Notes on the phylogenetic relationships of planthoppers of the family issidae (Hemiptera,Fulgoroidea) of the Western Palaearctic fauna,with descriptions of two new genera

Seven groups of genera are distinguished in the subtribe Issina Spinola. The new genera Apedalmus gen. n. (the type species Falcidius abruzicus Dlabola, 1983) and Thalassana gen. n. (the type species: Hysteropterum ephialtes Linnavuori, 1971) are erected. The subgenera Acrestia Dlabola, 1980, Montissus Gnezdilov, 2003, and Atticus Gnezdilov, 2003 are upgraded to genera. The subgenus Capititudes Gnezdilov, Drosopoulos et Wilson, 2004 is transferred to the genus Acrestia, and the subgenus Stygiata Emeljanov, 1972 to Celyphoma Emeljanov, 1971. The subtribe Agalmatiina Gnezdilov, 2002 and the subfamily Hysteropterinae Melichar, 1906 are placed in synonymy under Issina Spinola, 1839. Issus reticulatus Herrich-Schäffer, 1835 is designated as the type species of the genus Hysteropterum Amyot et Serville, 1843.     

New and little known species of the planthopper genus <Emphasis Type="Italic">Mycterodus</Emphasis> Spinola (Homoptera,Issidae) from the Eastern Mediterranean

Three new species of the genus Mycterodus Spinola are described: M. phoenicicus sp. n. from Lebanon and M. syriacus sp. n. from Syria, both belong to the subgenus Aegaeum Gnezdilov, and M. marki sp. n. from Turkey, belongs to the subgenus Aconosimus Dlabola. Figures of the described species and of the holotype of M. anaticeps Puton are given. M. efesicus Dlabola is for the first time recorded from Greece (Samos I.); M. alatus Logvinenko, M. caucasicus (Melichar), and M. johannesi Gnezdilov &; Drosopoulos are new to the fauna of Turkey.     

Improved tribal delimitation of the subfamily Dictyopharinae and description of new genera and new species (Homoptera,Fulgoroidea, Dictyopharidae)

Ten new genera, five new subgenera, and five new species are described in the family Dictyopharidae. Three generic names are synonymized. A new name is proposed for the generic homonym. Dictyophara kazeruna Dlabola is transferred to the genus Callodictya Melichar. The genus Sinodictya Matsumura, with the type species Sinodictya tukana Matsumura, is redescribed. A new key to the tribes of the subfamily Dictyopharinae is given. The composition and characters of the tribes Taosini and Lappidini are revised. All the genera of the subfamily Dictyopharinae are listed according to their tribal position. New records are given for some interesting species.     

The first record of <Emphasis Type="Italic">Callodictya kazeruna</Emphasis> Dlabola (Hemiptera,Auchenorrhyncha, Fulgoroidea: Dictyopharidae) from Turkey

Callodictya kazeruna (Dlabola, 1986) described from southern Iran is recorded here for the first time from Eastern Anatolia in Turkey. A key to the two known species of the genus Callodictya Melichar, 1912 is given.     

亚草叶蝉属分类订正（半翅目：叶蝉科：角顶叶蝉亚科：隆脊叶蝉族）

对亚草叶蝉属进行系统研究并首次报道其在中国分布,共记述5种：E. acarifer (Lethierry),E. hilaris (Melichar),E. magnus Mitjaev,E. medius (Mulsant & Rey) 和E. pratensis Emeljanov,其中E. hilaris (Melichar) 和E. medius (Mulsant & Rey) 为中国新记录种。文中提供了分种检索表和雄性外生殖器特征图,并首次记述和提供雌性生殖器特征。     

A new genus for <Emphasis Type="Italic">Hysteropterum boreale</Emphasis> Melichar, 1902 (Hemiptera,Auchenorrhyncha: Fulgoroidea: Issidae) from China

A new genus is erected for Hysteropterum boreale Melichar, 1902 which is redescribed. Lectotype is designated in the type series of the species deposited in the Zoological Institute of the Russian Academy of Sciences (St. Petersburg, Russia). Photographs of the female (paralectotype) and drawings of the male (lectotype) genitalia are provided.     

Revision of the tribe Colpopterini Gnezdilov, 2003 (Homoptera, Fulgoroidea: Nogodinidae)

The tribe Colpopterini Gnezdilov, 2003 is revised, upgraded to the subfamily Colpopterinae, stat. n., and transferred from the family Issidae Spinola to Nogodinidae Melichar. The tribe Colpopterini is recorded from the Afrotropical Region for the first time—Bumerangum deckerti gen. et sp. n. is described from Southern Africa. The tribe Colpopterini s. str. comprises 6 genera: Bumerangum gen. n., Caudibeccus Gnezdilov et O’Brien, 2008, Colpoptera Burmeister, 1835, Jamaha Gnezdilov et O’Brien, 2008, Neocolpoptera Dozier, 1931, and Ugoa Fennah, 1945. The genera Cheiloceps Uhler, 1895, Tempsa Stål, 1866, Eupilis Walker, 1857, and Gabaloeca Walker, 1870 are transferred to the tribe Issini Spinola of the family Issidae. Issus longulus Lethierry, 1890 is transferred to the genus Colpoptera Burmeister. A key to the genera and a list of the species of the tribe Colpopterini are given. Morphological data confirming independent evolution of similar ovipositor types in the families Issidae and Nogodinidae are provided. The term “styletization” is suggested for describing the transformation of the ovipositor from a rounded to an elongate type.     

A new subfamily of the planthopper family Ricaniidae Amyot et Serville (Homoptera,Fulgoroidea)

Pharsalinae subfam. n. is erected in the family Ricaniidae Amyot et Serville for two monotypical genera Pharsalus Melichar, 1906 and Silvanana Metcalf, 1947 distributed in South America. The members of the new subfamily are characterized by the submacroptery, simple radius and anterior cubitus of the fore wing, the style capitulum bearing a lateral tooth, and the subrectangular (in lateral view) gonoplacs with partly separated third lobes and without marginal teeth. Pharsalus repandus Melichar, 1906 is recorded from Uruguay for the first time.     

New and little known planthoppers of the subfamily Ommatidiotinae (Homoptera,Fulgoroidea, Caliscelidae) from Madagascar and South Asia

Cano merinus gen. et sp. n., belonging to the tribe Augilini Baker, is described from Madagascar. Tubilustrium typicum Distant, 1916 is transferred from the tribe Ommatidiotini Fieber to the tribe Augilini. Lasonia kirkaldyi Melichar, 1903 is transferred from the family Issidae to the family Caliscelidae, tribe Adenissini Dlabola. A key to the tribes of the subfamily Ommatidiotinae is provided.     

中国颖蜡蝉科一新纪录属及一新纪录种(半翅目:蜡蝉总科)

记述了中国颖蜡蝉科Achilidae1属:埃颖蜡蝉属EpiramaMelichar及1种:弗氏埃颖蜡蝉E.francescoides Fennah,1965。同时提供了该属属征及该种雄虫与雌虫的鉴别特征图。研究标本保存在中国科学院动物研究所(IZCAS)动物标本馆。     

Revision of the superfamily Rhynchoporoidae Muir-Wood, 1955

The revision of punctate rhynchonellids of the superfamily Rhynchoporoidea revealed that the superfamily is composite. Based on the detail study of shell interior, the punctate rhynchonellids of the genus Rhynchopora King, 1865, subfamily Rhynchoporinae Muir-Wood, 1955 are placed into the family Trigonirhynchiidae Schmidt, 1965, superfamily Rhynchotrematoidea Schuchert, 1913. The genera Greira Erlanger, 1993, Tchanakhtchirostrum Sartenaer et Plodowski, 2003, Sharovaella Pakhnevich, 2012, Zaigunrostrum Sartenaer et Plodowski, 2003 are also referred to this family within the subfamily Greirinae Erlanger, 1993. The only genus of the subfamily Tretorhynchinae Savage, 2002, Tretorhynchia Brunton, 1971, is moved to the family Leiorhynchidae Stainbrook, 1945, superfamily Camarotoechioidea Schuchert, 1929. Araratella Abrahamian, Plodowski, et Sartenaer, 1975 within the subfamily Araratellinae Erlanger, 1986 is referred to the family Septalariidae Havlicek, 1960 of the same superfamily. The position of Rariella Zhang, 1981, Yingtangella Bai et Ying, 1977, Momarhynchus Baranov et Sartenaer, 1996 in the order Rhynchonellida remains controversial. The punctation and exopunctation is found to arise parallel in different rhynchonellid evolutionary branches after the Frasnian-Famennian mass extinction.     

An annotated checklist of the planthoppers of Iran (Hemiptera, Auchenorrhyncha, Fulgoromorpha) with distribution data

A list of Hemiptera Fulgoromorpha (planthoppers) of Iran is provided, based primarily on literature records from 1902 to the present. In total 15 families and 235 species are recorded, with taxonomic details. Distribution data in Iran are given. Iranissus ephedrinus Dlabola, 1980 is transferred from Issidae to Nogodinidae. To resolve nomenclatural difficulty the following new combinations in Issidae are given: Iranodus dumetorus (Dlabola, 1981), Iranodus khatunus (Dlabola, 1981) and Iranodus repandus (Dlabola, 1981). Due to published generic synonomy the following are new combinations: Duilius seticulosus (Lethierry, 1874), Duilius tamaricis (Puton & Lethierry, 1887), Duilius tamaricicola (Dubovsky, 1966) and Duilius v-atrum (Dlabola, 1985).     

Two new replacement names for the planthopper genera in Dictyopharidae (Hemiptera,Fulgoromorpha)

New replacement names are proposed for two genera of the family Dictyopharidae (Hemiptera: Fulgoromorpha). The following changes are proposed: Neonotostrophia nom. n. for Notostrophia Emeljanov (not Waterhouse); Emeljanovina nom. n. for Glochina Emeljanov (not Meigen); Neonotostrophia nigrosuturalis (Melichar, 1912) comb. n. from Notostrophia nigrosuturalis (Melichar, 1912) = Dictyophara nigrosuturalis Melichar, 1912 and Emeljanovina dixoni (Distant, 1906) comb. n. from Glochina dixoni (Distant, 1906) = Dictyophara dixoni Distant, 1906.     

Addenda to the Revisions of the Genera <Emphasis Type="Italic">Gergithus</Emphasis> Stål and <Emphasis Type="Italic">Hemisphaerius</Emphasis> Schaum (Hemiptera,Auchenorrhyncha, Fulgoroidea: Issidae)

The genus Gnezdilovius Meng, Webb et Wang, 2017 is revised. Maculergithus Constant et Pham, 2016, which was described as a subgenus of Gergithus Stål, 1870, is upgraded to a genus. Ishiharanus Hori, 1969 is reinstalled from synonymy with Gergithus and considered a valid name. Two new genera are erected, Ceratogergithus Gnezdilov, gen. n. (type species: Gergithus spinosus Che, Zhang et Wang, 2007) and Ophthalmosphaerius Gnezdilov, gen. n. (type species: Hernisphaerius trilobulus Che, Zhang et Wang, 2006). Hernisphaerius bistriatus Schumacher, 1915, Gergithus carbonarius Melichar, 1906, G. rosticus Chan et Yang, 1994, G. nummarius Chan et Yang, 1994, and G. rotundus Chan et Yang, 1994 are transferred to the genus Epyhemisphaerius Chan et Yang, 1994, Gergithus quinquemaculatus Che, Zhang, Wang, 2007—to the genus Maculergithus, Gergithus chelatus Che, Zhang, Wang, 2007 and G. pseudotessellatus Che, Zhang, Wang, 2007—to the genus Ceratogergithus, Hernisphaerius binocularis Chen, Zhang, Chang, 2014—to the genus Ophthalmosphaerius, and Gergithus robustus hoozanensis Schumacher, 1915—to the genus Gnezdilovius. The male genitalia of Gergithus herbaceus (Kirby, 1891) and Hernisphaerius interclusus Noualhier, 1896 are illustrated for the first time.     

Review of the subgenus Semirodus Dlabola of the genus Mycterodus Spinola (Homoptera: Issidae)

The subgenus Semirodus Dlabola, 1987 of the genus Mycterodus Spinola, 1839 with all known species are redescribed. Four new species of the subgenus from Greece and West Turkey are described. A key to the species of the subgenus is proposed.