeast China, and Japan. 13 species of 12 genera are known only from Russia. Noteworthy is the low percent of endemic species (not more than 4%) and genera (one genus) in the Russian fauna. The principal centers of taxonomic diversity in the Palaearctic, including many endemic species and genera, lie in the Eastern Asian, Central Asian, and Turano-Iranian Subregions, outside Russia and the Euro-Siberian Subregion. The bulk of the Russian fauna is formed by the species and genera of the three largest flea families: Hystrichopsyllidae, Ceratophyllidae, and Leptopsyllidae. The family Ceratophyllidae has the greatest number of genera in the Russian fauna, and Hystrichopsyllidae, the greatest number of species. Western (Western and Western-Central Palaearctic; 84 species from 41 genera of 7 families) and Eastern (Central-Eastern and Eastern Palaearctic; 78 species from 42 genera of 6 families) species are nearly equally represented in the Russian fauna.     

The Palaearctic centers of taxonomic diversity of fleas (Siphonaptera)

The Palaearctic flea fauna includes 921 species and 479 subspecies from 96 genera of 10 families. Of them, 858 species (94%) from 43 genera are endemic to the Palaearctic; they comprise 40% of the Palaearctic Hystrichopsyllidae, 24% of Ceratophyllidae, and 20% of Leptopsyllidae. Ranges of 581 species (63% of the Palaearctic fauna) are situated within one province or subregion of the Palaearctic. Species with ranges including a part of Asia (592) comprise 87% of the total fauna; 72% of the species (517) are endemic to the Palaearctic. The largest centers of taxonomic diversity of Palaearctic fleas are situated in the East Asian, Central Asian, and Turano-Iranian Subregions: 320 species of fleas (214 of them endemic) from 59 genera (8 endemic) are known from the East Asian Subregion; 270 species (over 120 endemic) from 54 genera (5 endemic) are distributed in the Central Asian Subregion. The Turano-Iranian fauna comprises 213 species (103 endemic) from 47 genera (3 endemic); about 160 species occur in the Turanian Subprovince closest to the Russian borders, one-third of them (52 species, or 33%) are endemic; 69 species more are endemic to the entire Asian part of the Palaearctic. Extra-Asian and extra-Siberian ranges are known in 190 flea species. In the western Palaearctic, 76 species are endemic to the European Province, and 57 species, to the Mediterranean Province; 36 species have Euro-Mediterranean distribution. The fauna of the Saharo-Arabian Subregion comprises 30 species (12 endemic), 6 species have ranges of the Mediterranean-Saharo-Arabian type. Scenarios of the origin of the Siphonaptera at the Triassic-Jurassic boundary are hypothesized. Formation of the Palaearctic flea fauna was mostly supported by the Asian-Indo-Malayan and East Asian-Western American palaeofaunal centers of taxonomic diversity. The long history of faunal exchange between the east Palaearctic and the west Nearctic is manifested by the distribution of the parasites of rodents and insectivores, fleas of the genera Stenoponia, Rhadinopsylla, Nearctopsylla, and Catallagia, belonging to several subfamilies of the Hystrichopsyllidae, as well as members of a number of other flea families. A great number of endemic species in the genera Palaeopsylla and Ctenophthalmus (Hystrichopsyllidae), both in the European and Asian parts of the Palaearctic, can be explained by the junction of the European and Asian continental platforms in the late Cretaceous and their subsequent isolation during the Paleocene. A considerable contribution to the flea fauna in the Russian territory was made by the East Asian-Nearctic center of taxonomic diversity, with a smaller role of the European palaeofauna. Immigration of species of the family Pulicidae from the Afrotropical Region is restricted to the southern territories of Russia.     

陕西汉中地区蝗虫种类及区系特点

汉中地区的蝗总科昆虫有5科33属51种,锥头蝗科Pyrgomorphidae1属2种,斑腿蝗科Catantopidae9属19种,网翅蝗科Arcypteridae5属12种,斑翅蝗科Oedipodidae12属13种,剑角蝗科Acrididae5属5种。汉中地区蝗总科的昆虫以东洋界为主,共23种,占45.1%,其次为汉中地区特有种类10种,占19.6%,古北界和两界共有种类均为9种,均占17.6%。物种数量在各县区分布不均匀。     

Patterns of formation of the flea (Siphonaptera) fauna in the Caucasus

Fleas of the Caucasus belong to 155 species of 40 genera, constituting 17% and 43% of the species and generic composition of the Palaearctic fauna, respectively. The Caucasian fauna includes 23 endemic species but no endemic genera or subgenera. In the number of species, the Caucasian fauna is similar to that of the Mediterranean Subregion and is significantly poorer than the faunas of the Euro-Siberian (by 2.2 times) and Irano-Turanian (by 1.7 times) Subregions. Based on taxonomic diversity, we can propose a hypothesis on the West and East Palaearctic sources of the Caucasian fauna. The West Palaearctic source has determined the distribution of pulicomorph fleas of the families Pulicidae and Coptopsyllidae from Africa, on the one hand, and of fleas of the genera Ctenopthalmus and Palaeopsylla from Europe, on the other hand. Fleas of the Holarctic genera, such as Ceratophyllus and Megabothris, entered the Caucasus by the north Asian route; fleas of the genera Neopsylla, Rhadinopsylla, and Hystrichopsylla migrated to the Caucasus from east and central Asia by the south Asian route, through Middle and Western Asia.     

中国蚋类区系分布和地理区划(双翅目:蚋科)

报道我国蚋类的区系分布和地理区划,根据代表性类群的界间替代分布,对我国蚋类在东洋,古北两界的分布界限初步划线,分析表明,我国蚋类区系分布具有自北向南减弱,接壤区系的相似性和具有明显的地方性等特点,在区系分析的基础上,对我国蚋类进行三级地理区划。     

Oligochaete fauna of Lake Baikal

Oligochaetes are one of the most abundant groups of invertebrates in Lake Baikal. They compose up to 70–90% of the biomass and numbers of zoobenthos and are distributed from the water edge to the maximum depth. There are 207 identified species and subspecies belonging to 44 genera and seven families, of which 160 species and 13 genera are endemic. Many are relict species. The main peculiarities of oligochaete fauna of Lake Baikal are: immiscibility of its ecologically different complexes (Palaearctic and Baikalian), its antiquity and heterogeneity, and intense radiation of the species.     

Global diversity of water mites (Acari,Hydrachnidia; Arachnida) in freshwater

The Hydrachnidia (water mites) represent the most important group of the Arachnida in fresh water. Over 6,000 species have been described worldwide, representing 57 families, 81 subfamilies and more than 400 genera. The article analyzes extant water mite diversity and biogeography. Data on distribution and species richness of water mites are substantial but still far from complete. Many parts of the world are poorly investigated, Oriental and Afrotropical regions in particular. Moreover, information among different freshwater habitats is unbalanced with springs and interstitial waters disproportionately unrepresented. Therefore, more than 10,000 species could be reasonably expected to occur in inland waters worldwide. Based on available information, the Palaearctic region represents one of the better investigated areas with the highest number of species recorded (1,642 species). More than 1,000 species have been recorded in each of the Neotropical (1,305 species) and Nearctic regions (1,025 species). Known species richness is lower in Afrotropical (787 species) and Australasian (694 species) regions, and lowest in the Oriental region (554 species). The total number of genera is not correlated with species richness and is distinctly higher in the Neotropical (164 genera); genus richness is similar in the Palaearctic, Nearctic and Australasian regions (128–131 genera) and is lower in the Afrotropical and Oriental regions with 110 and 94 genera, respectively. A mean number of about three genera per family occur in the Palaeartic, Nearctic and Oriental while an average of more than four genera characterizes the families of Australasian and Afrotropical regions and more than five genera those of the Neotropical. Australasian fauna is also characterized by the highest percentage of endemic genera (62%), followed by Neotropical (50.6%) and Afrotropical (47.2%) regions. Lower values are recorded for the Palaearctic (26.9%), Oriental (24.4%) and Nearctic (21.4%). The Palaearctic and Nearctic have the highest faunistic similarity, some minor affinities are also evident for the generic diversification of Holarctic and Oriental families. The faunas of Southern Hemisphere bioregions are more distinct and characterized by the presence of ancient Gondwanan clades with a regional diversification particularly evident in the Neotropics and Australasia. This scenario of water mite diversity and distribution reflect the basic vicariance pattern, isolation, phylogenetic diversification, recent climatic vicissitudes and episodes of dispersal between adjacent land masses together with extant ecological factors can be evoked to explain distribution patterns at a global scale. Guest editors: E. V. Balian, C. Lévêque, H. Segers & K. Martens Freshwater Animal Diversity Assessment     

Geographical distribution of the weevil subfamily Ceutorhynchinae (Coleoptera, Curculionidae)

A brief general characteristic and review of distribution of the subfamily Ceutorhynchinae over zoogeographical realms are given, with an emphasis on the distribution within the Holarctic and Palaearctic. The potential of exploiting landscapes of all natural zones of the Holarctic by a low-rank taxon is exemplified by the Holarctic Ceutorhynchus cochleariae (Gyll.) species-group comprising ca. 20 species evenly distributed between the Palaearctic and Nearctic. Although neighboring with the powerful centers of tropical biota, the Holarctic fauna of the subfamily Ceutorhynchinae is formed mostly of endemic and subendemic genera which are especially abundant in the Palaearctic. This region possesses the most diversified generic and species composition of the fauna consisting of predominantly endemic and subendemic genera (in the Mediterranean and Saharo-Gobian regions, also of the highly diversified tribe Oxyonychini) up to its southern border. The existence of characteristic transitional faunas in the zones of contact of the Holarctic fauna with the faunas of the tropical regions in East Asia and Mexico is shown. These transitional faunas include a considerable number of endemic taxa of the genus and species groups. A conspicuous feature of the Palaearctic ceutorhynchine fauna is the rather numerous complex of the upland and high-latitude species.     

辽宁浑河源自然保护区寄蝇科昆虫区系资源

调查我国辽宁东北部清原浑河源自然保护区双翅目寄蝇科昆虫资源。网捕法采集寄蝇标本2553件,形态分类和动物地理研究法鉴定寄蝇4亚科23族83属158种,分别占中国已知族、属和种类的23.54%,29.47%和12.62%;给出该森林生态系统的寄蝇优势种15种和每种成虫发生期。该地区寄蝇区系主要组成有61古北界种,86古北、东洋两界共有种,合计147种,占本地区总种数93.04%。发现4中国新纪录种:乌苏等鬃寄蝇Peribaea ussuriensis(Mesnil,1963),弯叶芙寄蝇Phryno tenuiforceps(Tachi,2013),花截尾寄蝇Nemorilla floralis(Meigen,1824),巨角锥腹寄蝇Smidtia magnicornis Mesnil,1967,给出其主要鉴别特征和外形照片;辽宁新分布纪录28种。     

庐山蛾类区系研究

庐山,地处中亚热带北沿,北纬29.35°,东经115.59°,属亚热带东部季风区域。庐山襟江带湖,雨量充沛,植物种类繁多,种子植物有1800余种,隶属158科642属。因此,庐山蛾类也较丰富。笔者于1975年5月开始,对庐山蛾类进行了调查,共鉴定出蛾类455种。现将庐山蛾类区系报告如下。 区系分析 庐山蛾类已鉴定出32科340届455种。区系情况如表。 从表可以看出:     

Midges of the subfamily Tanypodinae (Diptera,Chironomidae) of Russia: Tribes Coelotanypodini Kieffer, 1913 and Tanypodini Meigen, 1803

It is established that in the territory of Russia the subfamily Tanypodinae (Diptera, Chironomidae) includes 80 species belonging to 10 subgenera of 8 genera and 5 tribes; 4 species are recorded for the first time in the Palaearctic Region, and 16 species, in Russia. In the present study, larvae, pupae, and male adults from different areas of Russia, belonging to 9 species of 2 subgenera, 2 genera, and 2 tribes, are examined. Keys to larvae, pupae, and male adults of these species is provided.     

世界蜘蛛的分布格局及其多元相似性聚类分析

蜘蛛是一类种类繁多、数量巨大、分布广泛的捕食性生物。至2012年底,全世界共有蜘蛛43678种（包括亚种）,隶属于112科3898属。科、属、种3个分类阶元的分布域非常悬殊,90%的种分布在一个界内,90%的科是跨界分布或全球分布。按行政区域,亚洲种类较多,欧洲较少,南极洲没有蜘蛛记录;按动物地理区域,古北界和新热带界较多,新北界较少。根据地理条件、生态条件和蜘蛛的分布状况,将全球陆地分为53个基础地理单元,用申效诚等新近提出的相似性通用公式和据此创立的多元相似性聚类分析方法,分别对属、种两级的分布进行分析,得到两个聚类结构相同、聚类关系合理的支序图,而且属级的支序图层次更为分明,在相似性水平为0.32时,53个基础地理单元聚为17个小单元群,在0,22水平上,又聚为8个大单元群。这些大、小单元群的组成单元地域相邻相连,生态条件相对一致,可以作为蜘蛛地理区划的界、亚界两个层级。和华莱士主要以哺乳动物建立的世界动物地理区划相比,主要差异是：1、古北界东、西两部分差异显著,可分设两界;2、新西兰和澳洲大陆相似性较低,可单独设界;3、新几内亚和太平洋岛屿与澳洲大陆的关系远于和东洋界的关系,华莱士线在两界间的作用似乎不存在; 4、新热带界的中美地区似乎属于新北界更为合适,并由此产生了南北美洲间的紧密联系;5、新北界与古北界的相似性关系弱于与新热带界的关系,全北界的概念几近消失。前两点差异可以从地球板块构造的变动得到解释,第3、5个差异已在植物和其它生物类群得到佐证,第4个差异尚不稳定,需要更多类群的比较与分析。使用多元相似性聚类分析方法对于如此典型的点状分布的生物类群和如此海量的数据,能够得到如此精细的,既符合地理学、统计学的逻辑,又符合生物学、生态学逻辑的定量分析结果,这在国内外都是首次成功尝试,其简便性和合理性将会促使在其它类群中的应用。     

陕西延安市区蝶类多样性初探

延安市区有蝶类6科25属36种,其中凤蝶科2属3种;粉蝶科5属8种;眼蝶科3属5种;蛱蝶科7属10种;灰蝶科7属9种;弄蝶科1属1种。陕西省新记录5种。区系分析表明,该地区以广布种为优势种。     

山西历山地区的两栖爬行动物

１９９３－ １９９７ 年在山西历山地区对两栖和爬行动物进行了调查。此区共有两栖动物１１ 种, 隶２ 目５ 科８属, 其中, 古北界种３ 种、东洋界种３ 种、广布种５ 种; 爬行动物２４ 种, 隶３ 目７ 科１４ 属, 其中, 古北界种７种、东洋界种１２ 种、广布种４ 种、特有种１ 种。     

广东丹霞山国家级自然保护区 蝙蝠物种多样性调查

于2017年至2018年对广东丹霞山国家级自然保护区的翼手目动物进行调查,主要采用日栖息地与夜栖息地、捕食区网捕等调查方法,共调查到翼手目5科13属23种,其中蝙蝠科(Vespertilionidae)9属15种,菊头蝠科(Rhinolophidae)1属4种,蹄蝠科(Hipposideridae)1属2种,假吸血蝠科(Megadermatidae)1属1种,犬吻蝠科(Molossidae)1属1种。从区系组成来看,以东洋界为主(19种),其次为广布种(3种),古北界仅1种。从栖息类型上看,分为洞栖型、树栖型及建筑物栖息型,以洞栖型为主(15种),建筑物栖息型其次(12种),树栖型最少(5种),但其中9个物种的栖息地类型同时包含了上述3种栖息类型中的2种。本研究在广东丹霞山发现中国蝙蝠新分布记录1种,为卡氏伏翼(Hypsugo cadornae);而中印鼠耳蝠(Myotis indochinensis)为中国分布的再次确认。     

Pugs of the tribe Eupitheciini (Lepidoptera,Geometridae): the history of study and distribution over biogeographic regions

The World fauna of the tribe Eupitheciini is the most species-rich in the family Geometridae. This tribe includes about 1900 species (almost 3000 species-group names) from 47 genera; about one third of the genera (15) are monotypic. The generic diversity of Eupitheciini is the highest in the Australian (38 genera, 11 of them endemic) and Oriental regions (32 genera, 4 endemic) and the lowest in the Neotropical Region (possibly one genus only). The faunas of different biogeographic regions can be arranged in following order by their species richness: the Palaearctic (487 species), Oriental (397), Neotropical (346), Australian (251), Afrotropical (198), and Nearctic Regions (166 species). Eupithecia is the most species-rich genus in the family Geometridae and the entire order Lepidoptera, and one of the largest genera in the whole World fauna of insects. The greatest number of species of this genus is recorded in the Palaearctic Region (466 species), where Eupithecia accounts for about 95% of the tribe Eupitheciini. The mainland of the Oriental Region (especially the Himalayas) is also very species-rich; however the proportion of the Eupithecia representatives decreases towards Malaysia, Sundaland, and the Australian Region (about 2% of the tribe). The Eupitheciini faunas have the greatest similarity at the generic level between the Oriental and Australian Regions (the Jaccard and Sørensen coefficient values being 0.62 and 0.77, respectively). The Palaearctic fauna is more similar to the Afrotropical and Oriental faunas at the genus-group level. On the whole, the fauna of the Nearctic Region is similar to that the West Palaearctic, with the exception of the fact that representatives of the genera Gymnoscelis and Chloroclystis are absent in North America, although two endemic genera Nasusina and Prorella are present. At the genus-group level, the Nearctic fauna of Eupitheciini is more similar to the Neotropical (the Jaccard and Sørensen coefficients 0.20 and 0.33, respectively) than to the Palaearctic fauna (0.17 and 0.29). The number of synonymies is very high in the tribe Eupitheciini because of the homogeneity of this group, whose species are difficult to identify without the use of elaborate anatomical techniques. Modern revisions, catalogues, surveys, and atlases on Eupitheciini are absent for many countries and large geographic regions. Revisions of pugs of the tribe Eupitheciini for some biogeographic regions are extremely difficult because of fragmentation of entomological collections including the type specimens of many species-group taxa. A large fraction of synonyms is characteristic of parts of the World with the best known faunas: Europe (64% of synonyms) and North America (39%). On the contrary, the lowest levels of synonymy are typical of the less known faunas of the regions situated at the equatorial latitudes, namely the Neotropical (9%) and Afrotropical (8%) ones.     

小腹茧蜂亚科的雄外生殖器及族级单元系统发育的研究

对分布在东洋区和古北区的小腹茧蜂亚科 (膜翅目: 茧蜂科)21个属的67个种及外群折脉茧蜂属 (膜翅目: 茧蜂科) 2个种的雄外生殖器的5个性状进行了比较研究。在形态学研究的基础上,通过选用头部、胸部和腹部(包括雌雄外生殖器的性状) 等34个性状,运用支序分析的方法探讨了分布在东洋区和古北区的小腹茧蜂亚科21个属以及它们所属的族间的系统发育关系,并对Mason (1981) 的分类系统进行了重新评价。雄外生殖器和支序分析基本上证实并恢复了由Mason (1981) 确定的2个主要分支,即绒茧蜂族Apantelini+小腹茧蜂族Microgastrini和拱茧蜂族Fornicini+盘绒茧蜂族Cotesiini+侧沟茧蜂族MicroplitinI。绒茧蜂族Apantelini、拱茧蜂族Fornicini和侧沟茧蜂族Microplitini为单系群也被支持,但小腹茧蜂族Microgastrini和盘绒茧蜂族Cotesiini是否为单系群尚难于在树形图中体现,而且族内各属间的分支关系有变动。因此,尽管Mason的族级分类单元有一些欠缺,但仍是可信、实用的,不同意Walker等 (1990) 认为不应再使用Mason分族系统的观点。     

青海湟水流域蝶类

1995～2003年调查并整理分析青海湟水流域的蝶类,有9科52属80种,其中凤蝶科1属2种,绢蝶科1属6种,粉蝶科8属22种,斑蝶科1属1种,眼蝶科12属13种,蛱蝶科15属18种,蚬蝶科1属3种,灰蝶科9属10种,弄蝶科4属5种.以古北界为主,青藏区最为丰富.     

On the distribution of broad-winged moths (Lepidoptera, Oecophoridae) of the Palaearctic fauna

The Palaearctic fauna of broad-winged moths (Oecophoridae with the subfamilies Oecophorinae, Pleurotinae, and Deuterogoniinae) comprises 47 genera and 329 species. The number of species rapidly increases from north to south, from 19 species in the Euro-Siberian taiga Region to 149 in the Mediterranean Region, but the Scythian steppe Region (30 species) and Sethian desert Region (49 species) have relatively poor faunas because the majority of oecophorid species are trophically associated with arboreal plants. The proper desert species are very rare among Oecophoridae moths, and the majority of species in the Sethian desert Region occur in the mountains. The number of endemic species is also greater in southern regions; it amounts for more than half of the total number of species in the Hesperian and Orthrian evergreen forest regions and in the Sethian desert Region. The faunas of European (62 species) and Far Eastern (67 species) deciduous forests comprise close numbers of species but are very different, having only 5 species in common. This fact indicates their long isolation. The Oecophoridae are represented in the Palaearctic mainly by the genera with a small number of species, except for 2 genera, Pleurota (100 species) and Promalactis (85 species). The distribution patterns of these genera are opposite: the species of Pleurota are numerous in the south of West Europe and in North Africa, their number quickly decreasing from west to east and only 2 species being present in China (Wang, 2006). The species of Promalactis are numerous in Southeast Asia, their number quickly decreasing from east to west: 3 species occur in Tibet and only 1, P. splendidella (Amsel, 1935), in Israel and Turkey.     

甘肃省蚜虫类物种多样性研究

在已有标本采集记录和资料的基础上,从分类阶元、寄主植物、寄生部位、动物地理区系的角度探讨了甘肃省蚜虫类的物种多样性。结果表明甘肃省蚜虫种类丰富,已知9科90属220种;分布在华北区、蒙新区、青藏区、华中区和西南区,其中华北区、蒙新区和青藏区的种类最为丰富,因此甘肃省蚜虫区系以古北界成分为主;寄主植物种类繁多,已知有46科111属;寄生部位多样化,以叶片和茎为主要为害部位。另外就甘肃省所处地理位置的特殊性,初步讨论了在不同地理区划交界处物种生物多样性所受的影响。