New and validated syntaxa for the checklist of Italian vegetation

We drew up a checklist of the Italian vegetation (http://www.prodromo-vegetazione-italia.org/), up to the syntaxonomical rank of alliance. During the compilation of this checklist, we observed that some syntaxa were invalidly published. For this reason, in this article we validated some syntaxa names and, at the same time, described new syntaxa of different hierarchical levels. Therefore, 10 new orders, 1 new suborder, 18 new alliances, 3 new suballiances and 5 new associations are described here. These new syntaxa belong to the following classes: Adiantetea capilli-veneris, Parietarietea judaicae, Thlaspietea rotundifolii, Artemisietea vulgaris, Stellarietea mediae, Galio aparines–Urticetea dioicae, Mulgedio alpini–Aconitetea variegati, Trifolio medii–Geranietea sanguinei, Festuco-Seslerietea, Salicetea herbaceae, Festuco valesiacae–Brometea erecti, Molinio-Arrhenatheretea, Cisto cretici–Micromerietea julianae, Rhamno catharticae–Prunetea spinosae, Salici purpureae–Populetea nigrae, Salicetea purpureae, Quercetea ilicis and Querco roboris–Fagetea sylvaticae.     

Halocnemum M. Bieb. vegetation in the Mediterranean Basin

The aim of the article is the syntaxonomic interpretation of hyperhalophilous woody or semi-woody vegetation with Halocnemum M. Bieb. along the coasts of the Mediterranean Basin. For this area, the two species of Halocnemum, H. strobilaceum (Pall.) M. Bieb. and H. cruciatum (Forssk.) Tod. are identified; their morphological characteristics, synonymy and distribution are here described. The study carried out of particular aspects of the plant morphology and the micromorphological characteristics of the seeds collected from different places in the Mediterranean allows to better differentiate the two species. Vegetation studies already done in many communities of the Mediterranean by several authors threw into great confusion because only H. strobilacem has been recognized as a dominant species. As regards the syntaxonomic analyses of the vegetation, the authors refer to the proposals already made by various scientists for the vegetation of inland salt basins of Eurasia and the Irano-Anatolian area, who suggested the classes Kalidietea foliate and Halocnemetea strobilaceiirano-anatolica, respectively. Conversely, in the Mediterranean Basin, the vegetation study is present only in coastal areas where it is considerably impoverished in the number of species. Therefore, the authors propose to include hyperhalophilous, woody and fruticose vegetation in the class Sarcocornietea fruticosae. However, the phytosociological and ecological diversity is highlighted proposing the order Halocnemetalia cruciati in which both the alliance Halocnemion strobilacei, for middle Eastern Europe, and the alliance Halocnemion cruciati, for North Africa with penetrations in the Western and Eastern Europe up to the Middle East coast, are included. This new interpretation has required the correction of the names of two associations (Frankenio corymbosae–Halocnemetum cruciati and Zygophylloalbi–Halocnemetumcruciati) and the proposal of two new associations Arthrocnemo machrostachyi–Halocnemetum cruciati and Halocnemo cruciati–Sarcocornietum fruticosae. A further proposal concerns the addition of the alliance Limoniastrion monopetali, previously included in the order Limonietalia, in the order Halocnemetalia cruciati.     

Phytosociology of Juniperus excelsa M.Bieb. forests in Alborz mountain range in the north of Iran

Juniperus excelsa is one of the specific tree species in the mountainous areas of Iran. It forms the mountainous forests of Irano-Turanian vegetation region. Despite the considerable spread of J. excelsa in Iran, there is no detailed and valid published study on the phytosociology of Iranian juniper stands. The aim of this research is to identify these associations and their characteristics in Alborz mountains in the north of Iran. To this goal, a set of data including floristic data according to Braun-Blanquet method, and important environmental characteristics, were collected. To classify the vegetation, TWINSPAN was used and phytosociology tables were analysed with JUICE software. Diagnostic species were determined by measuring the degree of fidelity and using statistically meaningful measurements. According to the results, the vegetation was classified into six associations and five subassociations. Among them are Cotoneastro nummulariis-Juniperetum excelsae ass. nov., Lonicero ibericae-Juniperetum excelsae ass. nov., Rhamno pallasii-Juniperetum excelsae ass. nov., Rhamno p.-Juniperetum e. ajugetosum chamaecisti subass. nov., Rhamno p.-Juniperetum excelsae gypsophiletosum aretioidis subass. nov., Amygdalo lycioidis-Pistacietum atlanticae hymenocratretosum bituminosoris Kashipazha et al. 2007, Feruletum ovinae ass. nov., Artemisio aucheri-Astragaletum veri Asri 2004, and Artemisio aucheri-Onobrychidetum cornutae ass. nov. The introduced juniper associations were placed in class Junipero-Pistacietea Zohary 1973 and order Juniperetalia excelsae nov.     

Phytosociological survey of Northern California dunes

ABSTRACT

The vegetation of sandy coasts of California between Point Arena and Morro Bay, was investigated. Near the tide line, was annual vegetation of the Cakile maritima community. Dunal vegetation included the new Abronio latifoliae-Ambrosietea chamissonis class with the new Ambrosietalia chamissonis order and the new Ambrosion chamissonis alliance (proposed). Foredune vegetation included the two new Abronio latifoliae-Elymetum mollis and Atriplici leucophyllae-Abronietum maritimae associations, with the new Polygono paronychiae-Artemisietum pycnocephalae and Erigero glauci-Abronietum latifoliae associations on more stabilised surfaces. In some localities, Ammophila arenaria var. australis occurs within a Lupino variicoloris-Ammophiletum australis association, which colonises the whole dune ecosystem. In the flat areas characterised by grit and affected by frequent sea ingression is an association characterised by Juncus lesueurii and Distichlis spicata (Junco lesueurii-Distichletum spicatae). In the hollows among inland dunes, affected by stagnation, thick meadows develop which contain two new associations: Stachydo rigidae-Iridetum douglasianae developing on flat areas with a stronger stagnation and Achilleo millefoliae-Elymetum pacifici.     

Synanthropic vegetation of the <Emphasis Type="Italic">Eragrostion cilianensi-minoris</Emphasis> alliance in the Czech Republic

A synthesis of the alliance Eragrostion cilianensi-minoris in the Czech Republic is presented on the basis of 82 relevés including new unpublished data. A TWINSPAN classification and detrended correspondence analysis were used to identify the main vegetation types included in the alliance Eragrostion cilianensi-minoris. A syntaxonomic revision of the data set revealed five associations of the alliance: Digitario sanguinalis-Eragrostietum minoris, Portulacetum oleraceae, Eragrostio poaeoidis-Panicetum capillaris, Cynodontetum dactyli, and Hibisco trioni-Eragrostietum poaeoidis. The latter was recently found in several arable fields in Southern Moravia (Czech Republic) and was newly characterized.     

Syntaxonomy of the Robinia pseudoacacia communities in the central peri-adriatic sector of the Italian peninsula

This study presents the results of a floristic vegetational study on Robinia pseudoacacia neoformation forests in the peri-Adriatic sector of central Italy. This has allowed the characterization of these coenoses at the ecological, biogeographic, syntaxonomic and landscape levels. These currently represent the southernmost syntaxa of the Robinietea class described for the Italian peninsula, and the first syntaxonomic contribution of this class in Europe for the Mediterranean biogeographical region. We propose here the new alliance Lauro nobilis–Robinion pseudoacaciae of the order Chelidonio–Robinietalia pseudoacaciae and class Robinietea, with two new associations: Melisso altissimae–Robinietum pseudoacaciae and Rubio peregrinae–Robinietum pseudoacaciae. The new alliance Lauro nobilis–Robinion pseudoacaciae (typus: Melisso altissimae–Robinietum pseudoacaciae) brings together neoformation forests and pre-forest dominated by R. pseudoacacia in those territories with a Mediterranean macroclimate of the peri-Adriatic sector of central Italy. The optimum is found for the alluvial plain and low-slope morphologies, on soils that are moist and rich in organic matter and in areas with anthropic disturbance. On the basis of comparisons with the European context, the alliance Bryonio–Robinion described for the temperate territories of northern Italy is here validated.     

Major plant communities of the Carson Desert (Nevada), North America's coldest and driest desert

This paper describes the vegetation of the Carson Desert (Nevada, USA) based on a phytosociological analysis of its major plant associations, as determined by the Braun-Blanquet method. Diagnostic tables, climatic, edaphic, and biogeographical data were used to establish floristic affinities among identified plant communities and to interpret their distributions in zonal gradients. Their syntaxonomic positions in the classes Allenrolfeetea occidentalis and Artemisio tridentatae–Juniperetea osteospermae, and in the new class Sarcobatetea vermiculati were also established. Three associations (Atriplici confertifoliae–Sarcobatetum baileyi, Atriplici canescentis–Psorothamnetum polydenii, and Suaedo moquinii–Sarcobatetum vermiculati) are described here for the first time along with the order Sarcobatetalia vermiculati and the alliance Sarcobation vermiculati.     

Rumex aquaticus L. at Loch Lomondside

Summary

A new alliance of spring vegetation dominated by the leafy liverwort Anthelia julacea is described from the montane regions of Britain. It is distinguished floristically and ecologically from similar alpine late snow bed communities in Scotland and Scandinavia. The classificatory position of the stands of this vegetation type is discussed with reference to continental and British systems and it is concluded that the erection of a new alliance is justified on floristic grounds.     

Vegetation of European springs: High-rank syntaxa of the Montio-Cardaminetea

Abstract. A survey of high-rank syntaxa of vegetation of European springs (class Montio-Cardaminetea) is presented. It shows the complexity of the class and the relations between the units. More than 1200 relevés were arranged into six phytosociological tables, each representing an alliance. A synoptic table comprising all alliances is presented. Bryo-phytes are the most important group in most of the communities, both regarding biomass and cover. The Adiantetea and the Montio-Cardaminetea are combined into one class. As to floristics and ecology, there are no obvious reasons to distinguish more than one order in this class. The vegetation of European springs is divided into six alliances: (1) Adiantion, split into two new suballiances, Adiantenion and Lycopodo-Cratoneurenion; (2) Cratoneurion commutati; (3) Cardamino-Montion - incl. three new suballiances: Mniobryo-Epilobienion hornemannii, Swertio-Anisothecienion and Cardamino-Montenion; (4) Dermatocarpion; (5) Epilobio nutantis-Montion; (6) Caricion remotae - incl. two new suballiances: Caricenion remotae and Cratoneuro filicini-Calthenion laetae. The alliances are floristically and ecologically characterized and their distributions in Europe shown on maps.     

Prodrome of the Italian vegetation: A new alliance for the high-mountain chamaephytic communities of central and southern Apennines

The new alliance Cerastio tomentosi-Globularion meridionalis and the new association Helianthemo alpestris-Globularietum meridionalis are here proposed as a contribute to the prodrome of Italian vegetation. The new syntaxa are related to the chamaephytic high-mountain vegetation and they are characterized by a wide presence of endemics, southern European orophytes, mountain-Mediterranean, and amphi-Adriatic species. The proposal aims to differentiate, on a structural basis, the high-mountain vegetation of the Apennines at higher syntaxonomical levels.     

Syntaxonomical study of Italian wall vegetation

A syntaxonomical study of Italian wall vegetation based on classification and ordination of a large phytosociological data set from various authors (using the IAHOPA program) has confirmed the alliances Centrantho-Parietarion and Galio-Parietarion (order Parietarietalia judaicae, class Parietarietea judaicae), as already proposed by traditional syntaxonomy. The variation in humidity and the north-south geographical gradient are shown to be the most significant factors for the differentiation of this vegetation in Italy. The presence of some associations belonging to the class Asplenietea trichomanis (alliance Cystopteridion, Potentillion caulescentis and Centaureo-Campanulion) shows a strong connection with natural rock vegetation.     

The Chrysopogono grylli-Nerietum oleandri association in Croatia as compared with other Rubo ulmifolii-Nerion oleandri communities (Tamaricetalia,Nerio-Tamaricetea) in the Mediterranean

Abstract

A new association, Chrysopogono grylli-Nerietum oleandri, situated in the south of Croatia, is proposed. According to its floristic structure, physiognomy and ecology, it belongs to the Rubo ulmifolii-Nerion oleandri alliance (Tamaricetalia, Nerio-Tamaricetea). It is the only association with Nerium oleander in Croatia. This community has developed along a temporary watercourse as a vegetation exclave (extrazonal stand) within the climazonal evergreen forest vegetation of the Oleo-Ceratonion alliance. The association has a thermophilous character and a large number of companions from the Quercetalia ilicis and Scorzonero-Chrysopogonetalia communities. This report compares the floristic composition of this association with others of the Rubo ulmifolii-Nerion oleandri alliance from other Mediterranean areas. The Croatian association is most similar to the Spartio-Nerietum oleandri association from the Ionian side of Calabria (south Italy). The syntaxonomic status of the Montenegrin association Andropogono distachyi-Nerietum oleandri was determined, and the status of the stands with Nerium oleander is discussed.     

Vegetation diversity of salt‐rich grasslands in Southeast Europe

Question

How does the plant species composition of Pontic–Pannonian salt‐rich habitats vary on a large geographical scale? Do the floristic differences between Pannonia and the Balkans correspond to the current phytosociological classification?

Location

Pannonia (Hungary, Slovakia, Austria, Czech Republic, Croatia, Serbia, Romania) and the Balkans (Bulgaria, Macedonia, Greece).

Methods

Two thousand four hundred and thirty‐seven relevés from halophytic and sub‐halophytic habitats were classified using a modified TWINSPAN. The crispness of classification was checked. DCA and CCA with climate data as explanatory variables were applied.

Results

The classification was best interpreted at the level of 15 clusters. The vegetation changed along the salinity gradient from sub‐halophytic grasslands (including Trifolion resupinati alliance of the Molinio‐Arrhenatheretalia class and Beckmannion eruciformis and Festucion pseudovinae p. p. alliances of the Festuco‐Puccinellietea class) and reed beds (Bolboschoenion maritimi p. p. alliance; the Phragmito‐Magnocaricetea class), through steppe and wet inland halophytic vegetation (Festucion pseudovinae p. p., Puccinellion limosae, Pucinellion convolutae, Bolboschoenion maritimi p. p. and Juncion gerardii of the Festuco‐Puccinellietea class) towards the extreme halophytic vegetation of the Thero‐Salicornietea, Crypsietea and Juncetea maritimi classes. This gradient was longer in the Balkan region, where it spanned from the sub‐mediterranean salt‐rich grasslands to the extremely halophytic vegetation at the Black Sea coast. The second most important gradient coincided with the water regime. Some vegetation types appeared to be confined to either the Pannonian or the Balkan region (especially within dry sub‐halophytic and steppe halophytic grasslands), while others were distributed across the entire study area. The above‐mentioned pattern did not always correspond with current classification systems.

Conclusions

Variation in salt‐rich vegetation predominantly follows the salinity and water regime gradients. Geographical variation, generally coinciding with climatic and historical effects, is also important, especially in drier salt‐rich habitats. Our large‐scale analysis of the floristic variation of salt‐rich habitats might be useful for the unification of classification systems that differ substantially between the countries involved. In addition, the analysis may be useful for adjustment of a classification system in the poorly explored Balkan region, where particular vegetation types were identified with, or delimited from, Central European vegetation types without detailed comparative analysis until now.

     

Variability of the <Emphasis Type="Italic">Molinion</Emphasis> meadows in Slovakia

Nine vegetation types were distinguished using cluster analysis within Molinion meadows in Slovakia. Vegetation of cluster 1 occurs on most acidic soils and is characterized by the occurrence of species of the Caricion fuscae alliance and of the Nardus grasslands. Vegetation of cluster 2 is also found on rather acidic soils but in contrast to cluster 1 vegetation it contains species of base-rich sites, such as Betonica officinalis, Galium boreale or Serratula tinctoria. Vegetation of cluster 3 occurs in wet base-rich habitats and often contains species of the Caricion davallianae alliance. Species of dry and Nardus grasslands are typical for vegetation of cluster 4, which is found at the driest sites and is confined to oligotrophic sandy soils. Vegetation of clusters 5 and 6 occurs on moist mesotrophic soils. Their species composition is quite similar, the main difference being that the former includes species-poor relevés and the latter includes species-rich relevés. Relevés of cluster 7 include species of dry grasslands and some ruderal species and represent degraded types of inundated floodplain meadows of the Deschampsion alliance. Vegetation of clusters 8 is characterized by species of the Phragmito-Magnocaricetea class and of the Deschampsion alliance, and occurs in wet nutrient-rich habitats. Vegetation of cluster 9, which usually develops from vegetation of cluster 8 due to decrease in the ground-water table, often contains species of dry grasslands and mesic meadows. Except for relevés of clusters 1 and 7, all others can be assigned to the Molinietum caeruleae Koch 1926 association. Cluster 1 corresponds to the Junco effusi-Molinietum caeruleae Tüxen 1954 association. Average Ellenberg indicator values for relevés, which were passively projected on the ordination biplot of detrended correspondence analysis, showed that the first ordination axis correlates with nutrients, soil base status and temperature, and second axis with moisture.     

The coastal plant communities of Juniperus macrocarpa in the Mediterranean region

Juniperus macrocarpa is a threatened taxon growing on coastal areas of the Mediterranean region. A remarkable part of its communities are included in Juniperion turbinatae, considered as priority habitats for conservation in the Habitats Directive 92/43 EEC. We have summarized the current status of this type of vegetation, including the described associations of this alliance in order to analyse the variability of the syntaxa, based on its floristic composition and geographic distribution. The available relevés were arranged in a synthetic table and subjected to a detendred correspondence analysis. Two new suballiances (Juniperenion turbinatae and Asparago orientalis-Juniperenion macrocarpae) have been proposed. A typological classification of these communities with ecological and biogeographical diagnoses is given.     

The Quercus cerris woods of the alliance Carpinion orientalis Horvat 1958 in Italy

Abstract

A phytosociological study of the alliance Carpinion orientalis in the hilly and subcoastal areas of the Italian peninsula is presented. These woods are included in the suballiances Laburno-Ostryenion and Lauro-Quercenion; the distinction between these two suballiances is not always easy also because of the conservation status of these forest ecosystems. The hilly and subcoastal Turkey oak woods of the alliance Carpinion orientalis are mainly distributed on the Adriatic side of the Italian peninsula, where they are found on neutro-basic substrata to which these coenoses are linked. The Turkey oak woods of the alliance Teucrio siculi-Quercion cerridis are strongly linked to the acidic substrata typical of the Tyrrhenian side, but are sporadically found on the Adriatic side within small enclaves with more acidic substrata. A numerical analysis allowed us to rearrange the lower hierarchical levels of the associations of Turkey oak woods of the alliance Carpinion orientalis. In particular, the association Lonicero xylostei-Quercetum cerridis is included in the alliance Carpinion orientalis, and the new subassociation festucetosum exaltatae is proposed. The subassociation rosetosum arvensis is proposed for the association Daphno laureolae-Quercetum cerridis, and the new subassociation cytisophylletosum sessilifolii of the association Aceri obtusati-Quercetum cerridis is described.     

Vegetation of the dry bioclimatic areas in the Dominican Republic

We conduct a study of the dry forest in areas of scant rainfall in the Dominican Republic; these are sites with particular endemic habitats, as the samples reveal a high rate of local and insular endemic species, with 84 (24.85%) endemisms. This work covers dry forests rich in coarse spiny plants and includes a statistical (multivariate analysis), phytosociological and catenal study of the vegetation. We analyse the floristic composition, ecology, distribution and richness in endemisms of each association. The bioclimatic analysis reveals the different thermotypes and ombrotypes on the island, and locates the proposed plant associations in the infra- and thermotropical thermotypes, and in the arid, semiarid and dry ombrotypes. As a result of this phytosociological analysis, we propose the new alliance Harrio nashii–Acacion skleroxylae and four plant associations: Harrisio nashii–Prosopidetum juliflorae, Crotono poitaei–Erythroxyletum rotundifolii, Lonchocarpo pycnophylli–Cylindropuntietum caribaeae and Neoabbottio paniculatae–Guaiacetum officinali; these associations connect catenally with the subhumid forest and mangrove swamps.     

Contribution to the knowledge of Bulgarian serpentine grasslands and their relationships with Balkan serpentine syntaxa

Serpentine areas, including those in Bulgaria, are rich in endemic taxa, and still remain to be investigated phytocoenologically. We analyse the vegetation types in various sites and compare them with those in other Balkan countries. The main objectives were (1) to explore and describe the relationships between the vegetation in the serpentine areas investigated in Bulgaria with those in the Balkan Peninsula and (2) to explore and classify the diversity of vegetation in grasslands on serpentine rocks in eastern Rhodope, Bulgaria. The classic methodology of the Braun-Blanquet school was applied to the exploration of the vegetation. Average linkage method (unweighted pair group method with arithmetic mean) and principal coordinate analysis were used to evaluate floristic and synoptic similarities. As a result, the new endemic association Onosmo pavlovae-Festucetum dalmaticae was proposed. This association can be included in the alliance Alyssion heldreichii Bergmeier et al. 2009, newly described on serpentine rocks in northern Greece. Our data confirmed the existence of similar or vicariant endemic syntaxa (associations) on isolated serpentine terrains in northern Greece and south-eastern Bulgaria.