The Origin and Early Radiation of Archosauriforms: Integrating the Skeletal and Footprint Record

We present a holistic approach to the study of early archosauriform evolution by integrating body and track records. The ichnological record supports a Late Permian–Early Triassic radiation of archosauriforms not well documented by skeletal material, and new footprints from the Upper Permian of the southern Alps (Italy) provide evidence for a diversity not yet sampled by body fossils. The integrative study of body fossil and footprint data supports the hypothesis that archosauriforms had already undergone substantial taxonomic diversification by the Late Permian and that by the Early Triassic archosauromorphs attained a broad geographical distribution over most parts of Pangea. Analysis of body size, as deduced from track size, suggests that archosauriform average body size did not change significantly from the Late Permian to the Early Triassic. A survey of facies yielding both skeletal and track record indicate an ecological preference for inland fluvial (lacustrine) environments for early archosauromorphs. Finally, although more data is needed, Late Permian chirotheriid imprints suggest a shift from sprawling to erect posture in archosauriforms before the end-Permian mass extinction event. We highlight the importance of approaching palaeobiological questions by using all available sources of data, specifically through integrating the body and track fossil record.     

Paleogeographic significance of Early Permian crinoids and blastoids from Oman

Early Permian crinoids and blastoids from Oman show relationship with Late Paleozoic Tethyan faunas of Timor and Western Australia and support an Early Permian age for part of the Basleo fauna of Timor. The camerate crinoidPlatycrinites omanensis n. sp. and the blastoidsTimoroblastus andDeltoblastus are reported for the first time from Sakmarian strata of northeastern Oman, doubling the known Permian echinoderms from Oman. The blastoids suggest an offshore lower energy shelf environment of deposition.      

Global deglaciation and the re‐appearance of microbial matground‐dominated ecosystems in the late Paleozoic of Gondwana

The extensive matgrounds in Carboniferous‐Permian open‐marine deposits of western Argentina constitute an anachronistic facies, because with the onset of penetrative bioturbation during the early Paleozoic microbial mats essentially disappeared from these settings. Abundant microbially induced sedimentary structures in the Argentinean deposits are coincident with the disappearance of trace and body fossils in the succession and with a landward facies shift indicative of transgressive conditions. Deposits of the Late Carboniferous–Early Permian glacial event are well developed in adjacent basins in eastern Argentina, Brazil, South Africa and Antarctica, but do not occur in the western Andean basins of Argentina. However, the deglaciation phase is indirectly recorded in the studied region by a rapid rise in sea level referred to as the Stephanian‐Asselian transgression. We suggest that an unusual release of meltwater during the final deglaciation episode of the Gondwana Ice Age may have dramatically freshened peri‐Gondwanan seas, impacting negatively on coastal and shallow‐marine benthic faunas. Suppression of bioturbation was therefore conducive to a brief re‐appearance of matground‐dominated ecosystems, reminiscent of those in the precambrian. Bioturbation is essential for ecosystem performance and plays a major role in ocean and sediment geochemistry. Accordingly, the decimation of the mixed layer during deglaciation in the Gondwana basins may have altered ecosystem functioning and geochemical cycling.     

Mesozoic tectono-sedimentary evolution of Rocca Busambra in western Sicily

The Rocca Busambra ridge in western Sicily is a shallow to pelagic Meso-Cenozoic carbonate structural unit of the Sicilian Chain with a variety of tectono-sedimentary features. Palaeofaults, unconformities (buttress unconformity, onlap, downlap), a network of neptunian dykes with several infilling generations, several large hiatuses, different facies and lateral facies changes, and erosional submarine and subaerial surfaces are observed. Detailed fieldwork and structural analyses have indicated the occurrence of fault planes with different orientations. These data, combined with facies studies and physical-stratigraphy analyses, allow for the distinction of different depositional regions. A lateral change from an open-marine carbonate platform with a stepped fault margin (located in the westernmost sector) to a deeper basinal depositional setting in the east, in the context of an upper slope scalloped margin and base-of-slope systems with talus breccias, is envisaged here. Extensional to transtensional tectonic pulses punctuated the sedimentary evolution during Early Toarcian, Late Jurassic, Early Cretaceous, Late Cretaceous, and Early Miocene times. The collected data show that most fault planes have preserved their original orientations throughout the reactivation processes. The reconstructed Meso-Cenozoic tectono-sedimentary evolution is closely related to the late syn-rift and post-rift tectonic evolution of the Tethyan continental margin.     

西藏申扎地区中二叠世(竹蜓)类动物群

结合新采集的材料与前人研究,综合分析拉萨地块申扎永珠乡下拉组中类动物群的构成与时代,并将其与同期华南类动物群进行对比。分析结果表明,申扎下拉组的类动物群共16属(含1亚属,2存疑属),70种(含比较种与未定种),时代为茅口期,大致相当于Roadian阶上部至Wordian阶。与同期华夏区华南的类动物群相比,下拉组的类动物群在属种构成,尤其是优势类群方面差异显著,其主要特征为Neoschwagerinidae与Verbeekinidae两科的种级分异度很低,Chusenella与Nankinella占优势地位,且缺失了约60%华南的常见属。     

滇西保山地区石炭纪、二叠纪古动物地理演化

探讨滇西保山地块晚古生代Ｔｉｎｇ类、有孔虫、珊瑚、牙形刺、腕足类等动物群的古生物地理属性,根据牙形刺和Ｔｉｎｇ类化石,确定长期争论的丁家寨组的时代为Ａｒｔｉｎｓｋｉａｎ期,小型单体珊瑚Ｃｙａｔｈａｘｏｎｉａ动物群可出现在从早石炭世到二叠纪的多种沉积环境中,不一定指示冷水冈瓦纳型。根据沉积特征及对环境特别敏感的珊瑚和Ｔｉｎｇ类动物群的分布特点,结合全球构造事件,恢复保山地块的古地理演化模式。早石炭世     

Early Permian facies and paleogeography of the Southeastern Russian craton

Summary During the Early Permian deep-water basins existed in the southeastern part of the Russian craton. North and west of the Cis-Ural foredeep and the Precaspian depression (micro-ocean) carbonate platforms were formed on a shallow-marine shelf during the Asselian, Sakmarian and Early Artinskian. Reefs developed on the margin of these platforms along the slopes of the Cis-Ural foredeep and the Precaspian depression. The reefs shifted platform ward in the eastern areas, due to the tectonic subsidence of the platform margin and at the same time, prograded basinward in the south. Movements of continental blocks from the south during the Late Artinskian and Kungurian caused the separation of the Early Permian basin of the Russian craton from the Palaeo-Tethys, followed by evaporite sedimentation in the restricted basins. The existence of source rocks (bituminous deep-water sediments), thick reservoir rocks (limestones and dolostones), evaporitic seals and structural as well as stratigraphic traps are responsible for large productive gas and oil fields (e.g., Orenburg field), some of which are distinctly associated with reef carbonates.     

Facies and biota of Anisian to Carnian carbonate platforms in the Northern Calcareous Alps (Tyrol and Bavaria)

Summary During the Middle and early Late Triassic carbonate ramps and rimmed platforms developed at the northwestern margin of the Tethys ocean. In the Northern Calcareous Alps, Anisian stacked homoclinal ramps evolved through a transitional stage with distally steepened ramps to huge rimmed platforms of Late Ladinian to Early Carnian age. Middle Triassic to early Late Triassic facies and biota of basin, slope and platform depositional systems are described. Special emphasis is given to foraminifers, sponges, microproblematic organisms and algae. The Ladinian to early Carnian reef associations are characterized by the abundance of segmented sponges, microproblematica, biogenic crusts and synsedimentary cements. Among the foraminifers, recifal forms likeHydrania dulloi andCucurbita infundibuliformis (Carnian in age) are reported from the Northern Calcareous Alps for the first time. Some sphinctozoid sponges likeParavesicocaulis concentricus were known until now only from the Hungarian and Russian Triassic.     

Triassic in Iran

Summary Except of the Nakhlak and Aghdarband regions, Lower and Middle Triassic strata in Iran consist almost exclusively of carbonate rocks built on vast platforms along the shelves of the Paleo- and Neotethys. The depositional environments varied from shallow shelf sea to lagoonal and near-shore tidal flats, becoming even evaporitic towards the coastal regions of the Persian Gulf. During the early Late Triassic the formerly dominating platform carbonates were strongly reduced, being restricted for the rest of the Early Mesozoic to Southwest Iran which remained on the northern passive margin of Gondwana (later Arabian Plate). On the Iran Plate otherwise, as a result of the closure of the Paleotethys and the continent-continent collision with the southern active margin of Eurasia (Turan Plate), there was a change to thick siliciclastic and molassic sediments with only minor carbonate content durig the Upper Triassic and Lower Jurassic. Geographically restricted and completely different sequences of thick volcanoclastic and basinal Triassic sediments are known from Nakhlak in Central and from Aghdarband in Northeast Iran, both interpreted as remanents of the northern Paleotethys margin (Turan Plate).     

A Late Permian flora with Dicroidium from the Dead Sea region, Jordan

Three new species of Dicroidium, D. irnensis, D. jordanensis and D. robustum, are described from the Um Irna Formation (Upper Permian) of the Dead Sea region, Jordan. The plant remains are preserved as compressions with excellent cuticles. These are the earliest unequivocal records of Dicroidium, a genus that is typical for the Triassic of Gondwana. It is also the northernmost occurrence of this genus that apparently originated in the Permian in the palaeotropics. Middle and Late Permian floras from the Arabian Peninsula and adjacent regions show a remarkable mixture of elements from different floral provinces, i.e. Euramerica, Cathaysia and Gondwana. The climatic amelioration in the Early Triassic apparently enabled Dicroidium to migrate southward and eventually colonise the entire Gondwana region. Dicroidium is one of the very few megaplant genera not affected by the end-Permian biotic crisis, the largest Phanerozoic extinction event.     

Biogeography of the Late Triassic wallowaconchid megalodontoid bivalves

Discoveries of Late Triassic alatoform bivalves (family Wallowaconchidae) in Alaska and in south Asia and Arabia reveal a broad distribution for these tropical bivalves, overturning the concept of these bivalves being endemics of offshore island arc terranes in eastern Panthalassa. They occupied an area extending from central Tethys to the eastern Pacific margin of Panthalassa. The Tethyan wallowaconchids occur in nearshore environments on the north margin of Gondwana, indicating an environmental tolerance for continental shelf settings in addition to their primary occurrence on isolated oceanic islands. The new sites reveal a taxonomic diversity among wallowaconchids. Wallowaconchids present in India are conspecific with Wallowaconcha raylenea, the type species of Wallowaconcha, whereas wallowaconchids in Arabia are an undescribed taxon and wallowaconchids in south-central Alaska are small in size and a different undescribed taxon. All known wallowaconchids are of Norian age and they appear to be a wide-ranging component of Late Triassic tropical biotas.     

Tethyan uppermost Permian (Dzhulfian and Dorashamian) foraminiferal faunas and their paleogeographic and tectonic implications

The foraminiferal faunas and biostratigraphic correlation of the Tethyan uppermost Permian (Dzhulfian and Dorashamian) provide important paleogeographic and tectonic data for the interpretation of the Palaeofusulina-bearing terranes in East and Southeast Asia. These interpretations have a significant bearing on understanding Japanese pre-Cretaceous tectonostratigraphic and micropaleontologic data, as well as the geodynamic evolution of the Japanese Palaeofusulina-bearing terranes. The Tethyan foraminiferal fauna in the uppermost Permian is characterized by the occurrence of provincial and endemic schubertellid genera, and the absence of neoschwagerinids and verbeekinids which had characterized the rapidly evolving Middle Permian Tethyan marine faunas until their extinction at the end of the Midian. Difficulties in world-wide correlation of the uppermost Permian have resulted because of different geographic faunal compositions and the geographic patterns of extinction of Permian marine faunas. The Palaeofusulina fauna is one of the most reliable indicators of the uppermost Permian. Its presence or absence serves as paleogeographic constraints on East and Southeast Asian terranes. For example, the absence of Palaeofusulina fauna and the presence of late Midian Lepidolina multiseptata faunas in the Lhasa Terrane in Tibet and the Wolya Terrane in Sumatra (the third continental sliver north of Gondwana) are important, particularly, for identifying the rift–drift–collision process of the Gondwana-affinity terranes. They suggest a Late Permian separation of the two terranes from Gondwana. Tethyan Palaeofusulina occur in the latest Permian tropical to subtropical latitudinal belt and along with other geologic data assist in paleogeographic reconstructions and in interpreting the possible movement and emplacement of the Palaeofusulina-bearing terranes such as the Maizuru, South Kitakami–Kurosegawa and Chichibu terranes in Japan. They reveal that: (1) the Upper Permian Maizuru Group was deposited on the eastern continental margin of South China; (2) the occurrence of the Lower Permian Cathaysian flora and a number of geologic data in the South Kitakami–Kurosegawa suggest an arc–trench system setting and the Late Permian deposition in a shallow open-marine environment in proximity to South China; (3) foraminiferal biogeograpic data and the reconstructed oceanic plate stratigraphy in the Chichibu Terrane constrain the location of the Chichibu Seamount Chains to the western part of the Panthalassan domain, as they moved westwards against the Cathaysian Continent until their Jurassic accretion.     

Paleobiogeography of Mesozoic brachiopod faunas from Andean–Patagonian areas in a global context

During the Mesozoic, the Andean region has played a hinging role between high- and low-latitude faunas, which are, respectively, characterized by stocks that display long-term fidelity. This paper is aimed at providing an updated review of Late Triassic to Late Cretaceous South American articulated brachiopods in the light of previous knowledge at worldwide scale. Late Triassic brachiopods from the Argentine–Chilean Andes show unmistakable Maorian (or Notal) faunal elements alongside some more cosmopolitan genera, with certain influence of Eastern Pacific taxa. By Early Jurassic times, differentiation of Tethyan and Boreal Realms became progressively evident in Europe. In South America, Hettangian–Sinemurian brachiopod faunules from the Argentinian Andes are somewhat impoverished, with mostly cosmopolitan genera showing certain affinities to Maorian species, and with the addition of some endemics later. Increasingly, diverse Pliensbachian Andean brachiopods denote close relationships to Celto-Swabian taxa, then by Domerian times, a certain degree of endemism was developed, though somewhat delayed Tethyan influences, and persistent links with New Zealand are subordinately recognizable, too; most Toarcian assemblages reveal basically Celto-Swabian and Iberian affinities as well. East-west austral links across the Pacific may have been favored by migratory routes fringing the Gondwana margin, whereas faunal exchange with the western end of the Tethys appears to reflect an intermittent shallow-marine connection through the Hispanic Corridor. During the Middle Jurassic, distinction of Tethyan and Boreal Realms was maintained in the northern Hemisphere, and the differentiation of an Ethiopian or Southern Tethyan fauna became better characterized. Aalenian and Bajocian brachiopods of the Andes display generic affinities mainly with those from western Europe, with some minor endemic developments; brachiopods recorded from the Bathonian–Callovian of Argentina (and Chile) also occur along the northern Tethyan margin, yet with some genera extending into Indo-Ethiopian areas. During the Late Jurassic, Boreal faunas from high-latitudes became even more strongly differentiated from low-latitude, Tethyan ones. Oxfordian and Tithonian brachiopods from the Andes apparently belong to genera of cosmopolitan or northern Tethyan affiliation, yet there are few elements in common with other eastern Pacific areas, such as Mexico. Early Cretaceous brachiopods, in addition to Andean basins of Chile and western Argentina, are known also from Patagonia and Tierra del Fuego. They belong mostly to widely distributed, mainly Tethyan genera, with some quasi-cosmopolitan and circum-Pacific components (some shared with Antarctica become noticeable). Late Cretaceous brachiopods from northern Patagonia show significant affinities to Maastrichtian ones of northwest Europe and central Asia, which calls for further assessing the potential role that may have played the trans-Saharan passageway in such dispersal. Broad aspects of Mesozoic brachiopod paleobiogeography are fairly well understood, yet details of ranking and naming of certain units are still in need of more agreement.     

Environmental change in the Early Permian of NE Svalbard: from a warm-water carbonate platform (Gipshuken Formation) to a temperate,mixed siliciclastic-carbonate ramp (Kapp Starostin Formation)

A detailed facies study of Early Permian strata within NE Svalbard reveals a fundamental change of the depositional setting, from a restricted-marine, warm-water carbonate platform to an open-marine, temperate-water, mixed siliciclastic-carbonate ramp. The uppermost strata of the Gipshuken Formation (Templet and Sørfonna members; Sakmarian–early Artinskian?) consist of microbialites (algal mats), mudstones, bioclastic/peloidal limestones, carbonate breccias and Microcodium facies reflecting peritidal platform areas and supratidal sabkhas. A mixed heterozoan/reduced photozoan assemblage indicates temperate-water conditions within neighboring deeper, open-marine mid-platform areas, while warm-water conditions still prevailed within inner platform zones. In contrast, the lowermost strata of the overlying Kapp Starostin Formation (Vøringen Member; late Artinskian?–Kungurian) show a fully heterozoan biotic assemblage reflecting temperate water conditions within open-marine, storm-dominated, nearshore to transitional offshore areas of a mixed carbonate-siliciclastic ramp. The Vøringen Member comprises three facies associations, which form a shallowing-upward sequence subsequent to an initial transgression. The sediments reflect bryozoan bioherms in most distal areas, followed by stacked tempestites of sandy brachiopodal shell banks and Skolithos piperocks, grading into broad sand flats in most proximal areas of the inner ramp. The above environmental change is regarded as a regional event taken place across the entire shelf along the northern margin of Pangea and is attributed to paleoclimatic, paleoceanographic, as well as paleogeographic changes, possibly related to the overall northwards drift of the supercontinent. An abrupt increase in terrigenous input coinciding with this change is ascribed to the uplift of a new local source area, probably to the north or east of the investigation area.     

Episodic sedimentation on a peri-Tethyan ridge through the Middle–Late Jurassic transition (Villány Mountains, southern Hungary)

The Villány area, as a central part of the Tisza microcontinent/terrane along the European margin of Tethys, was characterized by intense subsidence in the Early and Middle Triassic, followed by a long interruption of subsidence in the Late Triassic to Middle Jurassic. During the Middle–Late Jurassic transition, marine sedimentation started with three distinct sedimentary episodes dated as Late Bathonian, Early Callovian, and Middle–Late Callovian, respectively. The succession is terminated by a thick limestone of Middle Oxfordian age. The sedimentary features, microfacies, and macroinvertebrate associations of these four stratigraphic units are documented and illustrated. The Middle to Late Jurassic sedimentary episodes of the Villány succession record an interplay of local and global factors and paleogeographical changes. At the beginning, local tectonic movements governed the main features of sedimentation, though the role of eustasy was also essential. From the mid-Callovian onwards, global climatic, biotic, and paleoceanographical changes controlled the nature and formation of the local carbonate sediments. The Callovian stromatolites are attributed to the activity of sulphate-reducing bacteria in a deep sublittoral, current-swept environment. Upwelling of eutrophic Tethyan waters is recorded by the prevalence of the Bositra filament microfacies in the Callovian. The long submarine hiatus at around the Callovian–Oxfordian transition mirrors a serious restriction of the carbonate budget, due to sudden cooling and a change in the oceanic current system (opening of a circumglobal Tethyan Passage), and to a higher amount of dissolved CO₂. In the Middle Oxfordian, the carbonate production considerably increased in accordance with the sudden global warming.     

Earliest Triassic microbialites in the South China block and other areas: controls on their growth and distribution

Earliest Triassic microbialites (ETMs) and inorganic carbonate crystal fans formed after the end-Permian mass extinction (ca. 251.4 Ma) within the basal Triassic Hindeodus parvus conodont zone. ETMs are distinguished from rarer, and more regional, subsequent Triassic microbialites. Large differences in ETMs between northern and southern areas of the South China block suggest geographic provinces, and ETMs are most abundant throughout the equatorial Tethys Ocean with further geographic variation. ETMs occur in shallow-marine shelves in a superanoxic stratified ocean and form the only widespread Phanerozoic microbialites with structures similar to those of the Cambro-Ordovician, and briefly after the latest Ordovician, Late Silurian and Late Devonian extinctions. ETMs disappeared long before the mid-Triassic biotic recovery, but it is not clear why, if they are interpreted as disaster taxa. In general, ETM occurrence suggests that microbially mediated calcification occurred where upwelled carbonate-rich anoxic waters mixed with warm aerated surface waters, forming regional dysoxia, so that extreme carbonate supersaturation and dysoxic conditions were both required for their growth. Long-term oceanic and atmospheric changes may have contributed to a trigger for ETM formation. In equatorial western Pangea, the earliest microbialites are late Early Triassic, but it is possible that ETMs could exist in western Pangea, if well-preserved earliest Triassic facies are discovered in future work.     

Late Paleozoic faunal, climatic, and geographic changes in the Baoshan block as a Gondwana-derived continental fragment in southwest China

The Carboniferous and Permian of the Baoshan block consist of three major depositional sequences: a Lower Carboniferous carbonate sequence, a Lower Permian siliciclastic sequence, and a Middle Permian carbonate sequence. These three sequences were interrupted by two major regressive events: first, the Namurian Uplift ranging in age from Serpukovian to Gzhelian, and second, the Post-Sakmarian Regression occurring probably at Artinskian time in the Baoshan block, although the precise time interval of the latter event is still unclear. The Baoshan block is characterized by warm-water, highly diverse and abundant faunas during the Early Carboniferous, by cold-water and low diversity faunas during the Early Permian, and by possibly warm-water but low diversity faunas during the Middle Permian. The Sweetognathus bucaramangus conodont fauna constrains the upper boundary of the diamictite-bearing siliciclastic deposits (Dingjiazhai Formation) to the Sakmarian to early Artinskian, as well as the eruption of the rifting basalts (Woniusi Formation) to, at least, the post-early Artinskian. Paleozoogeographically, affiliation of the faunas in the Baoshan block changed from Eurasian in the Early Carboniferous, to Peri-Gondwanan in the Early Permian, and to Marginal Cathaysian/Cimmerian in the Middle Permian. Cimmerian blocks have more or less comparable geohistory to one another in the Carboniferous and Permian. During the Middle Permian, the eastern Cimmerian blocks such as Sibumasu (s.s), Baoshan, and Tengchong are not far from the palaeoequator, but apparently more distant than the western Cimmerian blocks based on the presence or absence of some index taxa such as the fusulinaceans Eopolydiexodina and Neoschwagerina, and the corals Thomasiphyllum and Wentzellophyllum persicum.     

Early Permian brachiopod faunas from Peninsular India: Their Gondwanan relationships

The Early Permian (Late Asselian? to Aktastinian?) brachiopod faunas of Peninsular India are revised in terms of current taxonomy. Genera such as Semilingula, Arctitreta, Etherilosia, Strophalosia, Aulosteges, Bandoproductus, Cyrtella, Neospirifer, Crassispirifer, Tomiopsis and Gilledia confirm the Gondwanan aspect of the faunas and a close relationship, at the generic level, to the Early Permian brachiopod faunas of Western Australia. Peninsular Indian Early Permian brachiopod faunas belong to a complex of Gondwanan and peri‐Gondwanan faunas from Oman and the Pamirs in the West to Australasia in the east. This distribution implies relative freedom of migration for the faunas along the northern margin of Gondwana during the Early Permian.     

Late paleozoic and Late Triassic limestones from north Palawan Block (Philippines): Microfacies and paleogeographical implications

Summary Selected Late Paleozoic and Triassic limestone exposures were studied on northern Palawan Island, Philippines, with regard to microfacies, stratigraphy and facies interpretation. Although some of the outcrops were already reported in literature, we present the first detailed microfacies study. Late Paleozoic carbonates in the El Nido area are represented by widley distributed Permian and locally very restricted Carbonifenous limestones. Of particular interest is the first report of Carboniferous limestones in the Philippines dated by fossils. Fusulinids indicate a ‘Middle’ Carboniferous (Moscovian-Kasimovian) age of the Paglugaban Formation only known from Paglugaban Island. The Permian Minilog Formation consists mostly of fusulinid wackestones and dasycladacean wacke-/packstones. Fusulinid datings (neoschwagerinids and verbeekinids) provide a Guadalupian (Wordian-Capitanian) age. The depositional setting of the Middle Permian carbonates corresponds to a distally steepened ramp with biostromes built by alatoconchid bivalves locally associated with richthofeniid brachiopods. Late Triassic limestones occur in isolated exposures on and around Busuanga Island (Calamian Islands). The age of the investigated carbonates is Rhaetian based on the occurrence ofTriasina hantkent Maizon. Microfacies data indicate the existence of reefs (Malajon Island) and carbonate platforms (Kalampisanan Islands, Busuanga Island, Coron Island). Reef boundstones are characterized by abundant solenoporacean red algae, coralline sponges and corals. Platform carbonates yield a broad spectrum of microfacies types, predominantly wacke- and packstones with abundant involutinid foraminifera and some calcareous algae. These facies types correspond to platform carbonates known from other parts of Southeast Asia (Eastern Sulawesi and Banda Basin; Malay Peninsula and Malay Basin). The Philippine platform carbonates were deposited on and around seamounts surrounded by deeper water radiolarian cherts. The new data on facies and age of the Philippine Permian and Triassic carbonates contradict a close paleogeographical connection between the North Palawan Block and South China and arise problems for the currently proposed origin of the North Palawan Block at the paleomargin of South China. We hypothesize that North Palawan was part of the Indochina Block during the Carboniferous and Permian, separated from the Indochina Block during the Middle Permian and collided with the South China Block in the Late Cretaceous.