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1.
The study purpose was to investigate the reproductive biology, growth and length‐weight of the Turkmenian crested loach, Metaschistura cristata, in the Radkan River of northeastern Iran. Age and growth are described for 1029 specimens from January 2009 to December 2010. The sex ratio was 1:1. Maximum age, based on opercula readings, was 6+ years for both sexes. Specimens ranged in size from 24 to 98 mm total length and weighed from 0.08 to 7.32 g. The length‐weight relationships were described as W = 0.005166 TL3.225 (R2 = 0.97) for males and W = 0.006192 TL3.125 (R2 = 0.97) for females. Growth was expressed in length and the von Bertalanffy growth parameters were estimated as L = 354.9 mm, k = 0.0038, t0 = ?26.82 for males and L = 339.0 mm, k = 0.0043, t0 = ?24.88 for females. Growth performance indexes were also estimated as Φ′ = 6.17 for males and Φ′ = 6.20 for females. The gonadosomatic index showed that peak reproduction occurred during April and June, with highest average values of 2.1% for males and 25.3% for females in May. Oocyte diameter ranged from 0.09 to 1.58 mm, with a mean value of 0.54 ± 0.42 mm.  相似文献   

2.
Age composition, growth, and reproductive biology of the non‐native Indian glassy fish Parambassis ranga (Hamilton, 1822) were surveyed in the Haebaru Reservoir on Okinawa‐jima Island, southern Japan. Standard lengths (SLs) of males and females ranged from 19.7 to 44.0 mm and from 19.2 to 52.4 mm, respectively. The overall sex ratio was significantly female‐biased, with the monthly percentage of females ranging from 71.4 to 100.0. Marginal growth analysis of sectioned otoliths revealed opaque zones formed annually from November to May. Observed age ranged from 0 to 3 years for both sexes, although the 1‐year age class comprised the majority of the sampled population. Von Bertalanffy growth parameters were L = 48.8 mm (SL), K = 0.43 year?1, and t0 = ?1.47 years for males, and L = 43.7 mm, K =0.72 year?1, and t0 = ?1.29 years for females. Length at 50% maturity was estimated to be 25.8 mm SL, and maturation was within 1 year after hatching. The main spawning season of P. ranga was estimated to occur from February to October, peaking in April.  相似文献   

3.
The life‐history traits of Aphanius danfordii in Hirfanl? Reservoir were studied on the basis of 2252 specimens caught between April 2008 and April 2009. Maximum age was determined as five years for both sexes. The overall sex ratio of males to females was 1 : 1.21; however, this ratio varied by age and season. Mean total length at age data derived from scale readings were used to estimate growth. The von Bertalanffy growth parameters were: L = 126.63 mm, = ?0.09, t0 = 2.35 for females; and L = 61.2 mm, = ?0.19, t0 = 2.76 for males. The gonadosomatic index (GSI) values indicated spawning to be from May to September. Absolute fecundity varied from a minimum of 143 eggs for age one to a maximum of 698 eggs for age five. Relationships between fecundity–length and fecundity–weight were described by the equations: = 0.0002 TL3.3222 and = 36.032 W1.0053, respectively. The equation of the relation between absolute fecundity and age was = 25.372 t1.2343.  相似文献   

4.
Moroccan white seabream Diplodus sargus cadenati (n = 603) were caught off the Canary Islands from April 2000 to March 2001. Total length ranged from 46 to 404 mm. The subspecies was characterized as being dygynous with partial protandry. Overall ratio of males to females was 1 : 2.9. The reproductive season extended from December to May, with a peak in spawning activity in January–February. Fifty per cent maturity was reached at 201 mm total length in males and 216 mm in females. The length–weight relationship for all individuals was described by the following parameters: a = 0.000023, and b = 2.96, when length is given in millimeters and weight in grams. Fish of 0–12 years in age were found. The von Bertalanffy growth parameters for the entire population were: L = 467 mm, k = 0.143 year?1 and t0 = ?2.14 year. Growth parameters differed between males and females. For all fish, instantaneous rates of mortality were Z = 0.68 year?1, M = 0.31 ± 0.6 year?1 and F = 0.37 ± 0.6 year?1; the exploitation ratio was E = 0.54 ± 0.9. Length at first capture for all individuals was 173 mm. The stock exploited above is an assumed optimum.  相似文献   

5.
Age, growth, and age at sexual maturity of the polkadot skate Dipturus chinensis, in the northern East China Sea were determined for a total of 614 specimens collected from April 2009 to December 2014. Vertebral centrum analysis was used to calculate the age of the skates. Annual band deposition was determined by marginal increment analysis. The von Bertalanffy growth model was fitted to the observed length‐at‐age data for each sex (males, L = 76.8, k = 0.109, t0 = ?1.28; females, L = 83.1, k = 0.103, t0 = ?1.20). Growth patterns of females and males were similar until the age of 6; thereafter, females grew larger than males. Maximum age recorded was 13 years for males and 15 years for females. Age at 50% sexual maturity was 8.22 years for males and 9.39 years for females. These results indicate that Dchinensis is slow growing, relatively long‐lived, and late maturing, and therefore vulnerable to exploitation.  相似文献   

6.
Under‐or non‐exploited areas are useful to aid in evaluating the potential productivity of fish stocks for sustainable fisheries. The Gulf of Castellammare (NW Sicily), where trawling has been banned since 1990, is a good site to study the biology and dynamics of low impacted fish populations. A total of 661 (595 female and 66 male) specimens of red mullet Mullus barbatus (95–245 mm total length, TL) obtained by monthly sampling from trammel net artisanal fishery, was collected in the Gulf of Castellammare from April 2006 to June 2007. Mature females occurred from April to September, with a peak in May. The mean Gonado‐somatic index (GSI) also showed a May peak for both sexes. From Sagittae (642) readings the age structure ranged from age class I to VII in females and I to V in males. Female growth parameters, estimated according to the classic von Bertalanffy model, were: L = 221.2 ± 11.51 mm standard error (SE), ky?1 = 0.38 ± 0.09 SE, t0y = ?0.94 ± 0.38 SE. The growth performance index (ø’ = 2.27) was included in the range of values obtained by hard structure readings in the Central Mediterranean. Natural mortality (My?1) of females estimated by different methods ranged between 0.62 and 0.87 (mean value = 0.71 ± 0.06 SE).  相似文献   

7.
Life history parameters associated with reproductive biology, age, and growth of the convict cichlid (also known as the zebra cichlid) Amatitlania nigrofasciata, which was introduced into the Haebaru Reservoir on Okinawa-jima Island, were estimated using 437 specimens that ranged from 13.7 to 82.9 mm standard length (SL). Lengths of females at first maturity (SL) and 50% maturity (L 50) were estimated to be 32.2 and 37.3 mm SL, respectively. The spawning period continued throughout the year, with a peak spawning cycle from March to May 2006–2007. Observations of postovulatory follicles and tertiary yolk stage oocytes indicate that convict cichlids spawn multiple times within a year. Female cichlids that hatched during the peak spawning seasons matured after October of the same year. Batch fecundity of females (32.2–61.2 mm SL) ranged from 65 to 345 (mean ± SD = 155 ± 63). Opaque zones along the outer margins of otoliths formed annually. The maximum age of male and female cichlids was 3 years. The von Bertalanffy growth formulae (VBGF) were expressed as Lt = 57.4( 1 - e - 0.78( t + 0.91 ) ) {{\hbox{L}}_{\rm{t}}}{ = 57}{.4}\left( {1 - {e^{ - 0.78\left( {t + 0.91} \right)}}} \right) for females and Lt = 69.5( 1 - e - 1.07( t + 0.24 ) ) {{\hbox{L}}_{\rm{t}}}{ = 69}{.5}\left( {1 - {e^{ - 1.07\left( {t + 0.24} \right)}}} \right) for males. Males grew larger than females beginning from the first year. Certain life history characteristics, such as year-round spawning and early maturation, probably contributed to the successful establishment of the convict cichlid, and this species in particular is thought to adapt and become established quickly upon introduction to freshwater systems on Okinawa-jima Island.  相似文献   

8.
The river goby Glossogobius callidus is native to freshwater and estuarine habitats in South Africa. Individuals [21.1–144.4 mm total length (LT)] were sampled from impoundments in the Sundays River Valley, Eastern Cape, from February 2014 to March 2015. The largest female was 137.2 mm LT, and the largest male was 144.4 mm LT. Length-at-50% maturity was 75.2 ± 2.1 mm LT for males and 76.2 ± 2.0 mm LT for females. Absolute fecundity was 1028.2 ± 131.7 oocytes per fish, and relative fecundity was 50.1 ± 18.1 oocytes per gram. The spawning season extended from October to December. Fish were aged using sectioned sagittal otoliths. The growth zone periodicity was validated using edge analysis. Longevity was more than 7 years for females and more than 6 years for males. Length-at-age was similar for the two sexes and was best described using the von Bertalanffy growth model as Lt = 74.7(1 − e–1.0(t + 0.1)) mm LT for the entire population. Using the population age structure, the mortality rate was estimated at 1.3 per year.  相似文献   

9.
This aim of this paper was the study of the reproductive biology and growth of the sand smelt, Atherina boyeri, in Mellah Lagoon (Algeria). These data are important for the sustainable exploitation of the stocks of this species. Examined was a total of 1402 Atherina boyeri specimens captured monthly from March 2010 to March 2011, in a population with a 3‐year life cycle. Length–weight relationship was estimated as W = 0.0047 L3.077 (r2 = 0.935) for males and W = 0.0047 L3.176 (r2 = 0.935) for females. Using scales, the von Bertalanffy growth function fitted to back‐calculated size‐at‐age data was Lt = 9.49 [1 ? e?0.316 (t + 0.928)] for males, and Lt = 11.67 [1 ? e?0.179 (t + 1.514)] for females; using otoliths this was Lt = 9.68 [1 ? e?0.3 (t + 1.02)] for males, and Lt = 11.93 [1 ? e?0.171 (t + 1.55)] for females. The growth performance index (Φ) indicated that males (Φscales = 3.34, Φotoliths = 3.33) grew at the same rate as females (Φscales = 3.19, Φotoliths = 3.24), with a sex ratio of 1 : 1.6 in favor of females. The reproductive season extended from February to June. Individual length at first sexual maturity was 4.20 cm for 1‐year‐old males and 4.35 cm for 1‐year‐old females.  相似文献   

10.
Reproductive biology of albacore Thunnus alalunga   总被引:1,自引:0,他引:1  
Reproductive variables in albacore Thunnus alalunga were evaluated by gonad histology in samples of 132 males (58–118 cm fork length, LF) and 112 females (59–101 cm LF) that were collected from the western North Pacific Ocean from 2001 to 2006. In the sex ratio examination, males greatly outnumbered females in large adult fish (LF > 100 cm). Thunnus alalunga exhibited a protracted spawning period from March to September in the waters off eastern Taiwan and the Philippines, and the peak spawning activity occurred in March and April. Minimum sizes associated with the classification of mature fish were 78 and 83 cm LF for males and females, respectively. In addition, the largest LF of immature fish were 93 cm for males and 94 cm for females. The spawning frequency estimate in April was 1·7 days. Batch‐fecundity estimates of 21 females (89–99 cm LF) ranged between 0·17 and 1·66 million eggs (mean ±s.d . = 0·94 ± 0·43). The relative fecundity estimates of the 21 females ranged between 9·2 and 92·4 oocytes g?1 body mass (mean ±s.d . = 50·5 ± 22·8). The results presented in this study provide increased information regarding this species' reproductive‐related characteristics than are currently available in stock status determinations.  相似文献   

11.
This study concerns the reproductive strategy of the freshwater fish, Leporinus piau, native to the Brazilian semi‐arid region. The hypothesis of this work predicts that the species would spawn during the rainy season, presenting a seasonal reproductive strategy. The study investigated the structure of total body length and body mass, sex ratio, body size at first sexual maturity, developmental stages of gonads, fecundity, gonadosomatic index (GSI), condition factor (K) and the spawning season of L. piau. Twelve monthly samplings (mean n = 18) were carried out in 2009 and environmental parameters such as rainfall, temperature, pH, electrical conductivity and concentration of dissolved oxygen of water were registered. The sampled population of L. piau (n = 211) showed a slight predominance of males (55%), but with larger and heavier females. First sexual maturation in males occurred earlier (16.5 cm total length) than in females (20.5 cm total length). The species is a total spawner with an average fecundity of 55 000 mature oocytes. Rainfall (225 ± 7.2 mm) and concentration of dissolved oxygen (8.5 ± 0.2 mg L?1) acted as influential factors during the spawning season. The monthly GSI shows that spawning starts when the rainy season is at its maximum and ends synchronously (females r = 0.91; males r = 0.89) with the end of the rainy season. L. piau presents a seasonal reproductive strategy with high reproductive output, providing a population increase and exploitation of the environmental resources.  相似文献   

12.
The age, growth, reproduction, sexual maturity and stomach content of John Dory (Zeus faber), caught in the Bay of Saros (North Aegean Sea) between September 2006 and September 2008, were investigated. The female‐male ratio was 1.6 : 1. The total length of females ranged from 13.8 cm to 52.8 cm, and of males from 12.2 cm to 42.9 cm. The length‐weight relationship was W = 0.0174*L2.936. Age data derived from vertebral centra readings were used to estimate the growth parameters of the von Bertalanffy equation. Growth parameters for females were L = 58.50 cm K = 0.11 year?1, t0 = ?0.99 year and for males L = 45.01 cm, K = 0.13 year?1, t0 = ?1.17 year. Maximum age was 18 years for females and 17 years for males. Length at first maturity for both females and males was 25.4 cm. Monthly values of the gonadosomatic index indicated that spawning occurred mainly in two peaks: one between January and June, and the other from August to September. Stomach content analysis showed the most‐preferred prey to be Pisces (IRI% 97.3).  相似文献   

13.
The aim of this work was to compare age determinations and precision using two deposition structures of trahira Hoplias malabaricus (Bloch, 1794): the scales, which are most frequently used, and otoliths (lapilli). The length‐age relationships were obtained with both structures and compared with results from previous studies. The 163 sets of trahira otoliths (lapilli) and scales were 17–46 cm standard length (SL) from Cayastá (Santa Fe) and Islas Lechiguanas (Entre Ríos), Paraná River, Argentina. Three independent readings of each structure were conducted. An age bias plot was performed to compare age estimations from scales and otoliths. To assess the precision of age determinations using both structures, the percent agreement among readers for both structures and the coefficient of variation were calculated (%CV). The age‐length relationships were plotted and fitted with the von Bertalanffy growth function for both structures and compared with previous works. Age readings recorded for scales were lower than those recorded for otoliths for ages above or equal to 3 years. Percent agreement among readers was higher than 80% for otoliths and less than 65% for scales. The%CV obtained for scales was 20% for young fish and 17% for adults. For otoliths the %CV was 7% for young fish and 3% for adults. The %CV obtained for scales was over the recommended limit (>7.6%). The von Bertalanffy parameters for scales were Linf = 45.80 mm; k = 0.29; t0 = ?1.34 and for otoliths were Linf = 40.76 mm; k = 0.39; t0 = ?1.05. Precision of age estimations assessed from the percent agreement and the coefficient of variation indicates that the scales of the trahira are inappropriate to estimate age in population studies for juvenile and adult specimens.  相似文献   

14.
Age, growth, and reproductive characteristics of blacktail snapper, Lutjanus fulvus, which is a commercial fish species, were investigated using 322 specimens (40–332 mm in fork length) caught around the Yaeyama Islands, Okinawa, Japan. Spawning season was estimated to be from April to October, and spawning was confirmed around the full moon and the last quarter moon. Age determination using sectioned otoliths revealed that ages ranged from 0 to 34 years and the majority was ≥ 3 years. Parameters of the von Bertalanffy growth function were estimated to be L  = 270 mm, k = 0.40, and t 0  = ?0.48 years for females, and 257 mm, 0.44, and ?0.42 years for males. Initial growth was rapid during the first 3 years, attaining over 200 mm for both sexes, and then females grew larger than males. Sizes (ages) at the first sexual maturity were 225 mm (4 years) for females and 207 mm (3 years) for males. The wide range of age composition in catch with majority of ≥3 years old implied that the current fishing effort to harvest was not sufficiently large enough to collapse the stock immediately.  相似文献   

15.
This study describes the reproductive biology of the Chinese minnow Hemiculterella sauvagei. The length‐weight relationship, sex ratio, spawning season, size at first maturity, and fecundity were analyzed based on 685 specimens collected from the Chishui River between July 2011 and July 2012. The relationship between standard length (SL) and body weight (BW) were estimated as BW = 2.14 × 10?4 × SL2.801 (R2 = 0.839; N = 413; P < 0.05) for females and BW = 1.31 × 10?4 × SL3.001 (R= 0.868; N = 272; P < 0.05) for males. The female to male sex ratio, 1.52 : 1, differed significantly from a 1 : 1 ratio. Females predominated in standard lengths >12 cm. Analyses of the monthly variation in the gonadosomatic index (GSI), the monthly proportions of macroscopic gonadal maturity, and the size distribution of oocytes consistently suggested a prolonged spawning season of H. sauvagei from March to August, with a peak from April to May. Logistic curves describing the relationship between proportion of maturity (Pr) at each length interval and standard length were estimated as Pr = 1/(1 + e26.867–0.306SL) (R= 0.999; N = 413; P < 0.05) for females and Pr = 1/(1 + e10.522–0.142SL) (R= 0.999; N = 272; P < 0.05) for males. Size at first maturity was estimated as 7.4 cm for males and 8.8 cm SL for females. Absolute fecundity varied from 563 to 5052, with a mean of 2413 ± 874 oocytes per ovary. The relative fecundity was estimated to be 41–299, with a mean of 171 ± 55 oocytes per ovary. The present study provides useful information for fishery management and resources conservation.  相似文献   

16.
The population parameters of stargazer (Uranoscopus scaber Linnaeus, 1758) were studied regarding age composition, sex ratio, growth, survival and mortality rates, and the exploitation rate in the southeastern Black Sea near the coast of Turkey during the 2011–2012 fishing season. According to ageing analysis of the samples, age group 1 is the most abundant (47.76%), followed by age 2 (41.04%), age 3 (9.33%), age 0 (1.12%) and age 4 (0.75%). Mean totals as well as total lengths and weights for males and females were 14.5 ± 0.22 cm and 12.9 ± 0.19 g, 16.6 ± 0.24 cm and 62.1 ± 2.76 g, and 37.2 ± 1.92 cm and 87.1 ± 3.88 g, respectively. The mean condition factor was K = 0.0167, while the sex ratio was 53.98% female, 38.75% male and 7.27% immature. Length‐weight, age‐length and age‐weight equations were W = 0.014 × L3.059, L(t)=44.5*[1?e?0.148*(t +1.242)] and W(t)=1544.4*[1?e?0.148*(t + 1.242)]3.059, respectively. The computed survival rate (S), instantaneous total mortality rate (Z), annual mortality rate (A), natural mortality rate (M) and fishing mortality rate (F) were S = 28.94%, Z = 1.24 year?1, A = 71.06%, M = 0.26 year?1 and F = 0.98 year?1, respectively. The exploitation rate was 0.79, which is above the optimum exploitation levels.  相似文献   

17.
Several environmental parameters related to the spawning season of the freshwater fish, Psectrogaster rhomboides were identified. The hypothesis predicts synchrony of the spawning season with the rainy period, presenting a seasonal reproductive strategy. This study investigated the total body length and body mass, sex ratio, body size at first sexual maturity, gonadosomatic index, condition factor, fecundity and reproductive period of P. rhomboides in the natural environment. Twelve monthly samplings (mean n = 20) were carried out from March 2009 to February 2010. Fish were captured using 4 cm diameter cast nets and environmental parameters such as rainfall, temperature, dissolved oxygen concentration, pH and electrical conductivity of the water were registered. The sampled population of P. rhomboides (n = 444) showed a sex ratio of 1 : 1. Females attained first sexual maturity at 15 cm total length and males at 15.3 cm total length. Spawning occurred from March to May, with high gonadosomatic index values during the rainy season. This species is a total spawner, with a mean fecundity of 6188 (±367) mature oocytes. P. rhomboides presents a seasonal reproductive strategy, providing a population increase and exploitation of the environmental resources during the rainy period. Rainfall (196.5 ± 4.9 mm) and the concentration of dissolved oxygen (7.82 ± 0.7018 mg L?1) were influential factors during the spawning season. Gonadosomatic index and the condition factor were negatively correlated during the gonadal development cycle of this species. The monthly gonadosomatic index values show that spawning begins when the rainy season is at its maximum and ends synchronously (females r = 0.9171; males r = 0.889) with the end of the rainy season.  相似文献   

18.
Age, growth and reproductive characteristics of creole perch, Percichthys trucha, were investigated in the Negro River, southern Argentina from samples collected seasonally, December 1994–December 1995. Age was estimated via scale and whole otolith reading methods. Total length (n = 413) ranged from 103 to 432 mm, and weight from 12 to 1042 g. Significant differences between the length‐weight relationships of males and females were detected (P < 0.05). Isometric growth was observed in juveniles and males, whereas total population and females exhibited positive allometric growth. The marking pattern in scales and otoliths followed an annual rhythm, with the formation of only one annulus in scales and only one hyaline band in otoliths during autumn‐winter. The oldest males were 5 years old whereas maximum age in females was 12 years from scales and 15 years from otoliths. Because scales were found to underestimate age in individuals older than 4 years, otoliths were considered to be the best structures for creole perch age determination. Gompertz growth parameters based on otolith data were L∞: 428.0 mm, k = 0.46 and t0 = 0.43 for total population (r = 0.90), L∞: 410.7 mm, k = 0.42 and t0 = 0.46 for males (r = 0.91), and L∞: 434.1 mm, and k = 0.49 and t0 = 0.43 for females (r = 0.91). Lengths at first maturity (TL50) were 260 and 241 mm in males and females, respectively, both of which corresponded to ages between 1 and 2 years. Macroscopic gonad inspection and the high percentage of juveniles captured during summer indicated that spawning begins at the end of spring.  相似文献   

19.
The grey gurnard Eutrigla gurnardus (L.) has been identified by ICES as a potential commercial species in the NE Atlantic with recommendations made to derive information on population biology for stock assessment purposes. However, data on the population biology of this species is limited. In this study, data on the age, growth and maturity of grey gurnard were collected by otter trawling in the coastal waters of northwest Wales and Eastern Anglesey. Total length (TL) of fish sampled ranged between 2.1–33.0 cm (male) and 1.9–36.9 cm (female) with the majority of female (70.8%) fish between 11 and 20 cm TL and male fish (70.5%) between 11 and 18 cm TL. The percentage of fish >20 cm TL was larger for females (30.4%) compared to males (17.6%). Total weight (TW) for female and male grey gurnard in the stratified subsample ranged from 1.9 to 499.9 g for females and 2.1–390.0 g for males, with the majority of female (66.3%) and male (76.1%) fish between 10 and 60 g. TL/TW relations for male and female fish and both sexes combined were: TW = 0.006TL3.07, TW = 0.007TL3.03 and TW = 0.007TL3.05 respectively. Age structure (based on otolith reading) ranged between 0.5 and 7.5 years old for females and 0.5 to 5.5 years old for male with the majority of female (41.7%) and male (46.0%) fish aged as 1.5 years old. The age structure of female and male grey gurnards was significantly different with the majority of older fish (>2.5 years) being female. The von Bertalanffy growth functions were calculated as Lt = 32.4[1 ? e?0.24(t + 1.41)] for males, Lt = 45.9[1 ? e?0.16(t + 1.37)] for females and Lt = 44.0[1 ? e?0.18(t + 1.20)] for both sexes combined. Instantaneous rates of total mortality were similar for males and females and the combined Z value 1.00 year?1 with the natural mortality rate estimated as 0.33 year?1. The size at 50% maturity (L50) was estimated to be 25.3 cm TL for males, females and for both sexes combined. Age at 50% maturity (A50) was 3.2 years for both males and females. The results of this study provide the first information on the population biology of E. gurnardus in the Irish Sea, the first detailed study in the NE Atlantic since 1985 and helps to address the data gap identified by ICES in knowledge of the population biology of this species.  相似文献   

20.
Age, growth, and mortality of the edible crab, Cancer pagurus, were determined for the native population in South Wales (UK). Sampling was carried out on a monthly basis between February 2001 and September 2002. Carapace width ranged between 10.4 and 163 mm. Based on the carapace width frequency distribution, the Swansea and Gower population was composed mainly of males belonging to the first and second age‐class (1 and 2), and of females belonging to the third and fourth age‐class (3 and 4). Sex ratio was 1.126 ± 0.27 in favour of males. Carapace width frequency distributions and weight‐at‐age data were used to estimate the von Bertalanffy growth equation parameters. For the population as a whole, these were: L∞ = 199 mm, W∞ = 1179.56 g, K = 0.24 year?1, t0 = ?0.1004 years. The overall carapace width–weight relationship was: W = 0.38(CW2.69). Analysis of covariance indicated a significant difference in the carapace width–weight relationship between males and females in the study area. Total mortality Z and natural mortality M rates for combined sexes were 1.245 year?1 and 0.567 year?1, respectively. The exploitation ratio E was estimated to be 54.43%.  相似文献   

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