In the Footprints of Our Ancestors: An Overview of the Hominid Track Record

Hominid footprints are particularly appealing and evocative of the living activity of our ancestors. The most famous and oldest (Late Pliocene, ca. 3.7 Ma) hominid footprints, from Laetoli in East Africa, have been attributed, with some uncertainly, to genus Homo or Australopithecus. The African track record also yields Early Pleistocene (～1.5 Ma) tracks attributable to Homo erectus. The only well-documented Middle Pleistocene tracks (age ～325,000-385,000 yrs) are reported from Italy and presumably represent a pre-Homo sapiens species.

The oldest Late Pleistocene tracks (～117,000 yrs), from southern Africa, may represent modern humans. However, the majority of Late Pleistocene sites are European, associated with caves in Romania, Greece, France and elsewhere, where hominid track preservation is often of high quality. Dates range from ～10,000 to ～62,000 BP Cavesite mammal tracks are almost exclusively those of carnivores, thus representing a distinctive underground ecology. Late Pleistocene open air sites are reported from widely scattered locations in Africa, Turkey, Tibet, Korea, Australia and even in the New World (Chile, Argentina and Mexico).

Early to Middle Holocene sites (> ～4,000 yrs BP) mainly occupy riparian, lacustrine, estuarine and littoral settings where the ichnofaunas are dominated by ungulates and shorebirds. Among these sites from England, Nicaragua, Argentina and Mexico and the United States, a few have been described in some detail. Younger Holocene sites are frequently associated with specified cultural periods (e.g., Neolithic, Bronze Age) or specific indigenous cultures, where supplemental archeological evidence may be directly associated with the footprint evidence.

At most surficial and some subterranean hominid tracksites, mammal and/or bird tracks are quite common and of use in creating a paleoecological picture of local faunas. The global distribution of human and hominid tracks is consistent with body fossil evidence and the record of archeological, cultural artifacts. However, in a few cases tracks suggest colonization of certain regions (Tibetan Plateau and the New World) earlier than previously thought. Tracks also give clues to behavior, age and health status of the trackmakers.     

Human temporal bone from the lower Paleolithic site of Castel di Guido, near Rome, Italy

The Castel di Guido site, with an estimated age of approximately 300,000 years, has yielded abundant animal remains, Acheulian stone and bone bifaces, and small tools. On the surface of the original deposit, turned over by agricultural activities, fragments of human remains were discovered between 1980 and 1986, including a temporal bone described here.     

Analysis and Preservation of Pleistocene Human and Animal Footprints: An Example from Toluquilla,Valsequillo Basin (Central Mexico)

Human and animal footprints found in the Valsequillo Basin were formed on the upper bedding planes surfaces of a volcanic ash (Xalnene Ash) deposited by a subaqueous volcano along the shores of a Pleistocene lake. The footprints were made on lake shorelines and the exposed lake floor during low stands associated, either with water displacement during the volcanic eruption, or due to climatically-driven fluctuations in the water balance. The Xalnene Ash has been dated to at least 40 K BP and consequently the human footprints provide evidence for much earlier colonization of the Americas than is often accepted. The methodology used to record, analyze and conserve these footprints used three-dimensional laser scanning, with sub-millimeter precision. This data were used to reproduce polymer models of individual footprints using the application of rapid-prototyping technology. This technology has wide significance for the study of ichnofacies in general. The characteristics of the human footprints and some problems associated with the volcanic ash as a molding medium for the clarity of these characteristics are given. These human footprints and their dating resolve the controversy related to the age of the archaeological and associated megafaunal remains that were reported in the Valsequillo Gravels in the 1960s and 1970s.     

A new dasycladalean alga from the Middle Triassic carbonates of Northern Calabria (Southern Italy)

Summary A new dasyclad alga—Acicularia boniae n.sp.—is discribed from Middle Triassic (?Ladinian) of the Piano del Minatore Formation, outcropping in the Cozzo del Pellegrino area (Calabria, southern Italy). The systematic position of this species, with respect to the genusAcicularia and morphogeneraAciculella andTerquemella, is discussed.

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Riassunto Viene descritta una nuova specie di dasicladale—Acicularia boniae—proveniente dal Triassico Medio della formazione del Piano del Minatore affiorante nel massiccio del Cozzo del Pellegrino (Calabria, Italia meridionale). Viene discussa la posizione sistematica rispetto al genereAcicularia ed ai morfogeneriTerquemella edAciculella.

     

A new large barn owl (Aves, Strigiformes, Tytonidae) from the Middle Pleistocene of Sicily, Italy, and its taphonomical significance

A new species of Tytonidae, Tyto mourerchauvireae, is described from the Sicilian cave deposits of Spinagallo, Luparello and Marasà, which have yielded a common vertebrate fossil assemblages referred to the early Middle Pleistocene. T. mourerchauvireae nov. sp. shows a pronounced increase in body size compared to other congeneric taxa. It is larger than the extant Tyto alba and the extinct Tyto balearica and Tyto sanctialbani and comparable in size among the Mediterranean taxa only with the extinct Tyto robusta, which differs in some morphological characteristics of the long bones. The insular adaptations of T. mourerchauvireae nov. sp. and the differences from the other forms of the genus Tyto which spread through the Mediterranean area during Neogene and Pleistocene are discussed. The fossil assemblages of the early Middle Pleistocene of Sicily are dominated by extinct giant Gliridae of the genus Leithia and Maltamys; these taxa are regarded as the primary prey items of T. mourerchauvireae nov. sp. The Gliridae remains were analysed microscopically to detect possible modifications of bones and teeth caused by ingestion and digestion processes. This type of analysis, together with the qualitative study of the fossil assemblages, allows to show the taphonomical importance of T. mourerchauvireae nov. sp. as agent of accumulation in such Sicilian fossil assemblages. Incidentally the taxonomic validity of T. robusta is discussed and confirmed.     

Dominant diatoms in the interglacial lake sediments of the Middle Pleistocene in Central and Eastern Poland

Diatom studies carried out in Central and Eastern Poland have shown a diversified composition and considerable changes of relative frequency of dominant diatoms in profiles of lake sediments of the Mazovian and Ferdynandovian interglacials. Cyclotella spp. are dominant in a large part of the sediment profiles at Adamówka, awki-7 and Wola Grzymalina-59 and, at Krpiec and Podlodów, Stephanodiscus is also common while at Biaa Podlaska and Falcice, Aulacoseira is most abundant. Cyclotella comta var. lichvinensis and C. vorticosa are the typical diatoms in the Mazovian Interglacial. In the sediments representing the Ferdynandovian Interglacial, the characteristic diatoms are Cyclotella cf. reczickiae and the Stephanodiscus rotula/niagarae complex.     

Occurrence of Theropithecus sp. in the Late Villafranchian of Southern Italy and implication for Early Pleistocene "out of Africa" dispersals

Three cervical vertebrae of a large primate have been identified in the revision of the faunal assemblage collected during the late '70s from fissure fillings in the area known as Pirro Nord in southern Italy. By size the three vertebrae are comparable to the large fossil species Theropithecus oswaldi, however we prefer here to attribute the remains to Theropithecus sp. The Pirro Nord Theropithecus represents the second finding outside of Africa of this genus in association with Megantereon whitei, after the site of 'Ubeidiya. The biochronology of Pirro Nord faunal assemblage demonstrates that the arrival into Europe of Theropithecus occurred earlier than thought: in the range of approx 1.6-1.3 Ma. The identification of an European "African assemblage" (Theropithecus associated with Megantereon whitei) has consequences for interpretation of an "out of Africa" (genus Homo) event around the Plio-Pleistocene transition.     

湖北郧西白龙洞中更新世古人类遗址的哺乳动物化石

白龙洞是位于鄂西北部郧西县的一处出土物丰富的古人类洞穴遗址,先后在此出土了8枚古人类牙齿化石、若干石制品和骨制品及大量哺乳动物化石。但有关白龙洞遗址的形成时间却一直存在争议,主要原因之一是动物群没有得到充分研究。基于郧西县文管所和中科院古脊椎所收藏标本,本文首次对白龙洞历年出土的哺乳动物化石进行了综合研究;研究结果表明,白龙洞动物群由32种（含未定种）哺乳动物组成,优势属种是谷氏大额牛、岩羊、野猪及各种鹿类(水鹿、斑鹿、麂、毛冠鹿);此外,还有豪猪、竹鼠、变异狼、貉、豺、猪獾、黑熊、大熊猫、中国硕鬣狗、锯齿剑齿虎、虎、豹、金猫、云豹、剑齿象、苏门犀、梅氏犀、巨貘、鬣羚及水牛等;该动物群缺乏猴类动物,并且豪猪化石也很少,这与其所处的时空位置很不相符;但总体面貌属于大熊猫-剑齿象动物群。白龙洞的大熊猫、中国硕鬣狗、猪獾、虎及巨貘的牙齿测量数值均达到或接近我国各自记录的最大值。白龙洞动物群组成与盐井沟动物群及南京猿人动物群最为接近;从动物群组合及有关属种的演化阶段来综合判断,白龙洞动物群的地质时代应当是中更新世中期。     

Late Quaternary Palaeoichnological Sites from the Southern Atlantic Coast of Buenos Aires Province,Argentina: Mammal,Bird and Hominid Evidence

Palaeontological and palaeoichnological studies were made along the southeastern coast of Buenos Aires province, Argentina. Continental Late Pleistocene and continental, transitional and marine Holocene stratigraphic units were identified. The former are represented by fossil mammal bones and ichnites dated from 16,000 to 12,000 y. B.P. Continental Early Holocene beds dated circa 8,100 years B.P. contain fish, rodent and bird remains. Middle Holocene sediments 6,930 to 6,570 years B.P. are recorded by deposits bioturbated by estuarine crabs. During the Late Holocene, between 5,000 and 4,800 B.P., a marine transgression is recorded by a very rich mollusc fauna. The palaeoenvironmental variety recorded along a few tens of kilometers of the coast is a distinctive feature of this part of the Argentine coastal region. At Monte Hermoso beach, in the eastern part of the studied zone, archaeological materials were dated 7,300 years B.P., and human footprints dated 7,125 years B.P. were also recorded. In January 2005 some isolated human footprints were found at Pehuen-Co palaeoichnological site, which are older than those found at Monte Hermoso beach and of the same age as the oldest human archaeological findings in Patagonia, that is about 12,000 yeas B.P.     

Pattern of dental development in Hominid XVIII from the Middle Pleistocene Atapuerca-Sima de los Huesos site (Spain)

. We describe the pattern of dental development of Hominid XVIII from the Middle Pleistocene Sima de los Huesos (SH) site of the Sierra de Atapuerca (Burgos, Spain). As expected, this pattern is similar to that of modern humans. A delay of development of the lower and upper canines was observed. In contrast, the relative advanced development of the lower second molars and, especially, the upper and lower third molars is noteworthy. This latter feature seems to be common in Pleistocene hominids, and suggests that the pattern of dental development evolved in the genus Homo during the Pleistocene. In European Middle Pleistocene hominids, this pattern probably was facilitated by the extra space available in the mandible and maxilla for developing teeth.     

中更新世晚期中国古人类化石的形态多样性及其演化意义

近年对许家窑、许昌、华龙洞、澎湖、夏河、哈尔滨等人类化石开展的系统研究,引发了学界对中更新世晚期人类演化及分类的不同认识。基于对相关中国人类化石形态特征的分析,作者提出这一时期中国人类化石形态特征表现为四种类型：1）以中更新世晚期人类共有特征为主;2）以原始特征为主;3）以现代特征为主;4）独特形态组合。多数化石形态特征表现为前三种类型,而许昌和许家窑这种以硕大的头骨和巨大颅容量构成的独特形态组合在其他同时期化石还没有发现。化石形态的多样性提示,不同类型的中更新世晚期中国古人类对现代人的形成贡献不同。作者认为在该时期的人类化石形态多样性规律还未阐明的情况下,将具有混合或镶嵌特征的相关人类化石归入分类地位不确定的人群较为合适。     

The cholera: The epidemics and their social-demographic features in Southern Italy

Cholera, as it was regarding each infective disease of which nobody knew its aetiologic agent or, in any way, if its agent was present really, was for the Calabrian people an unknown enemy that nobody knew anything about either its behaviour or the modality of its penetration into the human body. During many years characterized by a medical methodological obscurantism, Calabrian people lived, during the different choleric epidemics, on a daily life based upon the ignorance, and the pseudo-treatments, and pseudo-physicians. During the cholera epidemics, not only the people groped in the dark, but also the governmental authorities were confused because were subdivided into those who supported the theory of the epidemics spread and those who supported the theory of the contagion spread. It must be emphasized that from the XX century, when the pathologies due to natural agents were reduced and were substituted with those due to humans, cholera influenced very much the demography and the social-cultural features of the Southern Italy.     

The Lower-Middle Pleistocene succession of the Coastal Tuscany (Central Italy): new stratigraphic and palaeoecological data based on the ostracod fauna

The Lower-early Middle Pleistocene succession of the Coastal Tuscany is known to comprise three marine cycles: (I) a Santernian-Emilian cycle; (II) a Sicilian (“small Gephyrocapsa” Zone) cycle; (III) a third cycle, referred through stratigraphic and palaeoethnological arguments to the late Sicilian-early Middle Pleistocene, including the fluvial-transitional San Marco fm and the shoreface to backshore sandy-arenitic deposits of the correlatable Bibbona and Casa Saracino formations, outcropping in the Bibbona (Lower Cecina Valley) and Rosignano areas respectively. Conversely to the older cycles the third one has been poorly studied and its chronology and depositional history have remained somewhat uncertain. With the aim to fill this gap of knowledge the sedimentary record exposed in the Rosignano and Bibbona areas was the object of new on field investigations and microfaunal content (chiefly ostracods) analyses. Furthermore, this has represented a good opportunity to enhance our knowledge of the Pleistocene Mediterranean ostracods. The main results achieved are in synthesis the followings. (1) Stratigraphic and palaeoenvironmental significance of ostracods from the first cycle is consistent with literature data. Unexpectedly the recovered assemblages comprise both warm-temperate species (e.g. Cytherelloidea beckmanni Barbeito-Gonzales, Propontocypris solida Ruggieri, Verrucocythereis bulbuspinata (Uliczny), which suggest a relatively warm climate phase, and an yet undescribed species of Ruggieria, a genus previously thought to be represented in the Italian Lower Pleistocene only by Ruggieria nuda Moyes. (2) In agreement with previous studies, sediments of the San Marco fm in the Rosignano area are referable to a floodplain-coastal lagoonal setting. Divergently from literature data, in the Bibbona area the unit exhibits vuggy carbonate glaebules and rizhoconcrections and yields very rare fresh-brackish water ostracods and marine microfaunas regarded as reworked. Despite interpretation of these sediments still poses many problems, we speculate that they represent marine deposits reworked in a poorly drained continental-transitional environment, which experienced pedogenic alteration. Furthermore, the common occurrence of the ostracode Aurila puncticruciata Ruggieri seems to support the supposition that reworked deposits included marine Sicilian sediments completely eroded and presumably correlatable to the Fabbriche fm. (3) Lithological-sedimentological features and absence of autochthonous macro-microfossils indicate that the Casa Saracino fm and most of the Bibbona unit accumulated in a backshore environment dominated by aeolian deposition. Only locally the latter unit includes shallow marine deposits with fairly rich ostracod faunas, which confidently indicate an age not younger than the Sicilian sensu Ruggieri and Sprovieri [Riv. Mineraria Siciliana 151/153 (1975) 1]. Thus, it seems possible that the Coastal Tuscany succession includes two marine cycles, which developed within the Sicilian.     

Carbonate facies dominated by syndepositional cements: a key component of Middle Triassic platforms. The Marmolada case history (Dolomites, Italy)

Summary This article deals with the discussion of the role of the syndepositional cementation for the growth of the Middle Triassic pre-volcanic carbonate platforms of the Dolomites (Southern Alps, Northern Italy). The study is concentrated on the Marmolada Buildup, which escaped the facies destroying dolomitization which affected many surrounding platforms. The investigations took place within an almost isochronous uppermost Anisian palcogeographic transect, ranging from the platform-top to the margin and the upper slope. Methods used include geological mapping, sedimentological and paleontological studies, evaluation of the microfacies, as well as SEM and EDS epifluorescence analyses. The well bedded platform-top succession consists of intra-bioclast calcarenites and calcirudites, interbedded with subordinate boundstones, and organized in shallowing upward, meter scale depositional cycles, sometimes capped by subaerial surfaces. The platform margin belt is rich in boundstones and lacks a primary framework formed by organisms; metazoan skeletons form less then 5% of the rock volume. The outer margin and the uppermost slope are characterized by decimeter-scale boundstone blocks, coated and linked to each other by huge amounts of radiaxial fibrous calcite cements, arranged in concentric crusts. These cements (“evinospongiac”) represent the main component of the margin and upper slope facies. Epifluorescence analyses suggest the existence of abundant organic residual matter associated not only with the bioclasts and peloids, but also with the syndepositional cements. Organic matter likely played a significant role in carbonate cementation and was a key factor for the early lithification of the platform as well as for the sediment production. Minor element microanalyses reveal an uniform Mg content in different calcite types (2–4 Mole % MgCO₃), independently from the primary nature of the components. Late diagenetic sparry calcites exhibit similar Mg values but no iron. These data point to a homogenization of minor element distribution, probably associated with a slow but long-lasting semi-closed fluid circulation, possibly related with the Neogene uplifting of the Dolomite Mountains.     

更新世中期中国古人类演化区域连续性与多样性的化石证据

以往,在东亚大陆发现的更新世中期人类化石被分别归入直立人和古老型智人。这种分类的主要依据是化石形态特征以及年代。魏敦瑞对周口店第一地点人类化石研究描述的一些头骨、下颌骨和牙齿特征通常被作为判定直立人的标准。根据这些化石的年代分布,一般将30万年前的中更新世晚期作为划分直立人与古老型智人的大致年代界限。近20年来,在非洲、欧洲和东亚新发现了一些更新世中期人类化石,目前古人类学界对中国更新世中期人类化石特征及演化有了与以往不同的认识。最近对大荔、许家窑、盘县大洞、许昌、华龙洞等人类化石的研究显示,近30万年以来东亚大陆人类演化呈现复杂的多样性,将这一时期人类全部归入古老型智人难以准确反映更新世中期中国古人类演化模式及规律。本文结合近年中国更新世中期人类演化研究进展,选择部分具有演化及分类价值的形态特征,分析这些特征在更新世中期中国古人类化石的表现特点。在此基础上,对更新世中期中国古人类演化模式做了尝试性探讨。本研究发现,周口店、和县、沂源、南京等中更新世早期人类化石呈现有较多的区域性特征,形态特征表现相对稳定;而大荔、金牛山、许家窑、许昌、华龙洞、马坝、盘县大洞等中更新世晚期人类在化石形态特征表现复杂多样,变异范围大。此外,在这一时期人类化石上发现较多与生存活动、健康、环境适应有关的证据。根据这些发现,作者认为中国中更新世早期组人类演化以形态连续性为主;进入中更新世晚期,中国古人类演化区域性特征减弱,演化模式以多样性为主。一系列新的化石发现和研究证据提示中更新世晚期东亚大陆可能生存有不同的古人类成员。根据目前掌握的化石形态和年代证据,大约30万年前是中国更新世中期演化变化关键时间节点。     

The Casal de' Pazzi archaic parietal: comparative analysis of new fossil evidence from the late Middle Pleistocene of Rome

A fossilized fragment of human parietal bone has been recently recovered from the lowest layer of the Casal de' Pazzi fluvial deposit (stratigraphically dated at about 200–250 ky BP). The fossil presents characters-i.e., thickness, degree and development of curvature, type of endocranial vascularization-which distinguish it from the corresponding cranial regions of both Homo erectus and anatomically modern Homo sapiens. While a morphological orientation towards Neanderthal characters can be considered, the affinities of the Casal de' Pazzi parietal are primarily with other late Middle Pleistocene specimens. The authors conclude that the Casal de' Pazzi human find can be assigned to the “archaic Homo sapiens” group falling within the European pre-Neanderthal range. Its particular morphology constitutes new evidence of human evolution from the geographical area of Rome.     

Pleistocene camelids from the Syrian Desert: The diversity in El Kowm

The family Camelidae has been present in Eurasia since the latest Miocene, and several species are recognized, but their evolution is poorly known. The region of El Kowm, central Syria, includes several sites spanning the Early to Late Pleistocene and provides the only abundant fossil record of camelids in the Middle East. Our preliminary results show that several species are present over the sequence, revealing some surprising evolutionary trends.     

The late Oligocene molluscan fauna from Otranto (Apulia, Southern Italy): an example of alternating freshwater, lagoonal and emerged environments

Abstract:  Eighteen gastropod and five bivalve species are taxonomically described from an Upper Oligocene succession cropping out near Otranto (southern Salento, Apulia, Italy). Among these, Tectarius ( Echininus ) japigiae , Hydrobia dubuissoni hydruntina , Pseudamnicola messapica , Pseudamnicola palmariggii and Stenothyrella salentina are new taxa. Nonmarine and lagoonal environments characterize almost the whole succession, as testified by species-poor assemblages rich in specimens of freshwater and oligo- to mesohaline prosobranchs, such as Neritinidae, Hydrobiidae, Stenothyridae, Thiaridae, Potamididae, Batillariidae, pulmonates, including Planorbidae and bivalves comprising Dreissenidae and Cyrenidae. The genera Theodoxus , Hydrobia , Melanoides , Potamides , Terebralia , Batillaria , Granulolabium , Mytilopsis and Polymesoda alternately dominate. Littoral marine elements, such as Tectarius , Turritella , Barbatia , Anadara and Chama , are scattered present in the succession, but some prevail towards the top. The fauna is palaeobiogeographically significant having strong affinities with those of the Oligo–Miocene basins of Aquitaine, Mainz, Bavaria, North Alpine Foreland, Vienna, Greece and Turkey. The analysed Otranto succession is assigned to the Chattian Galatone Formation of southernmost Apulia (Salento) based on lithological evidence and supported by the palaeontological data.