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1.
Colonial island‐breeding birds can be particularly vulnerable to anthropogenic disturbance, which can adversely affect their nesting success. We studied Sooty Falcons Falco concolor breeding on 10 ground‐predator‐free islands in the Sea of Oman during 2007–2014 and evaluated spatio‐temporal trends in the number of breeding pairs occurring on the islands and the factors influencing nesting success. The number of breeding pairs on the islands declined during the study, due mostly to the decline on accessible islands; the rate of decline on islands accessible to humans was double that on inaccessible ones. The number of nests with one or more eggs declined during the study period, and the percentage of nests with eggs that produced one or more chicks showed an increasing trend over time. Sooty Falcon nests located farther away from beaches experienced a significantly higher probability of nesting success than those located closer to beaches. Our results suggest that the number of breeding Sooty Falcons on the islands of northern Oman is declining and that human disturbance may be a contributing factor; this probably mirrors the situation in other parts of the breeding range of this species.  相似文献   

2.
Amur Falcons (Falco amurensis) are a migratory species that face a variety of threats across their range, but little is known about their breeding ecology. These falcons breed in forest habitats in Eastern and Central Asia using nests constructed by corvids, including Eurasian Magpies (Pica pica). We monitored nests of 21 pairs of Amur Falcons at Hustai National Park in central Mongolia in 2017. Our objectives were to describe their basic nesting ecology, estimate nest survival by modeling the daily survival rate (DSR), examine nest selection by modeling it as a function of nest and site covariates, and use a spatial simulation to test hypotheses concerning intra‐ and interspecific avoidance. Clutch sizes averaged 4.1 eggs (= 21 nests), and incubation and nestling periods averaged 25.7 and 26.1 d, respectively. The daily survival rate was 0.98, with young in 12 nests surviving to fledging. Nest structures were more likely to be selected as percent cover of nest bowls increased, usually in the form of a dome of sticks with multiple side entrances. Closed nests likely provide increased protection from predators. In contrast to congeneric Red‐footed Falcons (F. vespertinus) that nest in large colonies, Amur Falcons nested no farther from or closer to nests of either conspecifics or congeners than expected by chance. One factor likely contributing to this difference is that Red‐footed Falcons often use the nests of colonial‐nesting Rooks (Corvus frugilegus), whereas Amur Falcons typically use the nests of non‐colonial Eurasian Magpies. The ongoing loss of deciduous trees like white birch (Betula platyphylla) across the breeding range of Amur Falcons, probably due to climate change and increased grazing pressure, is likely to reduce the availability of nesting habitat for Eurasian Magpies which, in turn, will likely reduce availability of nests for Amur Falcons and other small falcons.  相似文献   

3.
The selection of a nest site is crucial for successful reproduction of birds. Animals which re‐use or occupy nest sites constructed by other species often have limited choice. Little is known about the criteria of nest‐stealing species to choose suitable nesting sites and habitats. Here, we analyze breeding‐site selection of an obligatory “nest‐cleptoparasite”, the Amur Falcon Falco amurensis. We collected data on nest sites at Muraviovka Park in the Russian Far East, where the species breeds exclusively in nests of the Eurasian Magpie Pica pica. We sampled 117 Eurasian Magpie nests, 38 of which were occupied by Amur Falcons. Nest‐specific variables were assessed, and a recently developed habitat classification map was used to derive landscape metrics. We found that Amur Falcons chose a wide range of nesting sites, but significantly preferred nests with a domed roof. Breeding pairs of Eurasian Hobby Falco subbuteo and Eurasian Magpie were often found to breed near the nest in about the same distance as neighboring Amur Falcon pairs. Additionally, the occurrence of the species was positively associated with bare soil cover, forest cover, and shrub patches within their home range and negatively with the distance to wetlands. Areas of wetlands and fallow land might be used for foraging since Amur Falcons mostly depend on an insect diet. Additionally, we found that rarely burned habitats were preferred. Overall, the effect of landscape variables on the choice of actual nest sites appeared to be rather small. We used different classification methods to predict the probability of occurrence, of which the Random forest method showed the highest accuracy. The areas determined as suitable habitat showed a high concordance with the actual nest locations. We conclude that Amur Falcons prefer to occupy newly built (domed) nests to ensure high nest quality, as well as nests surrounded by available feeding habitats.  相似文献   

4.
Ground and aerial surveys of breeding Sooty Falcons Falco concolor were conducted along the western coast of Saudi Arabia and its offshore islands from August to October 1991 and 1992. An estimated population of 260–381 pairs was found. The known world breeding population is still below 1000 pairs. Most rocky islands were occupied except the largest islands of the Farasan archipelago. Among the many low islets surrounded with mangrove, only one small group near Al Lith harboured breeding Sooty Falcons. The distribution pattern, density and breeding success of the falcons seemed to be strongly influenced by food availability (autumn migration of small and medium-size birds) and the deterring presence of terrestrial predators. Human disturbance and trapping are still of local importance.  相似文献   

5.
Capsule: We report a significant reduction in population size and breeding success for the Lanner Falcon Falco biarmicus in Sicily, its biggest stronghold in Europe, since the latest coordinated survey.

Aims: To provide updated information on current population size of Lanner Falcon in Sicily and to compare breeding parameters with those obtained in previous studies.

Methods: We performed an intensive coordinated field survey and literature review of breeding success parameters across the species range.

Results: Overall, we monitored 126 territories throughout Sicily where the species had been reported in the last 15 years. Lanner Falcons were present only in 60 of them. Mean nest productivity (± standard deviation) was 1.09?±?1.18 fledged young/checked pairs, flight rate was 2.22?±?0.52 fledged young/successful pairs and breeding success was 49.0%.

Conclusions: Indirect measures aimed at preventing abandonment of occupied territories should be applied, for instance by developing a network of priority areas and slowing down degradation of the pseudo-steppe habitats by agri-environmental schemes. Additionally, direct measures aimed at preventing nest robbery, including the organization of nest guarding activities, and reduction of anthropogenic disturbance and illegal shooting, must be encouraged in order to avoid territory abandonment.  相似文献   

6.
Nestbox programmes are frequently implemented for the conservation of cavity‐nesting birds, but their effectiveness is rarely evaluated in comparison with birds not using nestboxes. In the European Palaearctic, Red‐footed Falcon Falco vespertinus populations are both of high conservation concern and are strongly associated with nestbox programmes in heavily managed landscapes. We used a 21‐year monitoring dataset collected on 753 nesting attempts by Red‐footed Falcons in unmanaged natural or semi‐natural habitats to provide basic information on this poorly known species; to evaluate long‐term demographic trends within this population; and to evaluate response of demographic parameters of Red‐footed Falcons to environmental factors including use of nestboxes. We observed significant differences among years in laying date, offspring loss and numbers of fledglings produced, but not in egg production. Of these four parameters, offspring loss and, to a lesser extent, number of fledglings exhibited directional trends over time. Variation in laying date and in numbers of eggs were not well explained by any one model of environmental factors, but instead by combinations of models, each with informative terms for nest type. Nevertheless, laying in nestboxes occurred 2.10 ± 0.70 days earlier than in natural nests. In contrast, variation in both offspring loss and numbers of fledglings produced were fairly well explained by a single model including terms for nest type, nest location and an interaction between the two parameters (65 and 81% model weights, respectively), with highest offspring loss in nestboxes on forest edges. Because, for other species, earlier laying dates are associated with more fit individuals, this interaction highlighted a possible ecological trap, whereby birds using nestboxes on forest edges lay eggs earlier but suffer greater offspring loss and produce lower numbers of fledglings than do those in other nesting settings. If nestboxes increase offspring loss for Red‐footed Falcons in heavily managed landscapes where populations are at greater risk, or for the many other species of rare or endangered birds supported by nestbox programmes, these processes could have important demographic and conservation consequences.  相似文献   

7.
The numbers of Falco cherrug and Falco peregrinus trapped during their migration over the Kingdom of Saudi Arabia (KSA) were investigated from published reports and through interviews with well-known trappers and dealers over several years (1989–2013). The number of trapped individuals increased for both species over a 23 year period, which is probably related to an enhanced trapping effort. Time series analysis suggests that the number of Saker Falcons being trapped is likely to be stable with annual fluctuations in the coming ten-year period, whereas the number of trapped Peregrine Falcons will probably decline with a small fluctuation initially. Using the population viability analysis suggests a high extinction rate for the Saker Falcon population migrating through KSA during the coming 10 and 20 years; whereas Peregrine Falcons probably take more than 100 years to reach the extinction threshold. However, the increase in the trapping period, especially in the spring, that has been observed during the last five years could increase the number of falcons trapped in the future. As both falcon species are migratory, implementing conservation actions across all range states is important to ensure a favourable conservation status for the Saker and Peregrine Falcons. Both species will benefit through the implementation of the Global Action Plan (GAP), developed by the Saker Falcon Task Force.  相似文献   

8.
Anthropogenic landscapes can be rich in resources, and may in some cases provide potential habitat for species whose natural habitat has declined. We used remote videography to assess whether reintroducing individuals of the threatened New Zealand falcon Falco novaeseelandiae into a highly modified agricultural habitat affected the feeding rates of breeding falcons or related breeding behavior such as nest attendance and brooding rates. Over 2,800 recording hours of footage were used to compare the behavior of falcons living in six natural nests (in unmanaged, hilly terrain between 4 km and 20 km from the nearest vineyard), with that of four breeding falcon pairs that had been transported into vineyards and nested within 500 m of the nearest vineyard. Falcons in vineyard nests had higher feeding rates, higher nest attendance, and higher brooding rates. As chick age increased, parents in vineyard nests fed chicks a greater amount of total prey and larger prey items on average than did parents in hill nests. Parents with larger broods brought in larger prey items and a greater total sum of prey biomass. Nevertheless, chicks in nests containing siblings received less daily biomass per individual than single chicks. Some of these results can be attributed to the supplementary feeding of falcons in vineyards. However, even after removing supplementary food from our analysis, falcons in vineyards still fed larger prey items to chicks than did parents in hill nests, suggesting that the anthropogenic habitat may be a viable source of quality food. Although agricultural regions globally are rarely associated with raptor conservation, these results suggest that translocating New Zealand falcons into vineyards has potential for the conservation of this species.  相似文献   

9.
The estimation of abundance is fundamental to ecology and conservation but often is difficult or impossible to accomplish reliably. Recent improvements in wildlife cameras and ecological modelling have allowed for improved accuracy in estimates of abundance. In this study, we paired nest captures and high-definition nest video camera monitoring with modelling for a novel approach to estimate survival and abundance of threatened Piping Plovers Charadrius melodus breeding on Missouri River sandbars. From 2005 to 2014, we captured individuals on nests and uniquely marked them and recaptured previously marked individuals. In 2015–2017, we resighted marked individuals using small, high-definition video cameras deployed at nests, and counted the number of marked and unmarked breeding individuals associated with nests. We estimated apparent survival and derived estimates of the abundance of breeding individuals and population growth each year using a state-space Jolly–Seber superpopulation model with the addition of a binomial band ratio model for data collected using nest video cameras. Apparent survival averaged 0.73 ± 0.03 (mean ± sd) throughout the study. The number of breeding individuals varied, with the population increasing from 2012 to 2017 following a major habitat creation event. This study provides one of the few examples of camera data being used to produce demographic parameter and abundance estimates for an avian species. The camera and modelling methods described in this study may be applicable to other avian species in which some portion of the breeding population is uniquely marked.  相似文献   

10.
The Spotted Barbtail (Premnoplex brunnescens) inhabits the understory of humid montane forests in Central and South America. Apart from basic information on eggs and nest form, little has been published on its breeding ecology. Using temperature sensors in nest cups, I have collected data on the diurnal patterns of egg-coverage from three nests in eastern Ecuador and reveal a remarkable incubation rhythm. After providing near-constant coverage during the morning, adults leave the eggs unattended for most of the afternoon, returning to the nest only in the late afternoon. The mean duration (±standard deviation) of this period of absence, across the entire incubation period at three nests, was 6.4 ± 1.9 h. These results are discussed in relation to their physiological and ecological significance.  相似文献   

11.
Red-breasted goose colonies have been studied near Medusa Bay (73°21′N, 80°32′E), on the northwestern Taimyr Peninsula, and along the Agapa River (70°11′N, 86°15′E) down to its mouth (71°26′N, 89° 13′E), in the central Taimyr Peninsula. Red-breasted geese nesting near peregrine falcons are protected by the falcons from arctic foxes; however, they are sometimes attacked by the falcons themselves. In the colonies near peregrine falcon nests, the vast majority of goose nests were situated no farther than 100 m from the falcon nest. When food is abundant, falcons protect a larger area around their nest. The distance between the falcon nest and the surrounding goose nests is inversely related to the falcon’s activity. In years of higher falcon activity, falcons prevent red-breasted geese from nesting as close to their nest as in years of lower falcon activity. Additional stimuli are required for red-breasted geese to form colonies near rough-legged buzzard nests. The distance between snowy owl nests and red-breasted goose nests was smaller when arctic foxes were abundant than when they were scarce.  相似文献   

12.
Provisioning regimes and growth of Light-mantled Sooty Albatrosses (LMSA) (Phoebetria palpebrata) were investigated on subantarctic Macquarie Island using an automatic tracking system and automatic weighing nests. The nests were deployed under five chicks in the post-brood provisioning period in 2000 and 2001. Adults typically utilised a cyclical foraging strategy consisting of a long foraging trip followed by three to four shorter trips. Chicks received an average (±SE) of 37.5±2.3 kg of food in the post-brood provisioning period and were fed every 1.6±0.1 days with a mean meal size of 520±10 g. Chicks grew at a rate of 61.3±1.0 g day-1 to a peak mass of 4.4±0.1 kg. Mean chick fledging mass was 3.0±0.1 kg. LMSA on Macquarie Island fed their chicks more frequently and showed a lower mean trip duration than conspecifics at South Georgia, which is likely related to proximity of productive Antarctic shelf waters, differences in prey availability and competition with other Procellariiformes.  相似文献   

13.
G. L. Maclean 《Ostrich》2013,84(3-4):241-253
Maclean, G. L. 1973. The Sociable Weaver, Part 4: Predators, parasites and symbionts. Ostrich 44: 241–253.

The main nest predator of the Sociable Weaver in the Kalahari sandveld is the Cape Cobra Nala nivea. This snake causes great losses of eggs and chicks; one cobra may eat the contents of an entire nest mass at one feed. Another nest predator which causes smaller losses of eggs and chicks but great destruction to the nest masses is the Honey Badger Mellivora capensis. These are the only two nest predators on the Sociable Weaver in the study area. Predators on adult Sociable Weavers include several birds of prey and some small carnivorous mammals.

Adult Sociable Weavers have few ectoparasites and hardly any Mallophaga. A common ectoparasite on the legs of chicks is a blood-sucking Dermestes larva, which appears not to be harmful. The only endoparasite found was the nematode, Diplotriaena ozouxi, which infected the abdominal air sacs of the adults.

The nest material of the Sociable Weavers' communal nest masses was inhabited by a wealth of invertebrate animals and a few harmless reptiles such as skinks and geckos.

Some of the chambers in a Sociable Weaver nest mass may be taken over by other species of birds. Most of these, such as Redheaded Finches Amadina erythrocephala, use the chambers for breeding purposes only, but the most important avian symbiont, the Pygmy Falcon Polihierax semitorquatus, is a permanent resident in the chambers. The presence of Pygmy Falcons is resented by the weavers but the falcons may help to keep snakes away from the nest mass. Adult Sociable Weavers are not normally preyed on by Pygmy Falcons, although the falcons may occasionally take young weavers in the nest chambers.

The tops of the nest masses may be used as nest sites by the Giant Eagle Owl Bubo lacteus. Barn Owls Tyto alba may use cavities in the superstructure of nest masses for roosting in. Neither of these owls appeared to prey on the weavers.  相似文献   

14.
Investigating the reproductive ecology of naturalized species provides insights into the role of the source population's characteristics vs. post‐release adaptation that influence the success of introduction programmes. Introduced and naturalized Mallards Anas platyrhynchos are widely established in New Zealand (NZ), but little is known regarding their reproductive ecology. We evaluated the nesting ecology of female Mallards at two study sites in NZ (Southland and Waikato) in 2014–15. We radiotagged 241 pre‐breeding females with abdominal‐implant transmitters and measured breeding incidence, nesting chronology and re‐nesting propensity. We monitored 271 nests to evaluate nest survival, clutch and egg size, egg hatchability and partial clutch depredation. Breeding incidence averaged (mean ± se) 0.91 ± 0.03, clutch size averaged 9.9 ± 0.1 eggs, 94 ± 2% of eggs hatched in successful nests, partial depredation affected 6 ± 1% of eggs in clutches that were not fully destroyed by predators, and re‐nesting propensity following failure of nests or broods was 0.50 ± 0.003. Nesting season (first nest initiated to last nest hatched) lasted 4.5 months and mean initiation date of first detected nest attempts was 28 August ± 3.3 days. Smaller females were less likely to nest, but older, larger or better condition females nested earlier, re‐nested more often and laid larger clutches than did younger, smaller or poorer condition females. Younger females in Southland had higher nest survival; cumulative nest survival ranged from 0.25 ± 0.007 for adult females in Waikato to 0.50 ± 0.007 for yearling females in Southland. Compared with Mallards in their native range, the nesting season in NZ was longer, clutches and eggs were larger, and nest survival was generally greater. Different predators and climate, introgression with native heterospecifics and/or the sedentary nature of Mallards in NZ may have contributed to these differences.  相似文献   

15.
Habitat availability might be the most important determinant of success for a species reintroduction programme, making investigation of the quality and quantity of habitat needed to produce self‐sustaining populations a research priority for reintroduction ecologists. We used a stochastic model of population dynamics to predict whether attempts to improve existing breeding territories using artificial nest platforms improved the population growth rate and persistence of a reintroduced population of Northern Aplomado Falcons Falco femoralis septentrionalis in South Texas. We further assessed whether the creation of new territories, i.e. conversion of entire areas to suitable habitat and not simply the erection of nest platforms, would lead to a subsequent increase in the nesting population. Our model was able to reproduce several characteristics of the wild population and predicted the number of breeding pairs per year strikingly well (R2 = 0.97). Simulations revealed that the addition of nest platforms improved productivity such that the population would decline to extinction without them but is stable since their installation. Moreover, the model predicted that the increase in productivity due to nest platforms would cause the population to saturate available breeding territories, at which point the population would contain a moderate proportion of non‐territorial birds that could occupy territories if new ones become available. Population size would therefore be proportional to the increase in available territories. Our study demonstrates that artificial nest‐sites can be an effective tool for the management of reintroduced species.  相似文献   

16.
Between 2004 and 2007, we studied density, habitat features and breeding parameters of the osprey (Pandion haliaetus) population in Boa Vista Island (Cape Verde). A total of 79 nest structures were identified, 37 of which were occupied for at least 1 year during the study period. The osprey population ranged between 14 and 18 pairs, and the mean density and distance between neighbouring occupied nests were 2.58 pairs per 100 km2 and 3089 m, respectively. Occupied nests were found to be significantly further from the coastline and roads than unoccupied nests, but the distances from villages were similar. The majority (81.1%) of the 37 occupied nests were easily accessible to humans. Mean clutch size was 2.59, average productivity was 0.76 young/active nest, and breeding success was 58.8% [Correction added on 13 May 2013, after first online publication: the average productivity was changed from 0.72 to 0.76]. Density in Boa Vista was higher than that in other sedentary island populations in the Western Palearctic, whereas the productivity was the lowest of this region. Clutch size did not vary among Western Palearctic populations, but the differences observed in productivity were likely influenced by local factors that in Boa Vista are attributed to nest depredation by the brown‐necked raven (Corvus ruficollis) and to direct human persecution.  相似文献   

17.
Andrew R. Jenkins 《Ostrich》2013,84(3-4):281-290
Jenkins, A.R. 1994. The influence of habitat on the distribution and abundance of Peregrine and Lanner Falcons in South Africa. Ostrich 65: 281–290.

The distribution and abundance of Peregrine and Lanner Falcons in South Africa was compared using recorded sightings from various sources, including the Southern African Bird Atlas Project. Falcon distributions were compared with the distribution of cliffs and vegetation, to quantify differences in the habitat preferences of the two species in the breeding and the non-breeding seasons. Lanner Falcons outnumbered Peregrine Falcons in most areas by at least 10:1. Peregrine Falcons were more habitat specific than Lanner Falcons, in terms of topographic and biotic requirements. Peregrine Falcons were largely restricted to high cliff areas throughout the year and there probably were no large-scale seasonal movements within the population. The bulk of the resident Peregrine Falcon population was found in the fynbos biome, in the southwestern Cape. Outside of this area, Peregrine Falcons were concentrated in woodlands. Lanner Falcons were less dependent on high cliffs, although cliff availability was important in defining the ranges of both species. Lanner Falcons were most common in the sour grasslands in the east of the country in the breeding season, with apparent movements in the non-breeding season into the fynbos, the Nama Karoo and the southern Kalahari. Overall, Peregrine Falcons favoured relatively closed habitats and Lanner Falcons favoured relatively open habitats. The differences in the two species' habitat preferences are proximate factors influencing distribution and abundance.  相似文献   

18.
Understanding breeding phenology and success can elucidate population dynamics, which is especially important for species in need of conservation. We describe the factors affecting the breeding biology of American Oystercatchers (Haematopus palliatus frazari) at El Rancho Island, a critical site that contains ~ 7% of the total estimated population, on the coast of Sinaloa, Mexico. We monitored 192 nests over four years (2016–2019). The breeding season lasted from March to June and mean laying dates differed among years, with the mean laying date in 2019 an average of 20 days earlier than in 2016. Clutch sizes decreased as the breeding season progressed. Both breeding success and productivity differed among years, with the lowest values in 2016 (30% hatching success and 0.6 chicks/nest) and the highest in 2019 (66% hatching success and 1.2 chicks/nest). Hatching success was affected by year, laying date, type of habitat, and distance to the high tide line. American Oystercatchers that laid eggs earlier in the season, used mixed marsh and dune habitat, and with nests relatively close to the waterline (< 50 m) had greater breeding success. Overall, however, the breeding success of American Oystercatchers was low and influenced by a combination of several intrinsic and extrinsic factors. Management measures may be required to increase breeding success and ensure the conservation of this subspecies.  相似文献   

19.
Nest attendance is an important determinant of avian reproductive success, and identifying factors that influence the frequency and duration of incubation recesses furthers our understanding of how incubating birds balance their needs with those of their offspring. We characterized the frequency and timing (start time, end time, and duration) of incubation recesses for mallard (Anas platyrhynchos) and gadwall (Mareca strepera) hens breeding in Suisun Marsh, California, USA, and examined the influences of day of year, ambient temperature at the nest, incubation day, and clutch size on recess frequency and timing using linear mixed models. Mallard, on average, took more recesses per day (1.69 ± 0.80, mean ± standard deviation) than did gadwall (1.39 ± 0.69), and 45% of mallard nest‐days were characterized by two recesses, while only 27% of gadwall nest‐days were characterized by two recesses. Mallard morning recesses started at 06:14 ± 02:46 and lasted 106.11 ± 2.01 min, whereas mallard afternoon recesses started at 16:39 ± 02:11 and lasted 155.39 ± 1.99 min. Gadwall morning recesses started at 06:30 ± 02:46 and lasted 91.28 ± 2.32 min, and gadwall afternoon recesses started at 16:31 ± 01:57 and lasted 192.69 ± 1.89 min. Mallard and gadwall started recesses earlier in the day with increasing ambient temperature, but later in the day as the season progressed. Recess duration decreased as the season progressed and as clutch size increased, and increased with ambient temperature at the nest. The impending darkness of sunset appeared to be a strong cue for ending a recess and returning to the nest, because hens returned to their nests earlier than expected when recesses were expected to end after sunset. Within hens, the timing of incubation recesses was repeatable across incubation days and was most repeatable for mallard afternoon recesses and on days in which hens took only one recess. Hens were most likely to be away from nests between 04:00 and 07:00 and between 16:00 and 19:00; therefore, investigators should search for nests between 07:00 and 16:00. Our analyses identified important factors influencing incubation recess timing in dabbling ducks and have important implications for nest monitoring programs.  相似文献   

20.
《Ostrich》2013,84(1-2):117-118
The Seychelles Fody, Foudia sechellarum, is a ploceid weaver occurring naturally on three islands in the Seychelles group in the Indian Ocean. The population on Cousine Island was studied between 30 June and 25 August 1997. The size of the population on the island was estimated at 458–614 individuals and densities varied in different habitat types. As Seychelles Fodies in non-breeding plumage are difficult to sex, we provide sexing criteria based on wing length. Breeding pairs form small, probably temporary, territories that are defended by both partners against other fodies, including the introduced Madagascar Fody, Foudia madagascariensis. The Seychelles Fody often breeds semi-colonially and we observed up to five nests close together. The birds are socially monogamous and both sexes share in nest building, nest defense and provisioning of the young. Only females incubate. Many nests were deserted before eggs were laid, including some that had been accepted by the female. Additional males and females were sometimes seen helping provision the young, but this cooperative breeding behaviour appeared to be uncommon. Non-breeding individuals congregate in large flocks, sometimes joined by breeding birds.  相似文献   

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