The Ordovician-Silurian boundary: new data, new concerns

Despite International Union of the Geological Sciences (IUGS) ratification of the International Commission on Stratigraphy recommendation that the base of the Parakidograptus acuminatus zone be the base of the Silurian and that the boundary stratotype be at Dob's Linn, Scotland, numerous concerns continue to be expressed over the timing of the selection as well as the boundary and its stratotype. The richness of both graptolite and non-graptolite biostratigraphic data in numerous stratigraphic sections from many areas in China published at about the same time that the boundary and stratotype were approved formally and shortly after approval suggest the possibility that boundary designation may have been somewhat premature. The new data from China document clearly the major extinction of characteristic Late Ordovician taxa significantly below the presently accepted System boundary. The new data indicate that the re-radiation of graptolites after the extinction took place in the Glyptograptus persculptus zone, the graptolite zone subjacent to the P. acuminatus zone. New geochemical data from Dob's Linn reveal a change in chemistry in strata subjacent to the base of the G. persculptus zone that could be used as an aid in correlation. The new data suggest the possibility that the base of the G. persculptus zone, the relatively traditional base of the Silurian, could be as appropriate a System boundary as the base of the P. acuminatus zone.     

The Ordovician-Silurian boundary stratotype: consequences of its approval by the IUGS

Lespérance, Pierre J., Barnes, Christopher R., Berry, William B. N., Boucot, Arthur J. & Mu En-zhi 1987 07 15: The Ordovician-Silurian boundary stratotype: consequences of its approval by the IUGS.
The Ordovician-Silurian stratotype at Dob's Linn, Scotland is the second systemic boundary approved by the International Union of Geological Sciences (IUGS). A review of the internationally accepted criteria required of a stratotype shows that few of these are possessed by the Dob's Linn section. The strongest attribute of the section is the presence of zonal graptolites, although the boundary is recognized primarily on a single biological event (base of acuminatus Zone) for which the evolutionary relationships of the taxa are not established. In approving this boundary proposal in 1985, which received only simple majority support within the Ordovician-Silurian Boundary Working Group (OSBWG) and which fails to meet most of the accepted prerequisites for a stratotype, the International Commission on Stratigraphy (ICS) and the IUGS have established an unfortunate precedent. It follows that future systemic boundaries need not meet the accepted standards. It raises serious questions on the assessment and voting procedures of the International Commission on Stratigraphy and on the credence accorded the recommendations developed by the International Subcommission on Stratigraphic Classification of IUGS.     

Evolution of the taxonomic diversity of Mongolian Ordovician-Silurian corals

Changes in the taxonomic diversity of Mongolian Ordovician-Silurian corals (Tabulatoidea, Heliolitoidea, Cyrtophyllida, Rugosa) are analyzed. Evolution of the taxonomic diversity is analyzed in two aspects: quantitative and qualitative changes. Changes in the diversity of Mongolian corals are caused by (1) evolutional developmental stage of the group and its adaptational possibilities (internal reason) and (2) changes in the environmental parameters, which reproduce regional and general global events (external reason).     

Distribution of Ordovician-Silurian ichnofossil assemblages in Wales - implications for Phanerozoic ichnofaunas

The stratigraphic succession of west Wales may be subdivided into two dominant lithologies, namely sandstone and mudstone. The distribution of these lithologies is largely determined by tectonic activity and eustatic sea-level changes in the area. The strata contain a diverse and relatively abundant ichnofaunal assemblage consisting, at the ichnogeneric level, of fifteen forms: Chondrites, Circulichnis , Cochlichnus, Cosmorhaphe, Desmograpton, Gordia, Helminthoida, Helminthopsis , Nereites, Neonereites, Palaeo-phycus, Paleodictyon (Glenodictyon), Paleodictyon (Squamodictyon) , Planolites, Protopaleodictyon and Spirophycus. The ichnogenera are not evenly distributed throughout the succession, the main controlling factors being toponomy, anoxia, water depth and a global extinction event. The relative importance of these controls is examined with reference to 33 previously described and taxonomically well-documented deep-water flysch ichnofossil studies. □ Trace fossils, Palaeozoic, Wales, toponomy, anoxia, depth, extinction.     

The relation of Late Ordovician glaciation to the Ordovician-Silurian changeover in North American brachiopod faunas

Sheehan, P. M.: The relation of Late Ordovician glaciation to the Ordovician-Silurian changeover in North American brachiopod faunas.
The Ordovician-Silurian changeover of brachiopod faunas in North American epicontinental seas involved the abrupt extinction of endemic Late Ordovician stocks and subsequent repopulation of North American seas by Old World taxa. The Late Ordovician Gondwanaland glaciation may have lowered sea levels sufficiently to place severe stress on the widespread shallow marine faunas in North America, resulting in their eventual extinction. The Late Ordovician depositional history in North America is not well enough known to establish the presence of a latest Ordovician regression, but the earliest Silurian was an interval of off-lap in North America. Therefore, the glacial lowering of sea level is considered to be the most likely cause of the faunal changeover.     

Convection-free boundary sedimentation

A simple technique is described that prevents the occurrence of convective flows in boundary sedimentation experiments. Superimposed density gradients employed as in band centrifugation are evaluated quantitatively. The usefulness of the method is demonstrated by comparison of stabilized and nonstabilized experiments run under identical conditions, with considerable variations in temperature introduced on purpose. The technique is shown to be of special value for the investigation of polydispersity problems.     

On boundary stimulation and optimal boundary control of the bidomain equations

The bidomain equations with Neumann boundary stimulation and optimal control of these stimuli are investigated. First an analytical framework for boundary control is provided. Then a parallel finite element based algorithm is devised and its efficiency is demonstrated not only for the direct problem but also for the optimal control problem. The computations realize a model configuration corresponding to optimal boundary defibrillation of a reentry phenomenon by applying current density stimuli.     

The scs' boundary element: characterization of boundary element-associated factors.

Boundary elements are thought to define the peripheries of chromatin domains and to restrict enhancer-promoter interactions to their target genes within their domains. We previously characterized a cDNA encoding the BEAF-32A protein (32A), which binds with high affinity to the scs' boundary element from the Drosophila melanogaster 87A7 hsp70 locus. Here, we report a second protein, BEAF-32B, that differs from 32A only in its amino terminus. Unlike 32A, it has the same DNA binding specificity as the complete BEAF activity affinity purified from Drosophila. We characterize three domains in these proteins. Heterocomplex formation is mediated by their identical carboxy-terminal domains, and DNA binding is mediated by their unique amino-terminal domains. The identical middle domains of 32A and 32B are dispensable for the functions described here, although they may be important for boundary element function. 32A and 32B apparently form trimers, and the ratio of 32A to 32B varies at different loci on polytene chromosomes as judged by immunofluorescence. The scs' element contains a high- and low-affinity binding site for BEAF. We observed that interaction with the low-affinity site is facilitated by binding to the high-affinity site some 200 bp distant.     

View from the boundary

Re-implementing biological mechanisms on robots not only has technological application but can provide a unique perspective on the nature of sensory processing in animals. To make a robot work, we need to understand the function as part of an embodied, behaving system. I argue that this perspective suggests that the terms "representation" and "information processing" can be misleading when we seek to understand how neurobiological mechanisms carry out perceptual processes. This argument is presented here with reference to a robot model of cricket behavior, which has demonstrated competence comparable to that of the insect, but utilizes surprisingly simple central processing. Instead it depends on sensory interfaces that are well matched to the task, and on the link between environment, action, and perception.     

Analysis of the midbrain-hindbrain boundary cell fate using a boundary cell-specific Cre-mouse strain

We describe here a transgenic mouse line MHB-Cre, which expresses Cre recombinase in a group of cells at the midbrain-hindbrain boundary. Using this mouse line, we studied the contribution of the boundary cells to distinct brain areas during development. Initially, the MHB-Cre expression coincides with that of Cdh22 and p21 around the Otx2 expression border in a narrow population of cells with reduced proliferative activity. Consistent with their location on both sides of the Otx2 expression border, the Cre expressing boundary cells contribute both to midbrain as well as hindbrain. However, the majority of recombinant cells remain close to the mid- and hindbrain border, suggesting very limited cell mixing within these brain compartments during development. Interestingly, dorsocaudally oriented fibers of the midbrain dopaminergic neurons follow the path marked by the boundary cells.     

The Cambro - Ordovician boundary

Henningsmoen, G.: The Cambro-Ordovician boundary.
The Cambrian and Ordovician Systems were established in Britain, and it seems appropriate to define the Cambro-Ordovician boundary in Britain or where there is a closely related faunal succession at the junction. When Lapworth erected the Ordovician System in 1879, his intention was that its lower boundary should separate the so-called primordial (or first) and second Palaeozoic faunas. He drew this boundary at the base of the Lower Arenig, but apparently included the Upper Tremadoc in the Lower Arenig. It seems problematic to base the Cambro-Ordovician boundary on grounds of historical priority, and we seem to be free to decide whether to have the boundary at the base of the Tremadocian, at the base of the Arenigian, or within the Tremadocian (e.g. at the base of the Upper Tremadoc). The alternatives are shortly discussed. Arguments are put forward for placing the boundary not at a break, but preferably in a sequence with cosmopolitan fossils, either in a sequence of uniform facies or perhaps rather in a sequence with frequently alternating facies.