Phu Din Daeng,a new Early Cretaceous vertebrate locality on the Khorat Plateau,NE Thailand

In this paper, we report on a new Early Cretaceous vertebrate locality, Phu Din Daeng, in Nakhon Phanom Province, NE Thailand. The Phu Din Daeng site has yielded a diverse vertebrate assemblage, including sharks (Heteroptychodus steinmanni), bony fishes (Pycnodontiformes; Sinamiidae cf. Siamamia and ?Vidalamiinae, and Ginglymodi), adocid turtles, indeterminate neosuchian crocodiles, pterosaurs and dinosaurs (spinosaurids and indeterminate theropods). A new adocid turtle, Protoshachemys rubra n. g. n. sp. is described on the basis of shell material. Field investigations on the geology and comparisons with other vertebrate faunas place Phu Din Daeng in the Sao Khua Formation (Barremian) of the Khorat Group.     

内蒙古查布地区下白垩统巨齿龙足印化石

内蒙古鄂尔多斯市鄂托克旗查布地区下白垩统中产有大量的蜥脚类和兽脚类恐龙足迹及鸟类足迹化石,作者对这个地区的足迹化石进行全面普查,发现许多新暴露出来的恐龙足迹化石,包括兽脚类恐龙足迹一新属新种Chapuslockleyiichnogen.etichnosp.nov.属于巨齿龙类(megalosaurid)所留。与兽脚类足迹保存在一起的蜥脚类足迹Brontopodusbirdi分布凌乱,反映了造迹恐龙搏斗的场面。通过测量连续的兽脚类足迹计算,Chapuslockleyi的造迹恐龙的行走速度为4.33km/h。通过足迹化石还识别出造迹恐龙的病态特征。     

The History of Dinosaur Footprint Discoveries in Wyoming with Emphasis on the Bighorn Basin

Dinosaur traces are well known from the western United States in the Colorado Plateau region (Utah, Colorado, New Mexico, and Arizona). Utah contains the greatest abundance of known and documented dinosaur footprints and trackways. Far less well known, however, is the occurrence and distribution of dinosaur footprint-bearing horizons in Wyoming. Scientific studies over the past 10 years have shown that three of the four Middle and Upper Jurassic formations in northern Wyoming contain dinosaur footprints. Two of the footprint-bearing horizons are located in geologic intervals that were once thought to have been deposited in offshore to nearshore marine settings and represent rare North American examples of Middle Jurassic (Bajocian and Bathonian) dinosaur remains. Some of these new Wyoming sites can be correlated to known dinosaur footprint-bearing horizons or intervals in Utah. Wyoming has a great potential for additional discoveries of new dinosaur footprint-bearing horizons, and further prospecting and study is warranted and will ultimately lead to a much better understanding of the geographic distribution and behavior of the potential footprint-makers.     

Deux nouveaux sites à traces de pas dinosauroïdes dans la Formation Dolomitique de l’Hettangien de Lozère,Languedoc-Roussillon,France

Recent ichnological studies revealed two new localities showing dinosaur footprints in the Hettangian Dolomitic Formation from the Causses Basin, southern France. The traces are reported from the northern part of the basin, in Lozère, an area where Lower Jurassic dinosaur ichnites were poorly documented. The surfaces bearing footprints are characterized by a large number of traces. The ichnites are ascribed to Grallator (Hitchcock, 1858), Dilophosauripus (Welles, 1971) and Eubrontes (Hitchcock, 1845) whose trackmakers were theropods. Although numerous tracksites were previously described in the southern part of the Causses Basin, these three ichnotaxa are for the first time observed in association into the Dolomitic Formation.     

First evidence of stegosaurian Deltapodus footprints in North Africa (Iouaridène Formation, Upper Jurassic, Morocco)

Abstract:  New findings of dinosaur footprints are described from the Upper Jurassic Iouaridène ichnosite of Morocco. On the top of two surfaces, stratigraphically close to that bearing the famous Breviparopus taghbaloutensis trackways, two footprints were excavated and assigned to the ichnogenus Deltapodus . This ichnogenus is well known from the Middle Jurassic of Yorkshire and also occurs in Upper Jurassic deposits from Iberia and the United States. This finding represents the first record of Deltapodus from Africa. These footprints, probably produced by stegosaurian dinosaurs, add new data on the distribution of this type of dinosaur and on the connection between the northern and southern margin of Tethys. 3D models have been generated to allow more detailed studies and to record these unique footprints.     

First record of a fossil bird from the Early Cretaceous of Thailand

We present the first known occurrence of a Mesozoic fossil bird from Thailand. The new specimen is the distal end of a left humerus, from the Early Cretaceous Sao Khua Formation in the Northeast of the country, and testifies to the presence of a medium-sized avian in these non-marine strata. This is also the first Mesozoic bird known from the whole of Southeast Asia. To cite this article: E. Buffetaut et al., C. R. Palevol 4 (2005).     

Dinosaur Tracks from the Kilmaluag Formation (Bathonian,Middle Jurassic) of Score Bay,Isle of Skye,Scotland, UK

Tracks of a juvenile theropod dinosaur with footprint lengths of between 2 and 9 cm as well as adults of the same ichnospecies with footprints of about 15–25 cm in length were found in the Bathonian (Middle Jurassic) Kilmaluag Formation of Score Bay, northwestern Trotternish Peninsula, Isle of Skye, Scotland, UK. Two footprint sizes occur together on the same bedding plane in the central portion of Score Bay, both in situ and on loose blocks. Another horizon containing footprints above this was also identified. The footprints from the lowest horizon were produced in a desiccated silty mud that was covered with sand. A close association of both adults and juveniles with similar travel direction indicated by the footprints may suggest post-hatching care in theropod dinosaurs. Other footprints, produced on a rippled sandy substrate, have been found on the slightly higher bedding plane at this locality. Loose blocks found 130 m to the northeast in the central part of Score Bay have not been correlated with any in situ sediments, but were preserved in a similar manner to those from the higher bedding plane. These tracks represent the youngest dinosaur remains yet found in Scotland.     

Aptian dinosaur footprints from the Apulian platform (Bisceglie,Southern Italy) in the framework of periadriatic ichnosites

New dinosaur tracks have been found near Bisceglie (Bari, Apulia), on loose blocks ascribed to the Corato Member (late Bedoulian to early Gargasian) of the Calcare di Bari Fm. The material consists of isolated footprints as well as of short trackways of quadrupedal and bipedal dinosaurs. The new tracksite has yielded a quite differentiated dinosaur ichnocoenosis, including theropod, sauropod, thyreophoran and ornithopod footprints.The discovery of early Aptian dinosaur footprints in the limestone of the carbonate platform of southern Italy gives new insights on dinosaur distribution, and new palaeontological constraints for the palaeogeographic reconstruction of the Mediterranean Tethys during the Cretaceous. The analysis of this and others ichnosites of the periadriatic carbonate platforms, gives evidence of repeated emersions and of widespread land-vertebrates dwelling. The characteristics of the associations suggest that the trackmakers did not constitute a real coevolved association but the occasional co-occurrence of taxa after migration.The results emphasize the need of both structural and environmental continuity and walking ways between a southern continent and the periadriatic carbonate platforms during the Early Cretaceous.     

安徽省黄山地区恐龙(足迹)脚印化石的初步研究

简要报道了安徽省黄山地区所发现的恐龙足迹化石。从脚印的形态和足迹上看,至少有三种不同的恐龙（蜥脚类、兽脚类、鸟脚类）共同生存过,其中多数恐龙为两足行走性的。记述了两个典型的小型兽脚类和小型鸟脚类恐龙所留下的脚印化石。黄山地区恐龙足迹、骨骼化石及其蛋化石的发现,对于研究晚白垩世恐龙生活习性以及古气候环境均有着一定的意义。     

An enigmatic early mesozoic dinosaur trackway from Zimbabwe

A trackway from Zimbabwe of probably the smallest dinosaur footprints recorded in Africa, is described and tentatively assigned to the Early Jurassic. The footprints are possibly those of a theropod and show strong negative (outward) rotation of the pes and are associated with manus prints. The shape of the footprints, unusual negative rotation, posterior curvature of digit IV and curious positioning of the manus prints in relation to the pes are enigmatic but somewhat reminiscent of Atreipus. Although a number of propositions are considered the most likely is that the animal was an immature dinosaur using a quadrupedal gait. A second trackway of slightly larger footprints of a bipedal theropod dinosaur is also recorded along with other diminutive tracks that suggest an early dinosaur assemblage, possibly dating from near the Trias‐sic‐Jurassic boundary.     

Los Cayos Dinosaur Tracksite: An Overview on the Lower Cretaceous Ichno-Diversity of the Cameros Basin (Cornago,La Rioja Province,Spain)

The Los Cayos dinosaur tracksite is located at the eastern sector of the Cameros Basin (Cornago Township, La Rioja province), NE Spain. The sediments consist of interbedded terrigenous siliciclastics and carbonates belonging to the Enciso Group (Early Cretaceous, Aptian in age). The sedimentological and faunal evidence suggests that these sediments were deposited in a low gradient lacustrine environment. Los Cayos constitutes a relatively wide area with at least 6 localities that have yielded dinosaur tracks. More than 2,000 dinosaur tracks have been discovered to date. Medium to large theropod dinosaurs constitute about 95% of the ichnofauna. Ornithopod dinosaur tracks have been reported from only one of the outcrops (Los Cayos D). Los Cayos S has yielded theropod and sauropod tracks. One sauropod trackway shows a narrow-gauge locomotion pattern and a manus print morphology suggesting that the trackmaker was a titanosaurid, or at least a titanosauriform sauropod. Some pterosaur manus impressions, avian-like footprints of small size and possible turtle tracks complete the assemblage of one of the most impressive and best-preserved dinosaur tracksites of the European Lower Cretaceous.     

Geographical distribution of Anopheles minimus species A and C in western Thailand.

Elucidating vector distribution based on an accurate species identification is important to understanding the nature of the species complex in order to achieve vector control. Morphologically, An. minimus s.l. is difficult to distinguish from both its species complex and its closely related species. A polymerase chain reaction-restriction fragment length polymorphism (PCR-RFLP) technique and a single multiplex-allele specific PCR developed for species identification were applied in this study in comparison with morphological identification. Both methods were used, combining with geographical information systems to determine the distribution of An. minimus species A and C. The investigation on the breeding habitats was performed in the malarious area of western Thailand. Anopheles larvae were collected from 36 bodies of water among five districts (Sangkhaburi, Thong Pha Phum, Si Sawat, Muang, and Sai Yok) of Kanchanaburi Province, Thailand. In this study, An. minimus A larvae were present in all study districts but the association differed when focusing on study sites within each district. Although there were many reports of An. minimus A in Ban Phu Rat and Ban Phu Toei villages in Sai Yok District, we did not find the breeding sites of species A in those two areas. An. minimus A and C were found in Ban Phu Ong Ka village in Sai Yok District. The breeding habitats of An. minimus C were present covering 30-40 km of distance in northern part of Sai Yok and this species was also found in the central and southern parts of Si Sawat District.     

New dinosaur tracksites from the Middle Jurassic of Madagascar: ichnotaxonomical,behavioural and palaeoenvironmental implications

Abstract: New dinosaur tracksites are described from the Bajocian–Bathonian Bemaraha Formation of western Madagascar. Two track‐bearing surfaces can be followed over a distance of at least 4 km, suggesting the existence of a hitherto unrecognized megatracksite. The track assemblage is theropod dominated, but sauropod tracks also occur at one site. Qualitative and quantitative analysis of the abundant theropod track material suggests that most, if not all, theropod footprints are attributable to a single trackmaker and are referred to Kayentapus isp. Although this ichnogenus, originally described from the Lower Jurassic of North America, has never been recorded from Gondwana nor from the Middle Jurassic, track morphology strongly suggests this attribution. Palaeogeographical, sedimentological and ichnological data suggest that the dinosaur tracks formed in an intertidal to supratidal setting where the coastline influenced the preferred walking direction of the animals.     

Dinosaur footprints from the Phra Wihan Formation (Early Cretaceous of Thailand)

The ichnological assemblage of the Phra Wihan Formation (early Early Cretaceous of northeastern Thailand) includes the first sauropod tracks ever found in Thailand. It is reminiscent of the fauna of the underlying Jurassic Phu Kradung Formation with theropods, small ornithischians of uncertain affinities (possibly Hypsilophodon-like ornithopods) and sauropods. It suggests that, in southeastern Asia, an important faunal change occurred in the Early Cretaceous among continental vertebrates, whereas little happened at the Jurassic–Cretaceous boundary.     

陕西子洲中侏罗世早期恐龙足迹初步研究

陕西子洲县中侏罗统延安组砂岩中,同一地点共厚约1.7 m的5个层面发现4种食肉类恐龙足迹,自上而下分别为:1)第5层面大型三趾型足迹,实雷龙足迹科王氏子洲足迹(新遗迹属、新遗迹种)Zizhoupus wangi ichnogen.et ichnosp,nov.;2)第4和第3层面中型三趾型足迹,实雷龙足迹科龙尾峁张北足迹(新遗迹种)Changpeipus longweimaoensis inchnosp.nov.;3)第2层面小型三趾或四趾型足迹,虚骨龙类小理河陕西足迹(新遗迹种)Shensipus xiaoliheensis inchnosp.nov.;4)第1层面小型三趾型足迹,虚骨龙类的铜川陕西足迹Shensipus tungchuanensis.     

辽西下白垩统九佛堂组尾羽龙类—新属(英文)

记述了一件采自辽西热河群九佛堂组一新的窃蛋龙类:义县似尾羽龙(Similicaudipteryx yixianensis gen.et sp.nov.),并依据其和尾羽龙属的相似特征,及其匕首状的尾综骨,肠骨、踱骨和脚趾的形态等将其归入窃蛋龙类的尾羽龙科,但其所具有的一些特征也不同于该科已建立的尾羽龙属,如尾综骨的形态和较大的个体等。该化石具有许多典型的窃蛋龙类的特征,如短尾、较短的前肢等,有别于其他窃蛋龙类的特征还包括耻骨和肠骨的长度比为1.46,背椎上发育2个大而深的椎体下突、背椎侧部具孔等。义县似尾羽龙是又一类具有真正尾综骨的恐龙,表明尾综骨这一曾经被认为是鸟类特有的结构可能是在恐龙中独自演化的。它具有一些进步的特征如具尾综骨等,但同时也具有一些原始的特征如5个愈合的荐椎,耻骨联合长,表明在窃蛋龙类中存在着特征的镶嵌进化现象。似尾羽龙与驰龙类、其他窃蛋龙类等恐龙一样,脚趾并没有对握,已有的证据说明完全的对握目前还只是出现于鸟类中。根据其短尾、中部收缩的趾骨和灵巧的身体等都表明它是一类适于快速奔跑的动物。目前为止,已知的尾羽龙类化石均发现于北票四合屯地区的义县组下部尖山沟层(段),距今约125Ma。新标本发现于义县西二虎桥地点,属于九佛堂组,距今约120Ma,这也是迄今为止在九佛堂组发现的惟一一件尾羽龙类化石,对研究早白垩世窃蛋龙类的演化和热河生物群恐龙组合的特征具有重要的意义。     

Orthogonius species and diversity in Thailand (Coleoptera, Caraboidea, Orthogoniini), a result from the TIGER project

The carabid genus Orthogonius MacLeay is treated, based mainly on materials collected in Thailand through the TIGER project (the Thailand Inventory Group for Entomological Research). Among 290 specimens, 20 species are identified in total, 10 of them are new species: Orthogonius taghavianaesp. n. (Nakhon Nayok: Khao Yai National Park), Orthogonius coomanioidessp. n. (Phetchabun: Thung Salaeng Luang National Park), Orthogonius similarissp. n. (Phetchabun: Thung Salaeng Luang National Park; Loei: Phu Kradueng National Park), Orthogonius setosopalpigersp. n. (Phetchabun: Thung Salaeng Luang National Park), Orthogonius gracililamellasp. n. (Loei: Phu Kradueng National Park; Chaiyaphum: Tat Tone National Park), Orthogonius pseudochaudoirisp. n. (Phetchabum: Thung Salaeng Luang National Park; Nakhon Nayok: Khao Yai National Park), Orthogonius constrictussp. n. (Phetchabum: Thung Salaeng Luang National Park), Orthogonius pinophilussp. n. (Phetchabum: Thung Salaeng Luang National Park), Orthogonius varisp. n. (Cambodia: Siem Reap; Thailand: Ubon Ratchathani: Pha Taem National Park; Phetchabun: Thung Salaeng Luang National Park) and Orthogonius variabilissp. n. (Thailand: Phetchabun: Thung Salaeng Luang National Park; Nakhon Nayok: Khao Yai National Park; Phetchabun: Nam Nao National Park; China: Yunnan). In addition, Orthogonius mouhoti Chaudoir, 1871 and Orthogonius kirirom Tian & Deuve, 2008 are recorded in Thailand for the first time. In total, 30 species of Orthogonius have been recorded from Thailand, indicating that Thailand holds one of the richest Orthogonius faunas in the world. A provisional key to all Thai species is provided. A majority of Thai Orthogonius species are endemic. Among the ten national parks in which orthogonine beetles were collected, Thung Salaeng Luang holds the richest fauna, including 16 species.     

Tetradactyl footprints of an unknown affinity theropod dinosaur from the Upper Jurassic of Morocco

Background

New tetradactyl theropod footprints from Upper Jurassic (Oxfordian-Kimmeridgian) have been found in the Iouaridène syncline (Morocco). The tracksites are at several layers in the intermediate lacustrine unit of Iouaridène Formation. The footprints were named informally in previous works “Eutynichnium atlasipodus”. We consider as nomen nudum.

Methodology/Principal Findings

Boutakioutichnium atlasicus ichnogen. et ichnosp. nov. is mainly characterized by the hallux impression. It is long, strong, directed medially or forward, with two digital pads and with the proximal part of the first pad in lateral position. More than 100 footprints in 15 trackways have been studied with these features. The footprints are large, 38–48 cm in length, and 26–31 cm in width.

Conclusions/Significance

Boutakioutichnium mainly differs from other ichnotaxa with hallux impression in lacking metatarsal marks and in not being a very deep footprint. The distinct morphology of the hallux of the Boutakioutichnium trackmaker –i.e. size and hallux position- are unique in the dinosaur autopodial record to date.     

Parataxonomy and palaeobiogeographic significance of dinosaur eggshell fragments from the Upper Cretaceous strata of the Cauvery Basin,South India

Abstract

In the past, fossilised dinosaur eggshells have been extensively documented from the Upper Cretaceous Lameta Formation of Central India and as many as nine oospecies are known at present from this formation. Compared to this, only one dinosaur oospecies has been described from the Cretaceous succession of the Cauvery Basin. However, the first fossil egg from India, identified as a chelonian egg, was documented from the Aptian – Albian Karai Formation of the Cauvery Basin in 1957. Following this, a solitary titanosaurid dinosaur egg was described from the Upper Cretaceous (Lower Maastrichtian) Kallankuruchhi Formation, Cauvery Basin in 1996. More recently, we have recovered isolated eggshell fragments from the marine part of the Upper Cretaceous (Late Maastrichtian) Kallamedu Formation. Based on eggshell morphology, microstructure and ultrastructure, these eggshell fragments are assigned to the oospecies Fusioolithus baghensis. The new find from the Cauvery Basin is important from palaeobiogeographic point of view as the oofamily Fusioolithidae is found in the Upper Cretaceous strata of India, France, Argentina and Morocco. Based on the common occurrence of similar oospecies in South America, Africa, Europe and India, a Late Cretaceous palaeobiogeographic connection between India and South America as well as Europe via Africa is suggested.