Probable Relationships between the Lower Jurassic Crocodilomorph Trackways Batrachopus and Selenichnus: Evidence and Implications Based on New Finds from the St. George Area Southwestern Utah

Some small and medium-sized crocodylomorph footprints are described from the Late Jurassic (Kimmeridgian) coastal and deltaic units of the northern Spain (Asturias). There are at least four footprint morphotypes. Three of them, with well preserved trackways, are included in the ichnogenus Crocodylopodus (Crocodylopodus isp. and Crocodylopodus meijidei); the fourth one, documented by some isolated large footprints, is referable to the ichnogenus Hatcherichnus. This ichnoassociation confirms the presence of small crocodilians in palaeoenvironments apparently dominated by dinosaurs. The presence of Hatcherichnus seems to confirm the affinity between the Iberian and North American ichnofaunas.     

Analysis and Preservation of Pleistocene Human and Animal Footprints: An Example from Toluquilla,Valsequillo Basin (Central Mexico)

Human and animal footprints found in the Valsequillo Basin were formed on the upper bedding planes surfaces of a volcanic ash (Xalnene Ash) deposited by a subaqueous volcano along the shores of a Pleistocene lake. The footprints were made on lake shorelines and the exposed lake floor during low stands associated, either with water displacement during the volcanic eruption, or due to climatically-driven fluctuations in the water balance. The Xalnene Ash has been dated to at least 40 K BP and consequently the human footprints provide evidence for much earlier colonization of the Americas than is often accepted. The methodology used to record, analyze and conserve these footprints used three-dimensional laser scanning, with sub-millimeter precision. This data were used to reproduce polymer models of individual footprints using the application of rapid-prototyping technology. This technology has wide significance for the study of ichnofacies in general. The characteristics of the human footprints and some problems associated with the volcanic ash as a molding medium for the clarity of these characteristics are given. These human footprints and their dating resolve the controversy related to the age of the archaeological and associated megafaunal remains that were reported in the Valsequillo Gravels in the 1960s and 1970s.     

Human Tracks from Quaternary Tufa Deposits,Cuatro Cienegas,Coahuila, Mexico

In 1961, two human footprints were excavated from the Cuatro Cienegas region of Coahuila, Mexico, but for decades were not studied scientifically or deposited in any museum. Consequently, knowledge of the locality where they were found was lost. Once the two tracks were relocated they found their way to the Museo del Desierto, Saltillo (Coahuila) where they were placed on display with a tentative label suggesting an early Holocene age (10,000 B.C.). This inference was based only on the known antiquity of humans in the region determined from previous archaeological work and two dated sediment cores close to the footprint site. It was not until 2006 that the presumed footprint discovery site was rediscovered, revealing more than 20 additional well-preserved prints, in at least four trackways representing several different individuals with foot lengths ranging from about 23–27 cm. The tracks include one well-preserved trackway providing step and stride measurements. A poorly preserved cervid trackway and a possible duck footprint have also been identified. The footprints are preserved in hard tufa. Geochemical evidence suggests the tracks were made during periods of increased aridity.     

Sauropod and Small Theropod Tracks from the Lower Jurassic Ziliujing Formation of Zigong City,Sichuan, China,with an Overview of Triassic–Jurassic Dinosaur Fossils and Footprints of the Sichuan Basin

A dinosaur footprint assemblage from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, comprises about 300 tracks of small tridactyl theropods and large sauropods preserved as concave epireliefs (natural molds). The theropod footprints show similarities with both the ichnogenera Grallator and Jialingpus. Three different morphotypes are present, probably related to different substrate conditions and extramorphological variation. A peculiar preservational feature in a morphotype that reflects a gracile trackmaker with extremely slender digits, is the presence of a convex epirelief that occurs at the bottom of the concave digit impressions. It is possibly the result of sediment compaction underweight load when the pes penetrated the substrate, being a resistant residue during exhumation and weathering. The sauropod tracks belong to a trackway with eight imprints consisting of poorly preserved pes and manus tracks and a better preserved set, probably all undertracks. The narrow-gauge trackway pattern resembles the ichnogenus Parabrontopodus well known from the Jurassic but other features such as the minor heteropody are different. The assemblage enriches the dinosaur record from the famous Zigong locality and the evidence from the Lower Jurassic in this area that was restricted to a few skeletal remains and footprints. Furthermore it proves the presence of small theropods, whereas skeletons of the group, well- known from the Middle-Upper Jurassic of Zigong, are of medium to large size only. Remarkable is the dominance of saurischians in these assemblages, which is characteristic of Jurassic dinosaur communities whereas the Cretaceous record shows an increase of ornithopod groups. An overview of the dinosaur trace and body fossil record of the Sichuan Basin supports this view. The paleoenvironment can be designated as a low-latitude tropical freshwater lake as it is indicated by bivalve shells.     

Vertebrate Tracksites in the Middle Jurassic-Upper Cretaceous of South Tunisia

Four vertebrate tracksites from the Middle Jurassic and Upper Cretaceous in the Tataouine basin of southern Tunisia are described. Approximately 130 tridactyl footprints distributed over an area of 200 square meters, preserved on Callovian beds exposed at the Beni Ghedir site, represent the oldest evidence of a dinosaur fauna in Tunisia. In addition, three tracksites—Chenini, Ksar Ayaat, and Jebel Boulouha—have been discovered in the Cretaceous beds of the upper Continental Intercalaire, previously considered as a strictly marine depositional sequence. In addition to dinosaur tracks, the Chenini tracksite (late Albian) includes poorly preserved crocodilian tracks, and footprints assigned to a pleurodiran turtle have been recovered at the Ksar Ayaat locality (early Cenomanian). The Jebel Boulouha tracksite is dominated by well-preserved tridactyl tracks referred to small-sized theropods. Depositional settings of each tracksite have been defined on stratigraphic and sedimentologic data, and tracks were ascribed to different ichnocoenoses in relation to their paleoenvironments. This new and differentiated track record gives important information on how the fossil vertebrate fauna changed in southern Tunisia during mid-Jurassic to mid-Cretaceous times. These data provide a unique and useful census of tetrapod associations along the southern margin of the peri-Mediterranean area.     

陕西子洲中侏罗世早期恐龙足迹初步研究

陕西子洲县中侏罗统延安组砂岩中,同一地点共厚约1.7 m的5个层面发现4种食肉类恐龙足迹,自上而下分别为:1)第5层面大型三趾型足迹,实雷龙足迹科王氏子洲足迹(新遗迹属、新遗迹种)Zizhoupus wangi ichnogen.et ichnosp,nov.;2)第4和第3层面中型三趾型足迹,实雷龙足迹科龙尾峁张北足迹(新遗迹种)Changpeipus longweimaoensis inchnosp.nov.;3)第2层面小型三趾或四趾型足迹,虚骨龙类小理河陕西足迹(新遗迹种)Shensipus xiaoliheensis inchnosp.nov.;4)第1层面小型三趾型足迹,虚骨龙类的铜川陕西足迹Shensipus tungchuanensis.     

Problems in discrimination of tridactyl dinosaur footprints,exemplified by the Hettangian trackways,the causses,France

Numerous tridactyl dinosaur footprints have been discovered in the Lower Jurassic (Hettangian) of the Causses, France. They exhibit the usual Grallator‐like morphology in which differences seem to be more a result of normal intraspecific variability than of inherent differences between skeletons. However, such subjective conclusions beg an analytical confirmation, which was the principal objective of this study.

For more reliable shape determinations, typical specimens from well‐preserved footprint ichnofaunas of the Connecticut Valley (USA) were used. Statistical methods are necessary to verify the homogeneity of such footprint populations. Determinations of the variability, confidence interval for the mean, and ratios between length characters are particularly important in reducing the influence of the size of each print. By this method, the ichnospecies Anomoepus intermedius, Eubrontes giganteus, and Grallator sillimani are statistically distinct; the equally distinct new ichnospecies from Saint‐Léons (Aveyron) can be designated as Grallator lescurei and the trackways of Saint‐Laurent as G. minusculus.

A problem remains in classifying the trackmaking dinosaurs. Their prior assignment to the theropods apparently should be revised because of the abundance of grallatorid footprints in the Lower Jurassic, which seems to contradict conclusions drawn from paleoecological data.     

Constructing and testing hypotheses of dinosaur foot motions from fossil tracks using digitization and simulation

Whilst bones present a static view of extinct animals, fossil footprints are a direct record of the activity and motion of the track maker. Deep footprints are a particularly good record of foot motion. Such footprints rarely look like the feet that made them; the sediment being heavily disturbed by the foot motion. Because of this, such tracks are often overlooked or dismissed in preference for more foot-like impressions. However, the deeper the foot penetrates the substrate, the more motion is captured in the sediment volume. We have used deep, penetrative, Jurassic dinosaur tracks which have been naturally split into layers, to reconstruct foot motions of animals living over 200 million years ago. We consider these reconstructions to be hypotheses of motion. To test these hypotheses, we use the Discrete Element Method, in which individual particles of substrate are simulated in response to a penetrating foot model. Simulations that produce virtual tracks morphologically similar to the fossils lend support to the motion being plausible, while simulations that result in very different final tracks serve to reject the hypothesis of motion and help generate a new hypothesis.     

Holocene Human Footprints in North America

Reports of human footprints from North American archaeological sites are summarized and described. Although some human footprints have been claimed to be of Pleistocene and earlier ages, the earliest authenticated prints in America north of Mexico, as distinct from Central or South America, date to the Holocene. The earliest of these prints is at least 5,070 years old, and the youngest are late prehistoric period, only about 400 years old. Footprint sites are reported in the U.S. Southeast, Southwest, and California. Activities indicated by ancient North American human footprints include the mundane, such as daily tasks in a riparian zone and domestic behaviors inside a habitation. Footprints were also left by prehistoric peoples undertaking more esoteric activities deep in cave interiors, including exploration, mineral extraction, and ritual.     

关于以色列国的基巴拉和哈约尼姆两洞穴的微矿物的研究：对于考古纪录中灰烬沉积的认识

关于以色列国的基巴拉和哈约尼姆两洞穴的微矿物的研究：对于考古纪录中灰烬沉积的认识斯·韦纳斯·希格尔奥·巴·约瑟夫关键词火,灰烬,矿物,红外光谱学,成岩作用火的考古纪录对于认识早期人类的技术能力是十分有趣的。通过研究烧骨、炭屑、石器、以及高温对于烧过的...     

Avian ichnia and other vertebrate trace fossils from the Neogene Red Beds of Tarom valley in north-western Iran

The Neogene Red Beds of the Tarom valley (north-western Iran) include conglomerate, sandstone, marl and gypsum. Avian and mammal footprints were discovered in one of the sandstone layers at the base of a third Miocene stratigraphical unit in the Gilankesheh area located in the east Tarom valley. The avian ichnia include Aviadactyla vialovi, Avipeda filiportatis, Charadriipeda disjuncta, Charadriipeda isp. A and B and cf. Ornithotarnocia lambrechti. Bird feeding traces are preserved as bilobate, loop-shaped, sinusoidal and ring-like traces. We have also identified a reticulate texture of sole scale imprints in some of the avian ichnia. Two mammal footprints of camelid-like artiodactyls are also present with the avian ichno-assemblage.     

Dinosaur Tracks from the Kilmaluag Formation (Bathonian,Middle Jurassic) of Score Bay,Isle of Skye,Scotland, UK

Tracks of a juvenile theropod dinosaur with footprint lengths of between 2 and 9 cm as well as adults of the same ichnospecies with footprints of about 15–25 cm in length were found in the Bathonian (Middle Jurassic) Kilmaluag Formation of Score Bay, northwestern Trotternish Peninsula, Isle of Skye, Scotland, UK. Two footprint sizes occur together on the same bedding plane in the central portion of Score Bay, both in situ and on loose blocks. Another horizon containing footprints above this was also identified. The footprints from the lowest horizon were produced in a desiccated silty mud that was covered with sand. A close association of both adults and juveniles with similar travel direction indicated by the footprints may suggest post-hatching care in theropod dinosaurs. Other footprints, produced on a rippled sandy substrate, have been found on the slightly higher bedding plane at this locality. Loose blocks found 130 m to the northeast in the central part of Score Bay have not been correlated with any in situ sediments, but were preserved in a similar manner to those from the higher bedding plane. These tracks represent the youngest dinosaur remains yet found in Scotland.     

Tetradactyl footprints of an unknown affinity theropod dinosaur from the Upper Jurassic of Morocco

Background

New tetradactyl theropod footprints from Upper Jurassic (Oxfordian-Kimmeridgian) have been found in the Iouaridène syncline (Morocco). The tracksites are at several layers in the intermediate lacustrine unit of Iouaridène Formation. The footprints were named informally in previous works “Eutynichnium atlasipodus”. We consider as nomen nudum.

Methodology/Principal Findings

Boutakioutichnium atlasicus ichnogen. et ichnosp. nov. is mainly characterized by the hallux impression. It is long, strong, directed medially or forward, with two digital pads and with the proximal part of the first pad in lateral position. More than 100 footprints in 15 trackways have been studied with these features. The footprints are large, 38–48 cm in length, and 26–31 cm in width.

Conclusions/Significance

Boutakioutichnium mainly differs from other ichnotaxa with hallux impression in lacking metatarsal marks and in not being a very deep footprint. The distinct morphology of the hallux of the Boutakioutichnium trackmaker –i.e. size and hallux position- are unique in the dinosaur autopodial record to date.     

Late Pliocene grassland from Olduvai Gorge,Tanzania

The Olduvai fossil plants documented by us in this paper are the first direct evidence for open grassland in the late Neogene of Africa based on macroplant remains. Silicified remains of herbaceous ground cover are exceptionally well preserved in situ within Late Pliocene sediments below the initial pyroclastic ash surge unit of Tuff IF in the uppermost part of Bed I, Olduvai Gorge, northern Tanzania. Published radiometric and palaeomagnetic dates place this grass layer between 1.839 + 0.005 Ma and 1.785 + 0.01 Ma. Exposed at localities on the south side of the Gorge this herbaceous ground cover grew on a floodplain developed on a dried out lake bed, following pronounced lake retreat of saline–alkaline palaeo-Lake Olduvai during a developing dry climatic phase. Sheathed basal culms, rhizomes and roots are interpreted as those of one or more small mat-forming grasses or less likely, sedges. Small dicotyledonous herbs were probably also present. The proximity of adjacent plants indicates a relatively dense ground cover. Roots extended at least 8 cm below the ground surface. Aerial parts of the plants were absent or were not preserved when the weathered basal culms were covered by a thin layer of brown waxy clay, followed by fallout of pyroclastic ash. Both units were mostly eroded away prior to emplacement of a wet, cool pyroclastic surge which then buried and preserved in situ remnants of the herbaceous ground cover. Preservation of the semi-woody rhizomes implies well-drained soils, otherwise the plant material would have quickly rotted. Collections from discontinuous exposures indicate the grassland covered an area of at least a few hectares. This open grassland would have provided grazing for the Late Pliocene fauna.     

A REVIEW OF TRIASSIC TETRAPOD TRACK ASSEMBLAGES FROM ARGENTINA

Abstract:  This paper contains the first comprehensive ichnotaxonomic review of the Triassic tetrapod track record in Argentina, including previous accounts and new material recently discovered, and an analysis of its composition and stratigraphic distribution. Triassic footprints have been recorded from three basins: the Ischigualasto-Villa Unión and Cuyo basins in north-west Argentina, and the Los Menucos depocentre in northern Patagonia. Most are in successions of Middle Triassic age; a lower number are from the Late Triassic, and there are two records from Early Triassic rocks. The known track types include: Brachychirotherium isp., cf. Brachychirotherium isp., Chirotherium barthii , Dicynodontipus ispp., Grallator isp., Rhynchosauroides isp., Rigalites ischigualastianus , Rigalites isp., Tetrasauropus isp., and bird-like, chirotheriid and unidentified tridactyl footprints. The ichnogenera Gallegosichnus Casamiquela, 1964, Calibarichnus Casamiquela, 1964, Palaciosichnus Casamiquela, 1964 and Stipanicichnus Casamiquela, 1975 are considered to be synonyms of Dicynodontipus (Hornstein, 1876). In addition, the abandonment of the following ichnogenera (and single ichnospecies) that are based on poorly preserved material is suggested: Ingenierichnus sierrai Casamiquela, 1964, Rogerbaletichnus aguilerai Casamiquela, 1964 and Shimmelia chirotheroides Casamiquela, 1964. At least eight Triassic ichnofaunas can be recognized. The most peculiar is that of the Late Triassic Los Menucos depocentre, which is characterized by the dominance of therapsid footprints ( Dicynodontipus ispp.). The track assemblages from the Cuyo Basin display the highest ichnodiversity, with five footprint types.