The value of the archaeobotanical analysis of desiccated plant remains from old buildings: methodological aspects and interpretation of crop weed assemblages

Plant material like threshing remains has been used systematically as filling and insulation material in buildings for a long time. Such plant material can be found in several places, for example in floor cavities (internal floor deposits, Fehlböden) or in beam holes. For the first time in Switzerland, desiccated plant material like this has been investigated. In this article the analysis of the material from three different sites is presented, each dated to a different period, from the late medieval period to the time just before the industrial revolution. The samples consisted of chaff from Triticum spelta in bulk, and smaller amounts of other cereals such as T. monococcum, Avena sativa, Panicum miliaceum and Secale cereale, plus various weeds. The assemblages in the late medieval samples show evidence of three-field rotation: the high percentage of grassland species is interpreted as an indicator of the fallow land phase within the three-field system. A higher ratio of annual to perennial plants in the more modern samples indicates the development of more intensive tillage and cultivation. Methodological questions regarding sampling strategies of the floor cavity subsamples are investigated. The analyses suggest that the volume of these subsamples should be more than two litres.     

Plant husbandry and vegetation of early medieval Douai,northern France

This paper describes the results of the archaeobotanical examination of early medieval occupation deposits (8th to 11th centuries A.D.) at Douai, northern France. Carbonized as well as waterlogged seeds and fruits were recovered in great numbers. In addition to the macrofossil analyses, a palynological examination of occupation deposits was carried out. Bread wheat (Triticum aestivum) was the most common cereal, followed by rye (Secale cereale). Common oat (Avena sativa) and hulled barley (Hordeum vulgare) must have played a fairly modest role. In addition to cereals, field pea (Pisum sativum) was an important crop plant. Four types of plum (Prunus domestica) fruitstones are distinguished. Sweet cherry (Prunus avium) as well as sour cherry (P. cerasus) were cultivated. Other cultivated fruit trees included peach (Prunus persica) and medlar (Mespilus germanica). Sloe (Prunus spinosa) fruits, including those of the var. macrocarpa, were gathered from the wild. The palynological data suggest that the Douai area had virtually been cleared of forest. A comparison between pollen and seed proportions points to serious discrepancies between the macrofossil and microfossil plant records. Weeds of arable land (Secalietea, Polygono-Chenopodietalia) and vegetations of ruderal habitats (Sisymbrietalia, Artemisietalia vulgaris) are well represented. Molinietalia vegetations in the valley of the Scarpe river were probably exploited as hay meadows; good pasture land was provided by the Arrhenateretalia grasslands.     

Archaeobotanical investigations of late neolithic lakeshore settlements (Lake Biel, Switzerland)

This paper describes the results of the archaeobotanical examination of four Late Neolithic lakeshore settlements on Lake Biel in Switzerland. Due to the excellent preservation conditions in lakeshore settlements, non-carbonized as well as carbonized seeds and fruits were recovered in large numbers. In addition to the diaspore analyses, some samples of charcoal and moss were identified. The spectrum of cultivated plants was markedly different between the sites. In the transition from the 34th to 33rd century B.C. (sites Nidau, Lüscherz and Lattrigen ‘VI’) naked wheat and barley were predominant, while in the 32nd century B.C. (Lattrigen'VII') emmer (a glume wheat) was most abundant. Flax and opium poppy were of great importance during the whole late Neolithic period. In addition to cereals, a large number of cereal weeds were detected. The wild flora included a high percentage of aquatic and lake-shore plants which results from the strong influence of water on the cultural layers. Taxa of flood-plain forest are also common. The proportion of potential grassland plants was low (in total only 13 taxa) which suggests that in addition to the cultivated fields only few pastures and grassland areas existed close to the settlements.     

The history of cereals in the region of the former Duchy of Swabia (Herzogtum Schwaben) from the Roman to the Post-medieval period: results of archaeobotanical research

Archaeobotanical results from the investigations of cereal finds in southwestern Germany and northern Switzerland have been mapped. Seven phases are distinguished, from Roman to early modern (Post-medieval). Methodological problems are discussed concerning sampling, identification, preservation, type of site, feature and assemblage, which influence the representation of the results. The main results are as follows. In the Roman period, Triticum spelta was the main crop, except in the upper Rhine valley (Oberrheinebene) where T. aestivum was more abundant. In the Migration period and Early Medieval period several crops are of similar importance. The reason was a rather simple economic system, a subsistence economy for each village, and perhaps cereal and grassland rotation (Feld-Gras-Wirtschaft). In the High Medieval period, Secale cereale was the dominant grain in the northern part of the region up to the Danube (Donau). In Switzerland, T. spelta dominated. In the landscape in between, from Lake Constance (Bodensee) to the upper Neckar valley, there was a mixture of both. These rather clear spectra, commonly dominated by one cereal species, express the changed economic system: the three field system (Dreifelderwirtschaft) and an increasing role of the market economy. Even in the Late Medieval and Post-medieval periods S. cereale and T. spelta remained the main crops. The present dominance of T. aestivum and Hordeum vulgare is a very recent development, less than a century old. The less important cereals, T. monococcum and Panicum miliaceum, occur regularly until Post-medieval times. T. dicoccum was very rare in this region from the Roman period onwards.     

Ethnic groups and biological diversity in medieval Spain: analysis of a Jew Necropolis from Sevilla

The skulls and mandibles of adults individuals from a Jewish necropolis of Sevilla (La Buhayra), (V–IX Cent. A.D.) have been analyzed. The results show that Sevillian Jews, both males and females, present narrow skulls with a notable dolichocrany. Sexual differences in both the height of the skull and the mandible robustness have been found. A comparison between Sevillian Jews and other Iberian samples from different epochs (Neolithic-Bronze to Centuries A.D.) has been carried out by means of a principal component analysis taken only into account the variability between groups. Results of the analysis showed a certain degree of biological differentiation between Sevillian and Catalonian Jews (XI–XIVth Centuries A.D.) which can be justified not only by the geographical distance and the temporal gap between the two groups, but also by the intrinsic heterogeneity of Jews (Nathan, 1961a, b; Nathan et al., 1963; Arensburg et al., 1980). The existence of some degree of biological variability between several ethnic-religious groups (Jew, Christians and Moslems) can be detected although they share a common Mediterranean background. Paper presented at the 4^th congress of the European Anthropology Association (Florence, Sept. 1984).     

Grain versus chaff: identifying a contrast between grain-rich and chaff-rich sites in the Neolithic of northern Greece

Recent archaeobotanical investigations of Late Neolithic sites in northern Greece have revealed a pattern as regards the crop parts represented at each site. Some sites appear to be dominated by chaff, mainly glume wheat chaff, while other sites are mainly characterised by cereal grain and pulses. This pattern could be the outcome of various factors: a. pre-depositional differential treatment of crops selecting for one category of plant remains to arrive at the site, b. depositional, therefore related to the type of activity represented at the excavated contexts and the use of space in relation to plants, and c. post-depositional factors such as erosion, or the type of features sampled. These alternative factors are considered in light of the preliminary results of the archaeobotanical investigation of seven Late Neolithic (5500–4000 cal. B.C.) sites from northern Greece. It is suggested that this pattern is the outcome of a differential use of space in relation to plants (storage, processing/food preparation, refuse disposal), probably related to aspects of the socioeconomic organisation of the settlements.     

Nutrition, aspects of land use and environment in medieval times in southern Germany: plant macro-remain analysis from latrines (late 11th–13th century a.d.) at the town of Überlingen, Lake Constance

From high medieval courtyards at überlingen, Lake Constance (Bodensee), 17 archaeobotanical samples from seven latrines (11th–13th century) were analysed for plant macrofossils. They contained small amounts of cultivated plants and many well preserved seeds and fruits of wild plants. The most numerous cereal finds were of Triticum spelta followed by Secale cereale. Recorded oil plants were Papaver somniferum and Linum usitatissimum. Cultivated and gathered fruits such as various Prunus species, Pyrus communis and Malus domestica, Rubus sp. and Fragaria vesca were eaten by the people of überlingen. Only one seed of Ficus carica was found which may have been imported; no other imported plants could be recognised. Agrostemma githago and Vaccaria hispanica were prevalent weeds. The presence of the latter probably indicates the cultivation of cereals on the steeper slopes above überlingen. It is shown that in central Europe, V. hispanica became a troublesome weed only around the 12th/13th century A.D. Grassland species—mainly from more or less wet locations—and ruderal plants of nitrogen-rich soils were well represented in the samples and characterize the surrounding area. Electronic Supplementary Material Supplementary material is available in the online version of this article at     

Arthropod diversity of exotic vs. native Robinia species in northern Arizona

1 Arthropods were collected on native locust, Robinia neomexicana A. Gray, and exotic Robinia pseudoacacia L. in northern Arizona over a 2‐year period to determine the number of arthropod species and number of individuals present. 2 More arthropod species were found on the native (251) than on the exotic Robinia (174). 3 Greater species diversity was likewise found on the native than the exotic. The five most numerous insects collected each year accounted for 81% to 91% of the total number collected on the exotic and native Robinia in 1997 and 1998. Only 12 species occurred on both the native and exotic Robinia in both years. 4 These findings are discussed in the context of using exotic trees in plantations and ecological theory regarding rates of arthropod species accumulation on exotic hosts.     

Plant remains from the early medieval lakeshore settlement Fonyód-Bélatelep (Lake Balaton,Hungary) with especial emphasis on the history of fruit cultivation in Pannonia

During the excavation of an early medieval lakeside settlement at Fonyód-Bélatelep, Balaton (7th-9th century A.D.), remains of cultivated and wild plants were found in the culture layer. The main cereal crops of the Avar culture were Hordeum, Triticum and Secale. Panicum also played an important role. In addition to the cereals, 13 species of fruit trees were present. It appears that fruit cultivation survived from Roman times into the early medieval period.     

Plant economy and village life in Neolithic lake dwellings at the time of the Alpine Iceman

This paper gives an overview of domesticated and wild plants used during the second half of the fourth millennium cal b.c. in a region within a semicircle of ca. 100 km radius around the find site of the Iceman. The landscape considered lies north of the main Alpine chain and includes central to eastern Switzerland and southwest Germany, mainly sites at the lakes Zug, Zürich, Constance (Bodensee), the Federsee region of Upper Swabia (Oberschwaben) and western Bavaria. From this part of central Europe, the plant remains of many well-preserved lake dwelling sites have been investigated during the last decades and much is known about daily life in these villages. Probably contemporary with the Iceman’s lifetime is one of the best-investigated settlements, Arbon Bleiche 3 on the southern shore of Lake Constance. A comparison of the Iceman’s artefacts and plant macro remains with the finds from the lake dwellings which have been studied shows striking similarities, but also differences. The Iceman’s southern provenance can be corroborated by the domestic plants found with him. He used domestic as well as wild plants, very similar to those of his contemporaries in the lakeside villages. The detailed knowledge of the manifold activities in the lakeshore village of Arbon Bleiche 3 allows us to show a rather realistic picture of daily life at the time of the Iceman.     

药用植物多样性特征分析——以陕北米脂县为例

为研究陕北米脂县野生药用植物资源现状及其分布规律,通过野外调查及查阅文献等方法,对该地区药用植物多样性及其特征进行研究。结果显示：（1）米脂县共有野生药用植物45科110属148种,其中裸子植物2科2属2种、被子植物43科108属146种,集中分布于单种科和寡种科及单种属内;（2）从生长型来看,草本植物最多,其次是乔木,灌木最少;（3）药用部位以全草类、根及根茎类最多,分别占药用植物总种数的52.03%和14.86%;（4）药用功效以清热和利水渗湿为主,分别占药用植物总数的23.65%和11.49%;（5）从药用植物属的分布类型来看,有13个分布区类型8个变型,区系成分复杂多样,温带性质明显。本研究表明米脂县药用植物资源多样性较高,生长型、药用部位、功效、区系成分丰富多样,明确了该地区野生药用植物资源的分布及生存现状,为加强资源的保护及开发利用提供了依据。     

Plant species diversity as a driver of early succession in abandoned fields: a multi-site approach

Succession is one of the most studied processes in ecology and succession theory provides strong predictability. However, few attempts have been made to influence the course of succession thereby testing the hypothesis that passing through one stage is essential before entering the next one. At each stage of succession ecosystem processes may be affected by the diversity of species present, but there is little empirical evidence showing that plant species diversity may affect succession. On ex-arable land, a major constraint of vegetation succession is the dominance of perennial early-successional (arable weed) species. Our aim was to change the initial vegetation succession by the direct sowing of later-successional plant species. The hypothesis was tested that a diverse plant species mixture would be more successful in weed suppression than species-poor mixtures. In order to provide a robust test including a wide range of environmental conditions and plant species, experiments were carried out at five sites across Europe. At each site, an identical experiment was set up, albeit that the plant species composition of the sown mixtures differed from site to site. Results of the 2-year study showed that diverse plant species mixtures were more effective at reducing the number of natural colonisers (mainly weeds from the seed bank) than the average low-diversity treatment. However, the effect of the low-diversity treatment depended on the composition of the species mixture. Thus, the effect of enhanced species diversity strongly depended on the species composition of the low-diversity treatments used for comparison. The effects of high-diversity plant species mixtures on weed suppression differed between sites. Low-productivity sites gave the weakest response to the diversity treatments. These differences among sites did not change the general pattern. The present results have implications for understanding biological invasions. It has been hypothesised that alien species are more likely to invade species-poor communities than communities with high diversity. However, our results show that the identity of the local species matters. This may explain, at least partly, controversial results of studies on the relation between local diversity and the probability of being invaded by aliens. Received: 13 July 1999 / Accepted: 4 February 2000     

Plant remains from a Bell Beaker site in Switzerland, and the beginnings of Triticum spelta (spelt) cultivation in Europe

Little is known of the archaeobotany of the Bell Beaker period. The village of Cortaillod/Sur les Rochettes-est in Switzerland is one of the first settlements of this date discovered so far; and one of the few sites where systematic archaeobotanical research could be undertaken. The 114 features, mainly postholes, produced 5080 charred plant macrofossils. The composition of the cereal spectrum was dominated by hulled wheats and notably by Triticum spelta (spelt). Other important components were Triticum dicoccum (emmer) and T. monococcum (einkorn), and probably also Hordeum (barley). The abundance of spelt represents a fundamental change compared with the plant finds from earlier sites of the region. The origin of this cereal species is discussed in the light of recent results of cytogenetic and molecular research.     

New primates from the late Eocene of Thailand: a contribution to primate diversity in the Paleogene of Asia

We report the discovery of two new primates from the late Eocene Krabi Basin, southern Thailand. One isolated upper molar displays morphological features (protocone united with hypocone by the prehypocrista, postprotocrista extended distobuccally) suggesting possible phylogenetic relationships with Amphipithecidae, while an isolated lower molar is tentatively referred to as a new tarsiiform, mainly on the basis of its paraconid and entoconid morphology. Although very scarce, these remains expand the record of Paleogene primates in Southeast Asia, and testify to their successful radiation in that area.     

中国南海北部陆坡沉积物古菌多样性及丰度分析

【目的】海洋沉积物中的古菌在全球生物地球化学循环中充当重要的角色,深入了解沉积物中古菌群落的结构及功能特征是探究海洋沉积物中古菌参与生物地球化学循环和生态学功能的基础。【方法】采用高通量测序技术,分别对南海北部陆坡不同海域(东部,西部和神狐海域的7个站位)沉积物中古菌16SrRNA基因进行Illumina Mi Seq测序。【结果】中国南海北部陆坡沉积物中古菌的主要门类是Bathyarchaeota、Thermoplasmata、Woesearchaeota(DHVEG-6)、Thaumarchaeota(Marine Group I)、Lokiarchaeota和Marine Hydrothermal Vent Group(MHVG),还存在少量的AK8、Marine Benthic Group A和Terrestrial Hot Spring Crenarchaeota Group(THSCG)等。在潜在水合物区沉积物中还发现了甲烷代谢相关古菌(Anaerobic methanotrophic archaea,ANME)类群,主要为ANME-1、ANME-2ab和ANME-2c等。甲烷代谢古菌的分布特征也从甲烷代谢保守功能基因mcr A(Methyl coenzyme-Mreductase A)的扩增中得到了验证。利用定量PCR对南海沉积物中的细菌、古菌的16SrRNA基因和mcrA基因进行了定量,发现细菌16SrRNA基因拷贝数为10~5-10~7 copies/g(湿重),古菌16SrRNA基因拷贝数为10~5-10~6 copies/g(湿重),潜在水合物区mcrA基因拷贝数为10~3-10~5 copies/g(湿重)。【结论】揭示了中国南海北部陆坡沉积物中具有丰富的微生物资源,其中古菌种类多样且丰度较高,同时发现冷泉特征古菌群落,为深入认识和理解南海沉积物中微生物丰度和古菌多样性,以及解析古菌地球化学功能奠定基础。     

青藏高原北部多年冻土区草地植物多样性

研究了青藏高原北部多年冻土区草地群落植物多样性的特征。研究表明 :草地群落间丰富度指数差异不显著 ,均匀度指数和多样性指数差异显著 (P<0 .0 5 )。均匀度指数表现为高山嵩草 (Kobresia pygmaea)草甸 <紫花针茅 (Stipa purpurea)草原 <矮嵩草 (K.humilis)草甸 <青藏苔草 (Carex moorcroftii)草甸 ,多样性指数表现为高山嵩草草甸 <矮嵩草草甸 <紫花针茅草原<青藏苔草草甸。修路时破坏的矮嵩草草甸在次生恢复过程中 ,离公路 10 0 m处群落的丰富度指数 ,均匀度指数和多样性指数大于原生群落 ,而原生群落的多样性又大于 30 m和 5 0 m处群落的多样性。地上草地群落植物多样性伴随地下冻土退化过程表现为 ,以 1m2样方统计时 ,各个演替群落间的丰富度指数差异不显著 ,而以 10 0 m2样条统计时 ,高寒草甸和草原化草甸的丰富度指数显著大于沼泽草甸和稀疏草原 (P<0 .0 5 ) ,但均匀度和多样性指数在两种统计面积时均表现为先增加后下降的变化趋势。     

Plant growth hormones suppress the development of Harpophora maydis,the cause of late wilt in maize

Late wilt, a severe vascular disease of maize caused by the fungus Harpophora maydis, is characterized by rapid wilting of maize plants before tasseling and until shortly before maturity. The pathogen is currently controlled by resistant maize cultivars, but the disease is constantly spreading to new areas. The plant’s late phenological stage at which the disease appears suggests that plant hormones may be involved in the pathogenesis. This work revealed that plant growth hormones, auxin (Indole-3-acetic acid) and cytokinin (kinetin), suppress H. maydis in culture media and in a detached root assay. Kinetin, and even more auxin, caused significant suppression of fungus spore germination. Gibberellic acid did not alter colony growth rate but had a signal suppressive effect on the pathogens’ spore germination. In comparison, ethylene and jasmonic acid, plant senescing and defense response regulators, had minor effects on colony growth and spore germination rate. Their associate hormone, salicylic acid, had a moderate suppressive effect on spore germination and colony growth rate, and a strong influence when combined with auxin. Despite the anti-fungal auxin success in vitro, field experiments with dimethylamine salt of  2,4-dichlorophenoxyacetic acid (that mimics the influence of auxin) failed to suppress the late wilt. The lines of evidence presented here reveal the suppressive influence of the three growth hormones studied on fungal development and are important to encourage further and more in-depth examinations of this intriguing hormonal complex regulatory and its role in the maize-H. maydis interactions.     

Branch wood from the lake shore settlements of Horgen Scheller, Switzerland: Evidence for economic specialization in the late Neolithic period

596 waterlogged, uncarbonized branch wood specimens from four cultural layers of the late Neolithic lake shore site of Horgen Scheller (between 3080 and 3030 B.C.) on Lake Zilrich, Switzerland, were examined. The following parameters were analysed: taxonomy, diameter and age of the twigs, cutting season, and distribution of the branches over the area as well as their distribution in the different cultural layers. The twigs were divided into the following three groups, based on differences observed in these analyses: conifers, deciduous trees with catkins, and deciduous trees without catkins. Branch diameter examinations revealed that branches of deciduous trees were thicker on average than those of conifers. While branches from deciduous trees without catkins were gathered at the age of seven years on average, those of conifers and deciduous trees with catkins were older. Waney edge analyses showed that the majority of the twigs were cut within the winter months. Among the deciduous trees without catkins there is, however, a markedly higher proportion of branches cut in summer than in the other two groups. The taxonomic examinations demonstrated that 70% of the specimens consisted of twigs of eitherCorylus avellana L. (hazel),Taxus baccata L. (yew), orAbies alba Mill. (silver fir), thus implying the great economic importance of these species in the settlements of Horgen Scheller. To our knowledge this is the first report that a remarkably high proportion of yew twigs (26.8%) has been found in an analysis of subfossil twigs. The possibility is discussed that the inhabitants had specialized in obtaining yew from the region. Concentrations of wood finds, possible remains of buildings, became evident from the analyses of the distribution of the branch wood over the excavated area. A very similar spatial distribution pattern of the taxa was found through the different cultural layers. Therefore it seems likely that existing structures were reused when a new settlement was built. The data also show that the building outlines slowly moved towards land over time. A rising lake water level could have led to gradual dislocation of the settlements.