A new species of Tococladidae CARPENTER, 1966 from the Permian of France (Insecta: Archaeorthoptera)

A new forewing venation pattern of the Tococladidae is proposed and the phylogenetic affinities of this family are discussed, based on Tococladus garrici nov. sp. This fossil comes from the Salagou Formation (Permian, Saxonian Group, Lodève basin, France). The Tococladidae is assigned to the Archaeorthoptera BÉTHOUX and NEL, 2002. The biostratigraphic implications of this discovery are briefly discussed, but it is still not possible to infer any definite conclusion about the age of this formation.     

The fourth new species of the leafhopper genus Thaioneura Song,Li & Dietrich from China (Hemiptera,Cicadellidae, Typhlocybinae)

In this paper, one new species of the leafhopper genus Thaioneura Song, Li & Dietrich, 2016 from Guizhou Province, China are described and illustrated: Thaioneura huajiangensis sp. nov. And the female ovipositors are photographed and described. A key to distinguish all known species of the genus is provided. The hind wing venation of the new species are the same as the T. suphanburia and T. sinuata, and not common in the tribe Erythroneurini.     

A new genus and species of the cockroach family Blattulidae from Lebanese amber (Dictyoptera,Blattina)

A new genus and species of the cockroach family Blattulidae, Ocelloblattula ponomarenkoi gen. et sp. nov., are described from the Early Cretaceous Lebanese amber. In the wing venation, the new genus is extremely similar to the Jurassic genus Blattula Handlirsch, differing from the latter in a number of characters in its body structure. This find reveals much about the body structure of the extinct family Blattulidae, which is related to ancestors of the suborders Mantina and Blattina.     

Discovery of the genus Crinoedischia in the Middle Permian of France (Insecta: Orthoptera)

New orthopteran Crinoedischia lapeyriei sp. nov. is described and illustrated from the Middle Permian (Guadalupian) of Lodève Basin in southern France. The new species is separated from the Lower Permian (Artinskian) Crinoedischia species by differences in wing venation pattern such as long ScP with anterior branches, RA with prominent distal branches, and MA ending with three branches. The significance and parallels of entomofaunas between Salagou and Wellington formations (Lodève, Elmo, Midco) are briefly discussed.     

Two new species of Archaeohelorus (Hymenoptera,Proctotrupoidea, Heloridae) from the Middle Jurassic of China

Two new fossil species, Archaeohelorus polyneurus sp. n. and A. tensus sp. n., assigned to the genus Archaeohelorus Shih, Feng & Ren, 2011 of Heloridae (Hymenoptera), are reported from the late Middle Jurassic, Jiulongshan Formation of Inner Mongolia, China. Based on the well-preserved forewings and hind wings of these specimens, the diagnosis of the Archaeohelorus is emended: forewing 2cu-a intersecting Cu and Rs+M at the same point or postfurcal, and hind wing may have tubular veins C, Sc+R, R, Rs, M+Cu, M and Cu distinct, or simplified venation. The new findings also elucidate the evolutionary trend of forewing and hind wing venation and body size for the Heloridae from the late Middle Jurassic to now.     

The beetle-like Palaeozoic and Mesozoic roachoids of the so-called “umenocoleoid” lineage (Dictyoptera: Ponopterixidae fam. nov.)

The position of Palaeozoic and Mesozoic roachoids and their relationships to crown-group Dictyoptera is an unresolved problem of insect systematics since Hennig's time. This contribution presents new data based on the wing venation of the Early Cretaceous group Cratovitismioidea (so-called “Umenocoleoidea”) supplemented with the discovery of the first Late Palaeozoic representative. As Umenocoleus, type genus of the Umenocoleidae, is considered as a Coleopterida, the roachoids currently included in the Umenocoleidae are transferred to the new family Ponopterixidae, in the new superfamily Cratovitismioidea. Permoponopterix lodevensis, a new ponopterixid genus and species, is described as the oldest representative of this superfamily, from the Middle Permian of the South of France. A new species Ponopterix burkhardi is described from the Lower Cretaceous of Crato Formation in Brazil. Convergent adaptations of tegmina in Coleopterida, Protelytroptera and Cratovitismioidea are discussed.     

A new genus and species of tettigarctid cicada from the early Miocene of New Zealand: Paratettigarcta zealandica (Hemiptera,Auchenorrhyncha, Tettigarctidae)

A new genus and species of primitive cicada (Hemiptera: Tettigarctidae) is described from the early Miocene of southern New Zealand. Paratettigarcta zealandica gen. et sp. n. is the first cicada (Cicadoidea) fossil from New Zealand and exhibits wing venation patterns typical for the subfamily Tettigarctinae. It differs from other fossil taxa and the extant genus Tettigarcta in the early divergence of CuA2 from the nodal line in the forewing, its parallel-sided subcostal cell, the early bifurcation of vein M and long apical cells of the hindwing, and in wing pigmentation patterns.     

The oldest bee fly in the French Paleocene (Diptera: Bombyliidae)

Paleolomatia menatensis gen. and sp. n., oldest Bombyliidae sensu stricto, is described from the Paleocene of Menat (France). The new genus, based on the wing venation, is attributed to the rather ‘derived’ subfamily Lomatiinae, strongly supporting a Late Cretaceous age for the diversification of the pollinator bee flies, in relation with the floristic changes and the angiosperm radiation that occurred at the same time.     

A new species of the genus <Emphasis Type="Italic">Stathmopoda</Emphasis> (Lepidoptera: Stathmopodidae) closely related to the persimmon pest, <Emphasis Type="Italic">S. masinissa</Emphasis>, from Ryukyu Islands,Japan

A new species of the genus Stathmopoda from the Ryukyu Islands, Japan, is described. The new species is associated with Diospyros maritima Blume and is very similar to S. masinissa Meyrick, which is a notorious pest of persimmon, D. kaki L., in Japan and Korea. The adult external features, wing venation, and male and female genitalia of both species are illustrated and compared in detail.     

A new rhopalosomatid wasps in the Paleocene of France (Hymenoptera)

The third oldest Rhopalosomatidae, Paleorhopalosoma menatensis n. gen., n. sp., is described from the Paleocene of Menat (France). In addition to the characters unique to rhopalosomatids the plesiomorphic complete wing venation together with plesiomorphic shape of metasoma show that this genus is very likely a more ‘basal’ clade of this family, together with the Early Cretaceous genus Mesorhopalosoma.     

Miocene ants (Hymenoptera,Formicidae) from Crimea

Ant imprints, new and previously known, from Middle Miocene deposits near Kerch (Crimean Peninsula, Russia) are described or redescribed. A new Myrmicinae species, Solenopsis atavinus sp. nov., is described based on a wingless female. A winged female similar to the earlier described Dolichoderus tauricus Dlussky, 1981 is found: due to the excellent preservation of this specimen, the new specimen and the holotype of D. tauricus are redescribed and can be reclassified as Ponerites tauricus (Dlussky, 1981), comb. nov. Oecophylla taurica sp. nov. is described based on a partly preserved imprint of a female thorax with a forewing, the venation of which allows it to be positively identified as a weaver ant. Two other species are transferred from the formal genus Camponotites to Oecophylla based on forewing venation: O. kraussei (Dlussky et Rasnitsyn, 1999), comb. nov. (Early Eocene, United States) and O. macroptera (Dlussky, 1981), comb. nov. (Middle Miocene, Stavropol, Russia). One of the studied forewing imprints is similar in venation to Paraphaenogaster microphthalmus Dlussky, 1981, described from the Middle Miocene of Vishnevaya Balka (Stavropol province, Russia) and so is attributed to this species. Dolichoderus tavridus sp. nov. is described based on a forewing.     

First record and taxonomic description of the genus Thysanoplusia (Fabricius) (Lepidoptera: Noctuidae: Plusiinae) from Pakistan

Species belonging to genus, Thysanoplusia Fabricius of the subfamily Plusiinae which are polyphagous in nature and pest of vegetables, foods, legumes, fodder, fruits, ornamental plants and cotton crops. Samples were collected from different localities of district Bahawalpur. For collection, comprehensive and comparative surveys were carried out during 2017–18 on taxonomic account of species of the genus Thysanoplusia Fabricius and resulted identified one species Thysanoplusia orichalcea (Fabricius) first time from Pakistan. Morphological characters viz., vertex, frons, labial palpi, antennae, compound eyes, ocelli, proboscis, wing venation, male and female genital characteristics were used for the identification and classification. Dichotomous keys and photographs are also provided. There is hardly any substantial research work on taxonomic studies of subfamily Plusiinae Pakistan. So to fill this gap the present proposal was designed to study the diversity of Noctuid moths from Pakistan and very fruitful results have been obtained.     

First report of the family Palaeosetidae (Lepidoptera: Hepialoidea) from mainland China,with description of a new Ogygioses species and its phylogenetic position within the genus

The family Palaeosetidae is newly recorded from mainland China. A new species of the genus Ogygioses, O. maoershana sp. nov., is described from Guangxi Province, China. The adults, male and female genitalia, venation, head and legs structures are illustrated. Some ecological information and a key to all species of the genus are also provided. The phylogenetic position of the new species in the genus Ogygioses is inferred as intermediate between primitive O. luangensis Kristensen and other four derived species based on morphological characters and DNA barcodes sequences.www.zoobank.org/urn:lsid:zoobank.org:pub:D1FD6818-500A-4AE1-9F0B-13CB5A6E1ED4.     

A new species of Halolaguna Gozmány from Thailand (Lepidoptera: Lecithoceridae)

A new species of the genus Halolaguna Gozmány, H. sanmaru sp. nov., is described from Thailand. The adult, hind tibia, male genitalia, and venation of the new species are illustrated and compared with those of H. oncopteryx (Wu).     

A new species of the bee genus Ctenoplectrella in middle Eocene Baltic amber (Hymenoptera, Megachilidae)

A new species of the extinct bee genus Ctenoplectrella Cockerell (Megachilinae: Ctenoplectrellini) is described and figured from two females preserved in middle Eocene (Lutetian) Baltic amber. Ctenoplectrella phaetonsp. n. is distinguished from its congeners on the basis of its body proportions, integumental sculpturing, wing venation, and pubescence, and is one of the more distinctive members of the genus. A revised key to the species of Ctenoplectrella is provided.     

Two new species of Antispila Hübner (Lepidoptera: Adeloidea: Heliozelidae) associated with Hydrangea (Hydrangeaceae) from China

The genus Antispila Hübner, [1825] is poorly recorded from China. Two new Antispila species, A. longcangensis, sp. nov. and A. emeishanensis, sp. nov. feeding on Hydrangeaceae, are described from Sichuan, China. The adult male and female, genitalia and venation of both species are illustrated. Diagnostic characters of these species are compared with other described Antispila species from China. DNA barcodes sequences of the two new species are also provided.     

Preliminary phylogenetic analysis of the Protanisoptera (Insecta: Odonatoptera)

The Permian suborder Protanisoptera (Insecta: Odonatoptera) is revised and a new phylogenetic hypothesis proposed after analyses based on wing venation and different outgroups. After our study the families Camptotaxineuridae and Kaltanoneuridae are excluded from the Protanisoptera. After a new phylogenetic analysis, the family Permaeschnidae is redefined and the families Pholidoptilidae, Polytaxineuridae, Callimokaltaniidae and Hemizygopteridae are restored, as already proposed for the latter three families by Bechly (1996). The new genus Proditaxineura is described. The genus Gondvanoptilon RÖSLER et al., 1981 is excluded from the Meganisoptera: Erasipteridae and re-included in the Permaeschnidae, as already proposed by Bechly (1998). Permaeschna proxima MARTYNOV, 1931 is considered as a junior synonym of Permaeschna dolloi MARTYNOV, 1931. Pholidoptilon camense ZALESSKY, 1931 is excluded from Permaeschna MARTYNOV, 1931 and the genus Pholidoptilon [Zalessky, 1931a] and [Zalessky, 1931b] is restored. Ditaxineurella stigmalisMARTYNOV, 1940 is excluded from the Hemizygopteridae and considered as a Protanisoptera Incertae sedis.     

中生代线蛉科化石(昆虫纲,脉翅目)

线蛉科是脉翅目中的一个灭绝科,仅发现于中国中侏罗统和蒙古晚侏罗统地层中。线蛉科昆虫脉序独特,为其分类地位的确定以及整个脉翅目翅脉演化的研究提供了重要的线索;此外,线蛉科昆虫前后翅上均具有条带状的斑纹,初步推测这种斑纹在其生存繁衍过程中发挥了积极作用,也可能是导致其灭绝的最直接因素之一。目前线蛉科已发表3属9种,本文对其进行了简要的回顾,并编制了线蛉科化石种检索表,对当时的古环境特征进行了简单推测,初步探讨了翅斑在其生存和灭绝中的作用。