孟家岗林场森林景观结构动态分析

以黑龙江省孟家岗林场1989年和2006年两期森林资源二类调查数据为基础,结合林相图,利用GIS技术和景观格局分析软件包FRAGSTATS进行分析,提取森林景观要素内部结构指标等景观指数,并对孟家岗林场近20年的景观格局的动态特征进行了系统的分析。研究结果表明：由于孟家岗林场森林景观要素类型的增加以及多样性和优势度指数的增高,使得森林景观的分离度和破碎度也在增加;虽然落叶松林为该研究区域的优势景观要素类型,但其优势度却在逐渐降低,表明落叶松景观要素在向其它景观要素类型转化;首次对孟家岗林场进行景观内部结构指数分析,表明阔叶树和混交林景观要素类型是以成、过熟林为主,三大主要针叶树种以中龄林和近熟林为主。人为干扰的加剧,使针叶林年龄结构趋于单一化。     

快速城市化阶段上海海岸带景观格局的时空动态

海岸带景观动态是海岸带综合管理的重要方面.本文利用多时相遥感影像解译的土地利用数据研究了1994-2005年快速城市化阶段上海海岸带的景观动态变化特征.结果表明:人类干扰导致了海岸带景观的破碎化程度增加,斑块隔离程度加大,景观多样性增大;海岸带基质景观(即农田)的面积持续减少,优势度明显降低,破碎化加大,反映城市化的人工景观类型(如工业用地、交通用地、公共设施用地、居住用地)面积不断增加,成为第二大景观类型,其他用地(主要是在建用地)的面积不断增加,显示出海岸带仍在经历快速城市化过程;导致海岸带景观变化的驱动力主要是国家政策、地方政府的决策、上海市的快速城市化,以及沿海大型开发项目(如浦东国际机场、洋山国际深水港、上海化学工业区等).在分析海岸带城市化带来的生态环境后果的基础上,建议实施综合海岸带管理以实现海岸带的可持续发展.

Abstract：

Coastal zone's landscape dynamics is recognized as one of the important aspects of in-tegrated coastal management. In this paper, the land use datasets derived from time series aerial images were employed to study the coastal landscape dynamics of Shanghai from 1994 to 2005, the period of rapid urbanization. During this period, human disturbances caused landscape frag-mentation and isolation, and increased landscape diversity. The area of main landscape (agricul-tural land) continued to decrease, its dominance declined, and became more fragmentation; while the areas of artificial landscapes reflecting urbanization (e.g.,lands for industry, traffic, public facility, and residence) increased, becoming the second dominant landscape type. The increasing area of other land uses (mainly constructing sites) implied that this coastal zone was still undergoing rapid urbanization. Central government policies, local government decision-mak-ing, rapid urbanization of Shanghai, and coastal big-scale development programs such as Pudong International Airport, Yangshan International Harbor, and Shanghai Chemical Industry Park were the main driving forces for these coastal landscape changes. Based on the analysis of the ecologi-cal consequences of this rapid urbanization, some suggestions for integrated management were made to promote the sustainable development of Shanghai coastal zone.     

毛乌素沙地景观格局变化研究

通过对毛乌素沙地不同时期的景观分类和制图以及景观格局计算,分析了50年代到90年代毛乌素沙地景观动态与景观格局变化的特征和模式。研究表明,50年代以来,毛乌索沙地景观发生了巨大变化,流动沙地、半固定沙地面积大幅度增加,固定沙地面积迅速减少,农田面积明显增加,草原、柳湾和盐湿低地等面积显著减少。50年代时,整个景观以流动沙地和固定沙地占优势,而90年代,则以流动沙地占绝对优势。伴随着荒漠化的发展,各种景观要素类型之间相互转化,景观格局发生显著变化。固定沙地面积减少伴随着斑块的破碎化,大型斑块数量减少,小型斑块数量增加;流动沙地面积增加的同时,一方面,较小的斑块不断联合形成更多的大型斑块,另一方面,流动沙地不断向固定、半固定沙地内部渗透,形成更多的小型斑块,使整个景观更加破碎。不合理的人类活动是荒漠化扩展及景观格局发生显著变化的主要驱动因素。     

绿洲景观动态变化分析

以新疆石河子150团场为研究地区,对其1982－1995年景观动态及其原因进行了研究。结果表明,景观变化主要体现在以下方面：首先是景观基质由耕地转变为荒草地;其次,从嵌块体类型看,在绝对面积上,荒草地、林地和居民点是增加的,其余的嵌块体类型均呈现出减少的趋势。第三从嵌块体的转换过程看,菜地、弃耕地、园地和沙地的转化率最高,均在40％以上,而荒草地和耕地的转化率无在20％左右。就其原因而言,主要有人工绿洲垦殖过程中非人力为主观原因引起的绿洲地下水高水位区的形成,以及人口变化、粮食单产提高和经济环境变动等。     

一种新的景观扩张指数的定义与实现

景观格局动态信息的定量表达始终是景观生态学研究的一个重要科学问题,景观格局指数是其中的一种重要方法,但其多是静态指数,难以有效定量表达景观格局的动态信息.因此,针对景观扩张过程以斑块扩张面积为基础提出了一种新的景观扩张指数,来表达景观格局的动态信息.并以妫水河流域1998-2009年的景观农田化过程为例,验证该指数的适用性,结果表明:该指数不仅能够定量表达斑块的空间扩张规模,而且可以准确识别斑块的空间扩张模式.根据扩张斑块与原斑块的空间位置关系,将景观的空间扩张模式划分为邻接扩张式和外部扩张式两种.提出的景观扩张指数在技术方法上计算简便,易于实现,完善了景观格局动态的量化表征科学方法.     

A modeling framework for the establishment and spread of invasive species in heterogeneous environments

Natural and human‐induced events are continuously altering the structure of our landscapes and as a result impacting the spatial relationships between individual landscape elements and the species living in the area. Yet, only recently has the influence of the surrounding landscape on invasive species spread started to be considered. The scientific community increasingly recognizes the need for broader modeling framework that focuses on cross‐study comparisons at different spatiotemporal scales. Using two illustrative examples, we introduce a general modeling framework that allows for a systematic investigation of the effect of habitat change on invasive species establishment and spread. The essential parts of the framework are (i) a mechanistic spatially explicit model (a modular dispersal framework—MDIG ) that allows population dynamics and dispersal to be modeled in a geographical information system (GIS ), (ii) a landscape generator that allows replicated landscape patterns with partially controllable spatial properties to be generated, and (iii) landscape metrics that depict the essential aspects of landscape with which dispersal and demographic processes interact. The modeling framework provides functionality for a wide variety of applications ranging from predictions of the spatiotemporal spread of real species and comparison of potential management strategies, to theoretical investigation of the effect of habitat change on population dynamics. Such a framework allows to quantify how small‐grain landscape characteristics, such as habitat size and habitat connectivity, interact with life‐history traits to determine the dynamics of invasive species spread in fragmented landscape. As such, it will give deeper insights into species traits and landscape features that lead to establishment and spread success and may be key to preventing new incursions and the development of efficient monitoring, surveillance, control or eradication programs.     

Heterogeneous folding of the trpzip hairpin: full atom simulation and experiment

The beta-hairpin trpzip2 can be tuned continuously from a two-state folder to folding on a rough energy landscape without a dominant refolding barrier. At high denaturant concentration, this extremely stable peptide exhibits a single apparent "two-state" transition temperature when monitored by different spectroscopic techniques. However, under optimal folding conditions the hairpin undergoes an unusual folding process with three clusters of melting transitions ranging from 15 degrees C to 160 degrees C, as monitored by 12 different experimental and computational observables. We explain this behavior in terms of a rough free energy landscape of the unfolded peptide caused by multiple tryptophan interactions and alternative backbone conformations. The landscape is mapped out by potentials of mean force derived from replica-exchange molecular dynamics simulations. Implications for deducing cooperativity from denaturant titrations, for the origin of folding cooperativity, and for the folding of thermophilic proteins are pointed out. trpzip is an excellent small tunable model system for the glass-like folding transitions predicted by landscape theory.     

Response of subarctic vegetation to transient climatic change on the Seward Peninsula in north-west Alaska

Understanding the response of terrestrial ecosystems to climatic warming is a challenge because of the complex interactions of climate, disturbance, and recruitment across the landscape. We use a spatially explicit model (ALFRESCO) to simulate the transient response of subarctic vegetation to climatic warming on the Seward Peninsula (80 000 km²) in north‐west Alaska. Model calibration efforts showed that fire ignition was less sensitive than fire spread to regional climate (temperature and precipitation). In the model simulations a warming climate led to slightly more fires and much larger fires and expansion of forest into previously treeless tundra. Vegetation and fire regime continued to change for centuries after cessation of the simulated climate warming. Flammability increased rapidly in direct response to climate warming and more gradually in response to climate‐induced vegetation change. In the simulations warming caused as much as a 228% increase in the total area burned per decade, leading to an increasingly early successional and more homogenous deciduous forest‐dominated landscape. A single transient 40‐y drought led to the development of a novel grassland–steppe ecosystem that persisted indefinitely and caused permanent increases in fires in both the grassland and adjacent vegetation. These simulated changes in vegetation and disturbance dynamics under a warming climate have important implications for regional carbon budgets and biotic feedbacks to regional climate.     

Test systems to determine the ecological risks posed by toxin release from Bacillus thuringiensis genes in crop plants

Procedures for the selection of species for ecotoxicological risk assessment of Bacillus thuringiensis (Bt) gene products in the epigeal and hypogeal environments are proposed. Although species can be selected on the basis of ecological realism and functional importance, the number of organisms requiring testing and the nature of the test procedures remain uncertain with such a selectively toxic material. The heterogeneity of the soil environment, the stratification of plant material at different stages of breakdown and decomposition and the aggregation and patterns of movement of the soil fauna and flora impose problems for the design of ecologically relevant test methods. Similarly, the impact upon beneficial invertebrates, if toxic effects are detected, will be mediated by the scale and pattern of transgenic plant release in the fragmented agricultural landscape. To properly assess the ecological risks posed by a widely released toxin with a narrow spectrum of effects, a combination of laboratory tests, field experiments and longer-term monitoring will be required.     

异质种群动态模型:破碎化景观动态模拟的新途径

景观破碎化导致物种以异质种群方式存活,使得基于异质种群动态模拟破碎化景观动态成为可能。异质种群动态模型的发展为景观动态模拟奠定了良好基础。根据空间处理方式的不同,异质种群模型可分为三大类,可不同程度地用于描述破碎化景观动态。(1)空间不确定异质种群模型,假定所有局域种群间均等互联,模型中不包含空间信息,仅能用于景观斑块动态描述;(2)空间确定异质种群模型,假设局域种群在二维空间上以规则格子形式排列,是一种准现实的空间处理方式,可用于景观动态的简单描述;(3)空间现实异质种群模型,包含了破碎化景观中局域种群的几何特征,可直接用于真实景观动态的模拟研究。空间现实的和基于个体的异质种群模型不但是未来异质种群模型发展的主流,也将成为未来破碎化景观动态研究的重要工具。为了更加准确完整地描述破碎化景观动态,不但应该综合运用已有的各种异质种群模型方法,更要引进新模型来刎画多物种、多变量、高维度、复杂连接的破碎化景观格局与过程。     

Genetic structure of the forest herb Primula elatior in a changing landscape

To investigate whether changes in land use and associated forest patch turnover affected genetic diversity and structure of the forest herb Primula elatior, historical data on landscape changes were combined with a population genetic analysis using dominant amplified fragment length polymorphism markers. Based on nine topographic maps, landscape history was reconstructed and forest patches were assigned to two age classes: young (less than 35 years) and old (more than 35 years). The level of differentiation among Primula populations in recently established patches was compared with the level of differentiation among populations in older patches. Genetic diversity was independent of population size (P > 0.05). Most genetic variation was present within populations. Within-population diversity levels tended to be higher for populations located in older forests compared with those for populations located in young forests (Hj = 0.297 and 0.285, respectively). Total gene diversity was also higher for old than for young populations (Ht = 0.2987 and 0.2828, respectively). The global fixation index FST averaged over loci was low, but significant. Populations in older patches were significantly more differentiated from each other than were populations in recently established patches and they showed significant isolation by distance. In contrast, no significant correlations between pairwise geographical distance and FST were found for populations in recently established patches. The location of young and old populations in the studied system and altered gene flow because of increased population density and decreased inter-patch distances between extant populations may explain the observed lower genetic differentiation in the younger populations. This study exemplifies the importance of incorporating data on historical landscape changes in population genetic research at the landscape scale.     

热带森林中的斑块动态与物种多样性维持

斑块作为景观要素之一,直接到景观结构的空间格局及其内部各要素之间的相互关系,其动态也将导致景观格局的变化。异质性的斑块在自然森林中是普遍存在的,这就是说,顶极森林中仍然存在着由不同种类或不同生长时期的植物种群组成的森林斑块。由自然或人为干扰所驱动的森林生长循环导致敢这些斑块在空间上的镶嵌,对于持定的地域片段,也导致了不同生长时期的森林斑块的周期性循环。在热带森林中,森林的生长循环由林窗期（ｇａｐ     

Lake Ladoga Region: human impacts and recent environmental changes

This paper presents a landscape-ecological analysis of Ladoga Region, a 30 000 km² territory around Lake Ladoga, which is the largest lake in Europe. The Region includes the contact zone between the crystalline bedrock of Baltic Shield and the sedimentary rocks of the East European plain, and the transition from Boreonemoral to Southern Boreal (Taiga) forest vegetation; it is characterized by abundance of lakes. The Region is divided into 17 discrete landscape areas that are classified in 6 groups.The author uses the landscape-dynamcal approach to the analysis of recent (from 2–3 to 40–50 yr) environmental changes in the region, caused by human activities: forest felling, fires, peat excavation, agriculture, recreational development etc. The following processes are observed: vegetation successions after clearcutting and selective logging; effects of forest fires on the spreading and conservation of pine forests; consequences of peatland drainage; landscape effects of collective farming and gardening; forest succession from pine and deciduous tree dominance towards spruce dominance (observable in all landscape regions); natural vegetation succession on former Finnish arable lands (two types of scenarios: forest succession or development of sedge meadows).     

深圳市1988—2007年间湿地景观动态变化及成因分析

城市湿地是城市生态安全体系构成的重要组成部分。以深圳地区为工作区,利用多时段遥感资料和土地利用变更调查数据,进行1988—2007年期间工作区内湿地景观动态变化特征分析,探讨快速城市化背景下湿地景观动态变化的成因。结果表明,研究时段内深圳市湿地总面积损失了35.7%,不同湿地类型及不同行政单元内湿地的动态变化表现出明显差异。建设用地扩张是工作区内湿地损失的主要原因并且其影响表现为一种非均匀过程;功能差异是导致不同湿地类型之间动态变化差异的主要成因;区域社会经济和自然环境条件差异是导致不同行政单元之间湿地动态变化差异的主要影响因素。     

Probing the high energy states in proteins by proteolysis

Unless the native conformation has an unstructured region, proteases cannot effectively digest a protein under native conditions. Digestion must occur from a higher energy form, when at least some part of the protein is exposed to solvent and becomes accessible by proteases. Monitoring the kinetics and denaturant dependence of proteolysis under native conditions yields insight into the mechanism of proteolysis as well as these high-energy conformations. We propose here a generalized approach to exploit proteolysis as a tool to probe high-energy states in proteins. This "native state proteolysis" experiment was carried out on Escherichia coli ribonuclease HI. Mass spectrometry and N-terminal sequencing showed that thermolysin cleaves the peptide bond between Thr92 and Ala93 in an extended loop region of the protein. By comparing the proteolysis rate of the folded protein and a peptidic substrate mimicking the sequence at the cleavage site, the energy required to reach the susceptible state (Delta G(proteolysis)) was determined. From the denaturant dependence of Delta G(proteolysis), we determined that thermolysin digests this protein through a local fluctuation, i.e. localized unfolding with minimal change in solvent assessable surface area. Proteolytic susceptibilities of proteins are discussed based on the finding of this local fluctuation mechanism for proteolysis under native conditions.     

Anthropogenic disturbance and the formation of oak savanna in central Kentucky, USA

Aim To deepen understanding of the factors that influenced the formation of oak savanna in central Kentucky, USA. Particular attention was focused on the link between historical disturbance and the formation of savanna ecosystem structure. Location Central Kentucky, USA. Methods We used dendrochronological analysis of tree‐ring samples to understand the historical growth environment of remnant savanna stems. We used release detection and branch‐establishment dates to evaluate changes in tree growth and the establishment of savanna physiognomy. We contrasted our growth chronology with reference chronologies for regional tree growth, climate and human population dynamics. Results Trees growing in Kentucky Inner Bluegrass Region (IBR) savanna remnants exhibited a period of suppression, extending from the establishment date of the tree to release events that occurred c. 1800. This release resulted in a tripling of the annual radial growth rate from levels typical of oaks suppressed under a forest canopy (< 1 mm year^?1) to levels typical of open‐grown stems (3 mm year^?1). The growth releases in savanna trees coincided with low branch establishment. Over the release period, climatic conditions remained relatively constant and growth in regional forest trees was even; however, the growth increase in savanna stems was strongly correlated with a marked increase in Euro‐American population density in the region. Main conclusions Our data suggest that trees growing in savanna remnants originated in the understorey of a closed canopy forest. We hypothesize that Euro‐American land clearing to create pasturelands released these trees from light competition and resulted in the savanna physiognomy that is apparent in remnant stands in the IBR. Although our data suggest that savanna trees originated in a forest understorey, this system structure itself may have been a result of an unprecedented lack of Native American activity in the region due to population loss associated with pandemics brought to North America by Euro‐Americans. We present a hypothetical model that links human population dynamics, land‐use activities and ecosystem structure. Our model focuses on the following three land‐use eras: Native American habitation/utilization; land abandonment; and Euro‐American land clearance. Ecological understanding of historical dynamics in other ecosystems of eastern North America may be enhanced through recognition of these eras.     

近40年艾比湖湿地自然保护区生态干扰度时空动态及景观格局变化

以新疆艾比湖湿地为研究区,利用1972、1998、2007年及2013年4个时期的Landsat遥感影像作为数据源,并结合湿地的土地覆被状况,参考《全国土地利用分类》建立艾比湖湿地生态干扰类型分类系统。借助生态干扰度指数、景观格局指数以及GIS空间分析方法,探讨艾比湖湿地的生态干扰度的时空动态及景观响应机制。结果表明:(1)1972—2013年,研究区的生态干扰度总体呈现较为稳定的趋势,但其空间分布发生变化。生态干扰度类型之间的转化速率有加快的趋势。(2)1972—2013年,边缘密度指数(ED),平均形状指数(MSI),面积加权的平均斑块分形指数(AWMPFD)及景观分离度(DIVISION)4项景观格局指数大体呈上升的趋势,2013年区域的景观指数较为稳定。(3)景观格局指数与生态干扰程度有密切的一致性。生态干扰度与景观格局指数空间分布相关性大小依次为:边缘密度指数(ED)景观丰度密度(PRD)香农多样性指数(SHDI)平均形状指数(MSI)面积加权的平均斑块分形指数(AWMPFD)景观分离度(DIVISION)。客观系统的认识和评价艾比湖湿地的生态系统及环境,可为干旱区实现自然环境的保护,协调土地利用及环境保护之间的关系提供较为实用的参考。     

Contagious Disturbance, Ecological Memory, and the Emergence of Landscape Pattern

Landscapes are strongly shaped by the degree of interaction between pattern and process. This paper examines how ecological memory, the degree to which an ecological process is shaped by its past modifications of a landscape, influences landscape dynamics. I use a simulation model to examine how ecological memory shapes the landscape dynamics produced by the interaction of vegetative regrowth and fire. The model illustrated that increased ecological memory increased the strength and spatial extent of landscape pattern. The extent of these changes depended upon the relative rates of vegetative recovery and fire initiation. When ecological memory is strong, landscape pattern is persistent; pattern tends to be maintained rather than destroyed by fire. The generality of the simulation model suggests that these results may also apply to disturbance processes other than fire. The existence of ecological memory in ecosystems may allow processes to produce ecological pattern that can entrain other ecosystem variables. The methods presented in this paper to analyze pattern in model ecosystems could be used to detect such pattern in actual ecosystems. Received 14 November 2000; accepted 21 September 2001.