A new species of comb-clawed beetle (Coleoptera: Tenebrionidae: Alleculinae) from the Lower Cretaceous of Yixian (China,Liaoning Province)

A new genus and species of comb-clawed beetles, Calcarocistela kirejtshuki gen. et sp. nov., from the Lower Cretaceous of Yixian (China) is described. The new genus is assigned to the tribe Gonoderini of the subfamily Alleculinae (family Tenebrionidae) based on the following characters: five visible abdominal ventrites, nonwidened tarsomeres without a membranous ventral lobes, and serrate antennae. The new genus differs from other members of the tribe Gonoderini in the long spurs of metatibia reaching a half of the first metatarsomere and the lamellate prosternal process. Calcarocistela kirejtshuki is the first member of Alleculinae from the Lower Cretaceous.     

New mesozoic diving beetles (Coleoptera,Dytiscidae) from China

A new subfamily of predaceous diving beetles, Liadytiscinae subfam. nov., including two genera and four species, Liadytiscus gen. nov. (L. cretaceus sp. nov., L. longitibialis sp. nov., and L. latus sp. nov.) and Liadroporus gen. nov. (L. elegans sp. nov.), from the Late Tithonian-Berriasian (Huangbanjigou, Yixian Formation) of China is described. Two new genera, Mesoderus gen. nov. with two species, M. magnus sp. nov. and M. ventralis sp. nov., and Sinoporus gen. nov. with one species, S. lineatus sp. nov., are also described; their position in the system of Dytiscidae remains uncertain. Relationships of the taxa described with Recent and fossil taxa of the same rank and some presumable ecological features of the new taxa are discussed.     

Notes on the genus Xenocerogria (Coleoptera,Tenebrionidae, Lagriini) from China

Three species of the genus Xenocerogria Merkl, 2007 have been recorded in China, Xenocerogria feai (Borchmann, 1911), Xenocerogria ignota (Borchmann, 1941) and Xenocerogria ruficollis (Borchmann, 1912). Xenocera xanthisma Chen, 2002 is proposed as a junior synonym of Xenocerogria ruficollis. Lectotype of Xenocerogria ignota is designated, and the species is transferred to the genus Lagria Fabricius, 1775. New Chinese province records of Xenocerogria ruficollis are provided.     

Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)

Six new species of Laena Dejean, Laena quadratasp. n. and Laena motoganasp. n.(China: Xizang), Laena chiloriluxasp. n., Laena dentatasp. n. and Laena liangisp. n. (China: Yunnan), Laena dentatocrassasp. n. (China: Hainan Island, representing new province record of the genus) are described, complemented with photos of habitus, illustrations of legs, antenna, aedeagus and last abdominal ventrite of male and female. Type specimens are deposited in both the Museum of Hebei University, Baoding, China and the Natural History Museum of Stuttgart, Germany.A key to the 102 Chinese species of genus Laena is provided.     

Two new species of the Stenochinus amplus species-group from China (Coleoptera,Tenebrionidae, Stenochiini)

Two new species of the Stenochinus amplus species-group are described, S. apiciconcavussp. n. (CHINA: Shaanxi) and S. xinyicussp. n. (CHINA: Guangdong). Also, some new distribution data are provided for S. cylindricus (Gebien, 1914), and a key to the seven species of the S. amplus species-group from China is given.     

中国广东优树甲属一新种(鞘翅目:拟步甲科)

记述采自中国广东的优树甲属1新种:高氏优树甲Uenostrongylium gaoi Lin&Yuan sp.nov.,该种因鞘翅刻点上缘具小颗粒而与浙江的粗皱优树甲Uenostrongylium scaber Yuan&Ren,2018相似.文中提供了新种的整体图和特征图.     

First fossil representative of the family omalisidae (Coleoptera,Elateroidea sensu lato) from the Baltic amber

The paper describes Jantarokrama utilis Kovalev et Kirejtshuk, gen. et sp. nov., the first fossil representative of the family Omalisidae from the Upper Eocene Baltic amber, which is similar to the Recent Phaeopterus unicolor Costa, 1856, but distinguished from the latter by the larger and not so slender body, smaller distance between antennal insertions, longer antennae, wider prothorax with very convex anterior edge of the pronotum, and particularly by five completely exposed abdominal ventrites. The diagnosis of the new genus among generic taxa of Omalisidae and its similarity to Berendtimirus Winkler, 1987 (Berendtimiridae) are discussed.     

Two new species of Itagonia Reitter (Coleoptera,Tenebrionidae, Blaptini) from China

Two new species of Itagonia Reitter, 1887, Itagonia tibialis sp. n. and Itagonia litangensis sp. n. are described from Sichuan, China. A key to the known species of Itagonia from China is given.     

Basal titanosauriform (Dinosauria, Sauropoda) teeth from the Lower Cretaceous Yixian Formation of Liaoning Province, China

The Yixian Formation of Liaoning Province, People's Republic of China has yielded a diverse fauna of non-avian dinosaurs, but is dominated by small-bodied taxa. Here, we describe a series of isolated teeth from the Lujiatun Beds of the formation that are referable to a basal titanosauriform sauropod. Some of the teeth possess a distinctive circular boss on the lingual surface, which suggests that they are referable to cf. Euhelopus sp. This identification provides some additional support for biostratigraphical correlations between the Jehol Group and the Mengyin Formation of Shandong Province that suggest an Early Cretaceous age for the latter unit. Moreover, the titanosauriform affinities of the teeth provide further evidence for the dominance of this sauropod clade in eastern Asia during the Cretaceous.     

New species of variegated mud-loving beetles (Coleoptera: Heteroceridae) from mesozoic deposits of China

A new species of variegated mud-loving beetles, Heterocerites magnus, sp. nov., is described from the Late Tithonian-Berriasian of China (Huangbanjigou, Yixian Formation). This is a second known representative of the family in the Mesozoic.     

First phylogenetic analysis of the subfamily Pachydeminae (Coleoptera,Scarabaeoidea, Melolonthidae): the Palearctic Pachydeminae*

This paper presents the first phylogenetic analysis of Pachydeminae Reitter, 1902 ; one of the least known subfamilies of Melolonthidae, `leaf‐chafers' (Scarabaeoidea, Coleoptera). Some species of Pachydeminae have recently become agricultural pests in southern Spain. We analysed the phylogenetic relationships among 49 species belonging to 16 genera in the Palearctic region, based on a set of 63 morphological characters from the adult external morphology, wing anatomy, mouthparts and male and female genitalia. The last three sets of characters are described here for the first time. The phylogeny shows that the Palearctic Pachydeminae are monophyletic within the subfamily. Mouthparts and male and female genitalia provide the best synapomorphies for intergeneric relationships. In contrast, most of the external morphological characters used in the taxonomy of Pachydeminae are highly homoplastic. The phylogeny shows a basal split between the genera Hemictenius Reitter, 1897; Pachydema Castelnau, 1832, and the monospecific Peritryssus Reitter, 1918; and a second clade including the rest of genera. The remarkable Peritryssus is confirmed as a Pachydeminae, being the sister group to the monophyletic Hemictenius . Except for the position of P. rubripennis (Lucas, 1848) and P. zhora Normand, 1951, the phylogeny supports the monophyly of Pachydema but rejects the traditional division into species groups and the monophyly of the endemic Canarian species. In contrast, Tanyproctus Faldermann, 1835, must be rejected as polyphyletic. Otoclinius Brenske, 1896, is also probably polyphyletic (two new species synonymies), whereas Leptochristina Baraud and Branco, 1991 , is either mono‐ or paraphyletic. The two Mediterranean genera Ceramida Baraud, 1897, and Elaphocera Gené, 1836, form a monophyletic group, this clade being the best supported by the data set. Ceramida is clearly monophyletic, whereas Elaphocera is probably monophyletic except for E. barbara Rambur, 1843, which shares with Ceramida the character state for numerous mouthpart and genitalic characters. The phylogeny questions the generic status of the small and monospecific genera of Pachydeminae. The monotypic Alaia Petrovitz, 1980 , and Brenskiella Berg, 1898, are merged with Europtron Marseul, 1867, into one clade, whereas Atanyproctus Petrovitz, 1954, is grouped with some species of Tanyproctus , and the monotypic Pachydemocera Reitter, 1902 , is proposed as a junior synonym of Elaphocera .     

New water scavenger beetles (Coleoptera: Hydrophilidae) from the mesozoic of Mongolia

A new genus and two new species of water scavenger beetles, Hydrophilopsia bontsaganica, sp. nov. and Prospercheus cristatus, gen. et sp. nov., are described from the Upper Jurassic and Lower Cretaceous of Mongolia. The systematic position of the new genus is discussed.     

中国逍遥蛛科化石新属种（蜘蛛目，逍遥蛛科）

记述产自中国辽西的逍遥蛛科化石1新属：白垩逍遥蛛属Cretadromus gen．nov．,1新种：辽宁白垩逍遥蛛Cretadromus liaoningensis sp．nov．：这是逍遥蛛科化石在中国的首次发现,也是逍遥蛛科在晚侏罗世-早白垩世的首次报道。标本采白辽宁省凌源市大王杖子晚侏罗世-早白垩世义县组地层,现存于大连自然博物馆：     

Forearm bone histology of the small theropod Daliansaurus liaoningensis (Paraves: Troodontidae) from the Yixian Formation,Liaoning, China

We present the first histological analysis of forelimb bones in a troodontid dinosaur, Daliansaurus liaoningensis, from the Early Cretaceous of China using osteohistological thin-sectioning and high-resolution synchrotron-based imaging. The thin wall compacta consists of primary bone, and three lines of arrested growth (LAG) in the radius (R) and two in the ulna (U) divide these into successive zones. Results show that the new fossil has four distinct bone depositional rates: (1) fastest deposition in inner zones R1 and U1 (fibro-lamellar bone with a plexiform-like vasculature); (2) slowed deposition in outer zones R1 and U1 + U2 (loss of vascular density and plexiform component); (3) fluctuating rates of deposition in zones R2 + R3 and in the inner zone U3 (alternating bands of circumferentially organised primary osteons and avascular bone); and (4) slowest deposition in zone R4 and the outer zone U3 (lamellar bone constituting the external fundamental system). Collectively, these growth characteristics suggest that the fossil is an individual that passed the exponential growth phase by the first year, and perished three years later. We conclude that the histology is consistent with an interpretation of this specimen as a late maturing individual that had not yet attained maximum somatic size.     

A new synonym in the Tribe Blaptini (Coleoptera,Tenebrionidae)

Type specimen of Platyscelis provostii Fairmaire, 1888 has been examined and is re-described herein. The following new synonymy is established: Platyscelis provostii Fairmaire, 1888 = Itagonia ganglbaueri Schuster, 1914, syn. n.     

Sperm morphology and phylogeny of lagriids (Coleoptera,Tenebrionidae)

The systematics of tenebrionids remain unclear, principally at the subfamily level, as is the case of the Lagriinae. Considering that sperm morphology has contributed to the various insect group phylogenies, in this work we describe the structure and ultrastructure of these cells in Lagria villosa. Sperm in this species exhibit a strong morphological similarity to those of Tenebrio molitor and Tribolium castaneum, the only two species of Tenebrionidae with previously described sperm. In tenebrionids, the flagellar components offer good diagnostic characters, e.g. the symmetry of mitochondrial derivatives in L. villosa differentiates sperm of this species from those of Te. molitor and Tr. castaneum. However, the lateral association of the nucleus with flagellar components, the form of accessory bodies, and the presence of material connecting the axoneme to the accessory bodies and mitochondrial derivatives indicate that the three species form a sister group. Therefore, the sperm morphology of L. villosa support lagriid beetles as a subfamily (Lagriinae) of Tenebrionidae.     

中国云南烁甲属分类研究及中国三新纪录种(鞘翅目:拟步甲科:烁甲族)

研究了中国云南拟步甲科Tenebrionidae烁甲族Amarygmini烁甲属Amarygmus Dalman,1823,发现中国3新纪录种,越北烁甲A.tonkineus Pic,1922、丝角烁甲A.filicornis(Gravely,1915)和肿腿烁甲A.nodicornis(Gravely,1915),云南2新纪录种,艾多烁甲A.adonis(Pic,1922)和中国烁甲A.sinensis Pic,1922,提供了该属云南已知种检索表、地理分布。检视标本保存于河北大学博物馆。