Palaeoenvironment and palaeoclimate in the western Liguria region (northwestern Italy) during the Last Glacial. The small mammal sequence of Riparo Mochi (Balzi Rossi,Ventimiglia)

Riparo Mochi is considered a key site for the Middle–Upper Palaeolithic chrono-cultural sequence in the Mediterranean area. Climatic and environmental conditions have been reconstructed after the study of the small mammal assemblage by means of linear regression method and the Habitat Weighting method. The climate proxies for the late Mousterian and Proto-Aurignacian (cultural units I to G) indicate an environment with a relative high percentage of forest component during two cold and one relative warm oscillations. Open environments with low temperatures are registered for the Gravettian (cultural Unit D) allowing a possible relation to the Heinrich 3 Event. The final Gravettian (cultural Unit C) shows a relative warm peak, tentatively related to Greenland Interstadial 2. The late Epigravettian (cultural Unit A) is characterized by a high percentage of forest-related environment suggesting that this horizon formed during the Bølling–Allerød Interstadial. Finally, the Riparo Mochi sequence contributes to reconstruct the processes of faunal changes along the northern Tyrrhenian coast in a period of climatic instability, especially during the end of Marine Isotope Stage 3 when foraging groups adopted different mobility strategies between southern France and the Italian peninsula.     

The fossil bat assemblages from the Grotta dei Pipistrelli in Pantalica (southeastern Sicily,Italy): Chronological and palaeoecological implications

Bat remains are useful for palaeoecological reconstructions, they provide independent information on palaeoenvironment and are good indicators for hypogean microclimates. Nine taxa of Chiroptera divided into three families (Rhinolophidae, Vespertilionidae and Miniopteridae) and four genera (Rhinolophus, Myotis, Plecotus and Miniopterus) were discovered in three fossil assemblages from the Grotta dei Pipistrelli, in Sicily, a key region for an understanding of Quaternary climates and environments. Bat remains were deposited during three distinct timespans: one late Pleistocene, MIS 2, around the Last Glacial Maximum, and two Holocene, both referable to the Atlantic chronozone. The presence of yearlings, adult and old individuals suggests hibernating colonies, but the cave was also utilized as a nursery. The diversity of taxa indicates a composite landscape with prevailing vegetation cover and relatively warm climatic conditions. The percentage variations of the frequencies of the recognized taxa suggest a progressive increase of open spaces in the middle Holocene.     

First occurrence of the extinct deer Haploidoceros in the Iberian Peninsula in the Upper Pleistocene of the Cova del Rinoceront (Castelldefels,Barcelona)

The Cova del Rinoceront is the first site in the Iberian Peninsula where the genus Haploidoceros has been documented. This discovery of abundant remains is also its first recorded occurrence in Europe during the Upper Pleistocene. The new fossil record strengthens claims that this genus was widely distributed during the Pleistocene, occupying a longer time span at least until MIS 5. Cranial and post-cranial skeletal remains ascribed to this species were recovered from the uppermost layers of the Cova del Rinoceront. The diagnostic features that allow these remains to be unequivocally assigned to H. mediterraneus, include their cranial morphology and antler shape, comprising two sickle-shaped beams, curving backwards and laterally, and a long frontal basal tine. This paper highlights the problems of deer systematics and identification given the high degree of fragmentation of the genus, their relatively homogeneous morphology, a virtual absence of reliable characters and the overlap of measurements between species.     

The open-air Paleolithic site of Mirak,northern edge of the Iranian Central Desert (Semnan,Iran): Evidence of repeated human occupations during the late Pleistocene

The northern edge of the Iranian Central Desert has provided valuable evidence of terminal Pleistocene human settlements. Mirak constitutes one of the largest open-air lithic scatters in the region, consisting of eight natural mounds. Fieldwork was initiated in 2015 by the joint Iranian-French program at Mirak 8. Preliminary results have demonstrated at least three successive phases of human occupation during the MIS3: an upper layer with clear Upper Paleolithic affinities and a maximum age of 28 ky, a lower layer with clear Middle Paleolithic affinities that dates around 47 ky, and an intermediate layer with mixed characteristics that can be seen as an intermediate Paleolithic phase which dates between 28 ± 2 and 38 ± 2 ky. At the time when Upper Paleolithic cultures originated in the Zagros Mountains, cultures with clear Middle Paleolithic affinities persisted nearby along the northern edge of the Iranian Central Plateau.     

Feeding ecology of Late Pleistocene Muntiacus muntjak in the Padang Highlands (Sumatra)

The Dubois Collectie comprises around 1000 isolated fossil teeth of Muntiacus muntjak from the Padang Highlands in Sumatra. The majority were retrieved from one of the three fossil cave sites Lida Ajer, Sibrambang or Jambu. Lida Ajer is the only cave, for which exact location, geology as well as the age of the fossiliferous deposits are known. Using the mesowear method, we searched for differences in the dietary signal of Muntiacus muntjak and thus in the paleoenvironments of the three caves. Muntiacus muntjak from either one of the caves does not differ significantly in the composition of its diet, showing a mixed feeder signal. The samples of Lida Ajer and Sibrambang illustrate an increase of the mixed feeding component, which could be the result of higher seasonality.     

The fossil bat assemblage of Sima del Elefante Lower Red Unit (Atapuerca,Spain): First results and contribution to the palaeoenvironmental approach to the site

The fossil bat assemblage from the Lower Red Unit of Sima del Elefante (TELRU) in Atapuerca (Burgos, Spain) has been exhaustively analysed for the first time. Bat fossil assemblages are of particular relevance to palaeoenvironmental approaches to sites. Here we integrate our new data on the chiropteran fauna with the data provided previously by other authors on the basis of the small-vertebrate assemblages and the palynology of the site. Our results are consistent with the earlier results in that they indicate a generally warmer climate than at present in the area and stable environmental conditions throughout the major part of the TELRU sequence. However, fossil bat assemblages sometimes lead to problems when they are used in landscape reconstruction. These problems are discussed here and should be taken into account in future works.     

A new Late Pleistocene non-anthropogenic vertebrate assemblage from the northern Iberian Peninsula: Artazu VII (Arrasate,Basque Country)

Late Pleistocene palaeontological sites without human intervention are limited in the Cantabrian region, and even more so those with a good state of preservation and rich biodiversity. A new vertebrate fossiliferous locality discovered at Kobate Quarry (Arrasate, northern Iberian Peninsula) is presented in this article. This site, in which remains of 40 different vertebrate taxa were accumulated, acted as a natural trap. The preliminary vertebrate faunal list includes five amphibian taxa, four reptiles, seven species of birds and 24 mammalian taxa. While small mammals are represented by 13 small mammal taxa (seven in the Order Rodentia, five in the Order Eulipotyphla, and one in the Order Chiroptera), the large mammal fauna comprises eleven species, including ungulates and carnivores. The palaeoecology inferred from this faunal assemblage suggests the existence of large forested areas with some grassland and a watercourse nearby, within a notably warm and humid climate. These palaeoenvironmental conditions, combined with AMS and AAR results carried out in macrofaunal bone samples, suggest that the deposit from Artazu VII would be located in the first half of the Late Pleistocene, in the Marine Isotope Stage (MIS) 5c.     

A new hawker dragonfly from the Middle Jurassic of China (Odonata: Aeshnoptera)

A new genus and species Linqibinia panae of paracymatophlebiid hawker dragonfly is described from the Middle Jurassic Haifanggou Formation (Inner Mongolia, China). Previously only known from Karatau in Kazakhstan, the discovery of another member of this family extends its range across Central Asia. It confirms that the Aeshnoptera was among the most diverse odonatan clades during the Middle-Late Jurassic.     

The Vallonnet cave on the northern Mediterranean border: A record of one of the oldest human presences in Europe

Prehistoric sites testifying to human presence older than one million years in Europe are rare, and in the current state of knowledge, the oldest of them have been dated to around 1.4–1.5 Ma. The Vallonnet cave at Roquebrune-Cap-Martin in the Alpes-Maritimes, on the Mediterranean border, is one of the oldest sites in France to have yielded evidence of human activity: a lithic assemblage of about a hundred pieces and traces of butchery on bones of an Epivillafranchian fauna. The archaeological levels of this small cave were recently dated between 1.1 and 1.2 Ma by U-Pb correlated with paleomagnetic data. The site was occupied alternately by large carnivores that used it as a den or a lair, and by hominins that stayed there briefly in bivouac. The lithic remains are mainly percussion tools, shaped pebbles, flakes and cores, whose raw materials are local, or even semi-local, and on the whole not very diversified with mainly limestone, and to a lesser extent sandstone, quartzite, flint and quartz. This assemblage is attributed to a Mode 1 technology (Oldowayen), among which macro-tools (hammerstones, shaped and fractured pebbles) are found alongside rarer elements resulting from debitage operating chains aimed at producing sharp-edged flakes, very rarely retouched. The bipolar-on-anvil flaking technique could be identified from the characteristics of some artifacts. Several refitting flakes on shaped pebbles or percussion tools attest to knapping and percussion activities in the cave. Hominins consumed the remains of large herbivore carcasses, as attested by the presence of cutting and fracturing marks on some bones. The presence of a freshwater source in the immediate vicinity, and the knapping and butchering activities here therefore document the subsistence behavior of Lower Pleistocene human groups, certainly in competition with the carnivores present.