Structure and Secretion of the Graptolite Prosicula, and its Application for Biostratigraphical and Evolutionary Studies

Studies on graptolite taxonomy and phylogeny in recent years have placed great emphasis on the proximal development of the rhabdosome, particularly the presence or absence of a virgella and early thecal growth patterns. As the prosicula was the earliest part of the graptolite skeleton to be secreted, it may also reveal fundamental information about evolutionary relationships within the Graptoloidea. The prosiculae from a variety of Ordovician taxa ranging in age from Tremadoc to Caradoc have been examined using a combination of light microscopy, SEM and TEM. Parameters investigated include the overall morphology, transition into the nema, pattern of longitudinal ridges and spiral line. Taxa show a change from early Tremadoc graptoloids which have a low diaphragm, prominent spiral line and lack longitudinal ridges, through late Tremadoc and early Arenig taxa which have longitudinal cortical bandages or spiralled, paired longitudinal ridges, into later Arenig and Llanvirn forms which have simple longitudinal ridges and indistinct spiral line and diaphragm. With additional work at higher stratigraphical levels, graptolite prosiculae may prove to be useful biostratigraphically when more complete material is absent, such as in palynomorph preparations from subsurface studies.     

华南特马豆克期笔石地层分带及全球对比

特马豆克阶是奥陶系底部第一个阶,笔石是特马豆克阶高分辨率地层划分与对比的重要化石类群。江南斜坡带是我国早奥陶世特马豆克期漂浮笔石分异度和丰度最高的相区之一,位于该区的湖南益阳南坝剖面,发育有完整的上特马豆克阶笔石地层,特马豆克阶-弗洛阶界线附近地层连续,上特马豆克阶笔石地层研究已取得较大进展,但下特马豆克阶地层缺乏系统研究。近年来,通过对该剖面笔石标本的不间断采集,新识别出下特马豆克阶笔石带Rhabdinopora flabelliformis parabola带。到目前为止,湖南益阳南坝剖面特马豆克阶可以识别出5个笔石带,自下而上依次为:Rhabdinopora flabelliformis parabola带、Adelograptus tenellus带、Aorograptus victoriae带、Araneograptus murrayi带以及Hunnegraptus copiosus带。基于目前已识别出的笔石带,参考国内外同期笔石地层资料,本文详细讨论华南特马豆克期笔石带序列,并与国内外同期地层进行精确对比。     

Billingen (Lower Arenig/Lower Ordovician) acritarchs from the East European Platform and their palaeobiogeographic significance

Billingen (Lower Arenig/Lower Ordovician) sediments of the St. Petersburg region, northwest Russia and the Leba area, northern Poland of the East European Craton yield acritarch assemblages, which are largely homogenous though displaying minor compositional differences that probably reflect a gradient from inner to outer shelf environments. Comparison with coeval acritarch microflora from the Yangtze Platform, South China, shows an overall similarity between Baltoscandian and South Chinese phytoplankton. The widespread uniformity in the fossil microphytoplankton may be related to the extensive global 'evae' sea-level transgression, which characterized the Billingen time. This suggests that during the Tremadoc through early Arenig times, acritarch assemblages displayed essentially an undifferentiated cold-water and oceanic character along the whole margin of Perigondwana in the South, as well as on the South Chinese and Baltic platforms, at middle latitudes (Mediterranean oceanic Realm). Despite this overall similarity, however, some typical taxa of the high-latitude Mediterranean Province (Arbusculidium, Coryphidium and Striatotheca) occur in South China, but are absent in Baltica. This discrepancy is explained as caused by differences in climatic and physiographic conditions that prevailed at the two palaeocontinents at this time. The inferred pattern of oceanic circulation during the Lower Ordovician is consistent with the palynological evidence of a prevailing warmer climate in Baltica than in South China, although the two palaeocontinents occupied the same palaeolatitudinal position.     

贵州三都地区阿伦尼克期疑源类的发现及其意义

本文首报报道了发现贵州三都地区同高组下段的疑源类化石,属种丰富,保存完好,它们多见于阿伦尼克期至兰维尔期,其中有些分子仅限一阿伦尼克期,根据组合中出现较多的Ｓｔｒｉａｔｏｔｈｅｃａ,Ｃｏｒｙｐｈｉｄｉｕｍ和Ａｒｂｕｓｃｕｌｉｄｉｕｍ等,认为本区可能属地中海疑源类生物区,详细研究这些化石对认识疑源类组合与笔石带的关系有重要意义。     

湖北宜昌奥陶系庙坡组疑源类

湖北宜昌地区庙坡组是达瑞威尔阶/桑比阶界线附近的地层,该段地层产出分异度较高,丰度适中的疑源类组合。该组合包括16属,28种,其中7个未定命名种,可与国内、外同期疑源类组合进行对比。庙坡组疑源类组合既产出晚奥陶世特征分子,也产出阿伦尼格期(弗洛阶上部—达瑞威尔阶底部)的特征分子,显示出一定的过渡特色。组合以Baltisphaeridium(17%—52%),Leiosphaeridia(6%—78%)占优势;反映了其沉积环境为离岸较远的外陆棚环境。     

Cambrian-Ordovician lingulate brachiopods from Scandinavia, Kazakhstan, and South Ural Mountains

Lingulate brachiopods are described from the Upper Cambrian - Lower Ordovician (Tremadoc-Arenig) of Scandinavia (Sweden, Denmark, and Norway), South Ural Mountains, northeastern Central Kazakhstan, and the southern Kendyktas Range in southern Kazakhstan. The faunas comprise a total of 56 species of which 20 are new these are assigned to 40 genera, of which the lingulids Agalatassia and Keskentassia , the siphonotretid Siphonotrerella. and the acrotretids Galinella, Longipegma, Ottenbyella, Akmolina, Mamatia, Sasyksoria , and Otariella are new. The new Subfamily Elliptoglossinae is proposed. The poorly known Cambrianardovician stratigraphy of the South Urals. northeastern Central Kazakhstan, and the southern Kendyktas Range is reviewed. Many sequences in these areas that were previously referred to the Upper Cambrian and Tremadoc can now be correlated with the lower Arenig Hunneberg Stage in Baltoscandia. Three main types of faunal assemblages can be distinguished: (1) the Broeggeria assemblage; (2) several microbrachiopod assemblages; and (3) the Leptembolon-Thysanotos assemblage. The Broeggeria assemblage is distributed world-wide in the Tremadoc of the southern Kendyktas Range, Scandinavia, Belgium, Great Britain, Canada, and Argentina, while the Leptembolon-Thysanotos assemblage is confined to the Arenig of an area surrounding the East European platform, including northern Estonia, Poland, Germany, Bohemia, Serbia, and the South Urals. The microbrachiopod assemblages are known mainly from the Upper Cambrian - Arenig of Scandinavia, South Ural Mountains, northeastern Central Kazakhstan, and the southern Kendyktas Range.     

Upper Tremadoc–Lower Arenig? Anisograptid–Dichograptid fauna from the Cabitza Formation (Lower Ordovician,SW Sardinia,Italy)

The discovery of the anisograptid graptolite Araneograptus murrayi (Hall, 1865), the dichograptids Clonograptus (Clonograptus) cf. rigidus (Hall, 1858) and Clonograptus (Clonograptus) cf. multiplex (Nicholson, 1868) and of two unassigned species of Didymograptus in the Late Tremadoc–Early Arenig? sediments of the Fluminese area (SW Sardinia) adds new data on the biostratigraphy and palaeoecological setting of the Cabitza Formation, as well as on the paleobiogeographical distribution of these taxa.     

Late Tremadoc to early Arenig graptolite faunas of southern Bolivia and their implications for a worldwide biozonation

The graptolite faunal succession of the late Tremadoc to early Arenig in southern Bolivia is established from the Cieneguillas and Culpina sections and compared with faunas from other regions. The succession yields important data for the discussion on the proposed GSSP at the base of the Tetragraptus approximatus Zone and the completeness of the successions in eastern North America and Scandinavia. The graptolite faunas of the Aorograptus victoriae, Kiaerograptus supremus, Araneograptus murrayi, Hunnegraptus copiosus and Tetragraptus phyllograptoides zones are reviewed and the faunal associations discussed. The Kiaerograptus supremus Zone is regarded as a local Scandinavian biozone.     

浙江常山黄泥塘全球层型剖面的中奥陶世疑源类

浙江常山黄泥塘剖面在1996-1997年被国际奥陶系分会和国际地质科学联合会(IUGS)确立为全球中奥陶统达瑞威尔阶(Dariwillian)层型剖面,从该剖面宁国组Azygograptus suectcus-Exigraptus clavus笔石带至Undulograptus sinicus笔石亚带下部的页岩及灰岩中获得丰富的,保存不很好的疑源类化石,它们包含26个形态属,41个已知种和12个未定名种.该宁国组疑源类组合极少有环冈瓦纳古陆的“冷水“类型,根据疑源类类型在地层柱的首次出现,可划分两个与Azygograptus suecicus和Undulograptus austrodentatus笔石带相吻合的疑源类生物地层单元,结合中国南方在奥陶纪的古地理位置加以讨论,当前疑源类组合表明,中奥陶世环冈瓦纳古陆的“冷水“洋流至下扬子区已很微弱.     

Early Paleozoic (Late Cambrian-Early Ordovician) acritarchs from the metasedimentary Baden-Baden-Gaggenau zone (Schwarzwald, SW Germany)

The metasediments of the low-grade metamorphosed Baden-Baden-Gaggenau zone of the northern Schwarzwald (southwestern Germany) have been analyzed palynologically. From 133 samples representing different metasedimentary units, only three samples of the upper part of the Traischbach Serie provide extremely poorly-preserved palynomorphs. The assemblage consists of the galeate acritarch genera Caldariola, Cymatiogaleaand Stelliferidium, as well as specimens of diacromorph and polygonomorph acritarchs. Although determinations are difficult at the generic level and essentially impossible at the specific level, the assemblage can be attributed to an interval between the Late Cambrian and Early Ordovician. This is the first biostratigraphical age assignment for the metasediments of the Baden-Baden-Gaggenau zone, which can possibly be correlated with the Villé Unit of the northern Vosges Mountains (eastern France).     

Stratigraphic distribution of selected acritarchs in the Ordovician subsurface,Canning Basin,Western Australia

Palynological investigation of 80 core samples from four boreholes, drilled through the Lower to Middle Ordovician marine sections of the central-northeastern Canning Basin, northwestern Australia, reveals diverse acritarch associations including a number of stratigraphically significant species. The stratigraphic succession studied, embracing the Willara Formation and conformably overlying Goldwyer Formation, has been independently dated as early Arenig to Llanvirn by conodont faunas (Oepikodus communis through Phragmodus-Plectodina zonal interval). Nine morphologically distinctive acritarch species have relatively narrow vertical ranges within the study interval and likely constitute serviceable palynostratigraphic indices for the Ordovician in and possibly beyond the Western Australian study area. Three new species, Comasphaeridium setaricum, Polyancistrodorus kunzeanensis, and Stelomorpha calix, are established; and one new combination, Baltisphaeridium variocavatum (Playford and Martin, 1984), is proposed. Other stratigraphically significant species include Petaloferidium comptum Playford and Martin, 1984 and Peteinosphaeridium sp. cf. P. exornatum Tongiorgi et al., 1995; together with Dorsennidium symmetricum (Lu, 1987) Sarjeant and Stancliffe, 1994, Peteinosphaeridium coronula Yin et al., 1998, and Striatotheca rarirrugulata (Cramer et al., 1974) Eisenack et al., 1976, which are found for the first time in Australia. The vertical distribution, and hence the stratigraphic utility, of each of the above species is documented through the studied sections.     

Upper Ordovician microphytoplankton of the Bill's Creek Shale and Stonington Formation,Upper Peninsula of Michigan,U.S.A.: Biostratigraphy and paleogeographic significance

An abundant, diverse, and well-preserved organic-walled microphytoplankton assemblage is described from the Upper Ordovician Bill's Creek Shale and the lower Stonington Formation (Bay de Noc Member) in the Upper Peninsula of Michigan, U.S.A. Based on graptolite and conodont evidence, the Bill's Creek Shale and Stonington Formation are Richmondian (=Ashgill) in age. The assemblage is dominated by acritarchs, which comprise 29 species (including the enigmatic palynomorph Gloeocapsomorpha prisca) assigned to 20 genera. The prasinophyte phycomata are represented by undifferentiated species of Leiosphaeridia and Tasmanites. In addition, chitinozoans are abundant, and scolecodonts and graptolite fragments are common. Paleontologic-palynologic and sedimentologic evidence indicates that the Bill's Creek Shale was deposited in a low-energy, shallow, nearshore marine environment. The overlying Bay de Noc Member of the Stonington Formation also accumulated in a low-energy, normal marine environment, but in a more offshore, somewhat deeper water setting. Both formations experienced minor transgressive and regressive episodes as indicated by fluctuations in the composition of the palynoflora. The combined Bill's Creek/Stonington acritarch assemblage closely resembles those described from the Richmondian-aged Maquoketa Shale (Missouri and Kansas), Sylvan Shale (Oklahoma), and Vauréal Formation (Anticosti Island, Québec, Canada). The overall composition of the acritarch assemblage from these four formations reflects a distinctive, recognizably Laurentian character. Nonetheless, many of the Bill's Creek/Stonington acritarchs have been reported from Upper Ordovician localities elsewhere, providing additional evidence for Late Ordovician cosmopolitanism of the marine microphytoplankton community. Additionally, the restricted stratigraphic range of many of the taxa further enhances their biostratigraphic application, both regionally and globally, and reaffirms the Richmondian (=Ashgill) age of the Bill's Creek Shale and Stonington Formation.     

湖北宜昌地区下-中奥陶统界线疑源类生物地层学意义

根据牙形类Baltoniodus? triangularis的首现,中奥陶统和大坪阶底界已界定于湖北宜昌黄花场剖面大湾组底界之上10.57m.对黄花场剖面以及邻近的大坪剖面大湾组疑源类研究表明,某些疑源类属种具有生物地层学意义,其首现可用于识别奥陶系下/中统界线.     

BASAL CAMBRIAN MICROFOSSILS FROM THE YURTUS AND XISHANBLAQ FORMATIONS (TARIM, NORTH-WEST CHINA): SYSTEMATIC REVISION AND BIOSTRATIGRAPHIC CORRELATION OF MICRHYSTRIDIUM-LIKE ACRITARCHS

Abstract:  Micrhystridium -like acritarchs are widely distributed in basal Cambrian cherts and phosphorites in South China. This paper describes similar acritarchs from the basal Cambrian Yurtus and Xishanblaq formations in Tarim, north-west China. The taxonomy of these acritarchs is revised. The basal Cambrian acritarch assemblage in Tarim and South China is characterized by three genera: Asteridium Moczydłowska, Heliosphaeridium Moczydłowska and Comasphaeridium Staplin, Jansonius and Pocock. This assemblage is named the Asteridium - Heliosphaeridium - Comasphaeridium (AHC) acritarch assemblage. In both South China and Tarim, the AHC acritarch assemblage is associated with the tubular microfossil Megathrix longus Yin L. and the small shelly fossil Kaiyangites novilis Qian and Yin G. This assemblage also occurs in the Lower Tal Formation in the Lesser Himalaya. Correlation with small shelly fossil (SSF) assemblages indicates that the AHC assemblage is restricted to the Meishucunian Stage, and possibly to the lower Meishucunian ( Anabarites trisulcatus - Protohertzina anabarica and Siphogonuchites triangularis - Paragloborilus subglobosus SSF assemblages). The AHC assemblage is broadly similar to the Asteridium tornatum - Comasphaeridium velvetum (acritarch) Zone in the East European Platform, which is considered to be Nemakit-Daldynian (and possibly Tommotian) in age.     

河南汝州罗圈村剖面汝阳群白草坪组的微体化石

河南汝州罗圈村白草坪组微体化石丰度大,但分异度低,仅包括具细网状纹饰的Dictyosphaera delica-ta和具刺饰的Shui yousphaeridium pilatumsp.nov.以及少量的带藻Taeniatum sp.。虽然化石组成单调,但总体上与山西永济水幽沟剖面和河南渑池麦尽沟剖面白草坪组的疑源类组合面貌一致。目前,此套化石组合的地质时代有中元古代和新元古代早期两种不同的认识,通过对化石形态特征分析以及与苏格兰西北部早新元古代Torri-don群中发现的一个微体化石组合的对比,文中暂将汝阳群地质时代置于新元古代早期。     

Aeronian (Llandovery, Lower Silurian) palynomorphs and graptolites from the Lipeón Formation, Eastern Cordillera, north-west Argentina

The lower levels of the Lipeón Formation, in the Eastern Cordillera, north-west Argentina, yield a marine-dominated palynomorph assemblage, together with graptolites of mid to late mid Llandovery age (Demirastrites convolutus and probably Stimulograptus sedgwickii zones). The palynomorph assemblage is dominated by acritarchs, but also contains algae and terrestrial cryptospores. Crassiangulina variacornuta, considered a potentially good global biostratigraphical marker for the Upper Llandovery is recovered for the first time from the Silurian of Argentina. The occurrence of this species in strata not younger than late Aeronian, and independently dated by graptolites, indicates an early first appearance for Crassiangulina variacornuta, in the Lipeón Formation, below the Aeronian/Telychian boundary. The lower part of the unit corresponds to a quiet marine environment; thus supporting that Crassiangulina variacornuta is a facies-sensitive acritarch.     

The acritarchs of the South Chinese Azygograptus suecicus graptolite Biozone and their bearing on the definition of the Lower–Middle Ordovician boundary

In southern China, well preserved acritarch assemblages have been recovered from numerous sections in the Yangtze Platform and the Jiangshan–Changshan–Yushan (JCY) area crossing the interval where the Lower–Middle Ordovician boundary should be defined, i.e., roughly at the base of the Chinese Azygograptus suecicus graptolite biozone. The acritarch taxa Aureotesta clathrata and Arbusculidium filamentosum first appear below the suecicus zone, while the genus Ampullula and the species Barakella felix have their First Appearance Datum (FAD) in the suecicus zone. These latter taxa, that are also present in Baltica and peri-Gondwana, respectively, thus probably indicate the base of the Middle Ordovician. To cite this article: J. Li et al., C. R. Palevol 1 (2002) 75–81.