用光合-蒸散耦合模型模拟冬小麦CO2通量的日变化

根据SPAC理论建立了一个冬小麦光合和蒸散的耦合模型.冬小麦CO2通量包括冠层光合、呼吸和土壤呼吸.冠层光合采用了Farquhar光合作用生化模型,并通过冠层阻力的参数化将光合作用与蒸腾作用耦合起来.用涡度相关方法观测了CO2通量,对模型进行了验证,结果显示模型可以较好地模拟CO2通量日变化过程.对模型的敏感性分析发现日间CO2通量最敏感的参数是初始量子效率.其次,CO2通量对光响应曲线凸度、CO2补偿点、凋萎点和叶面积指数的变化也有着较强的敏感性;夜间CO2通量敏感的参数是最适温度下Rubisco催化能力和暗呼吸参数.     

用光合-蒸散耦合模型模拟冬小麦CO2通量的日变化

根据 SPAC理论建立了一个冬小麦光合和蒸散的耦合模型。冬小麦 CO2 通量包括冠层光合、呼吸和土壤呼吸。冠层光合采用了 Farquhar光合作用生化模型 ,并通过冠层阻力的参数化将光合作用与蒸腾作用耦合起来。用涡度相关方法观测了 CO2通量 ,对模型进行了验证 ,结果显示模型可以较好地模拟 CO2 通量日变化过程。对模型的敏感性分析发现日间 CO2 通量最敏感的参数是初始量子效率。其次 ,CO2 通量对光响应曲线凸度、CO2 补偿点、凋萎点和叶面积指数的变化也有着较强的敏感性 ;夜间 CO2 通量敏感的参数是最适温度下 Rubisco催化能力和暗呼吸参数     

生态模型在农田蒸散及土壤水分模拟中的适用性评价

为评价生态模型在农田蒸散及土壤水分运动模拟中的适用性,利用2013—2015年南京农业气象测站观测数据,评估了BEPS(Boreal Ecosystem Productivity Simulator)模拟冬小麦农田生态系统逐日蒸散及与土壤水分动态的可靠性,并进一步开展了植被冠层蒸腾和农田土壤蒸发分离。模拟结果表明:BEPS适用于研究冬小麦农田蒸散量及土壤水分运动规律;由于考虑了叶片聚集指数和冬小麦根系垂直分布递减系数随生育期变动的参数化改进,BEPS分别可以解释2013—2014年和2014—2015年两个生长季农田生态系统蒸渗仪实测蒸散量变化的83%和74%,参数化改进前后模型效率ME相当(前:0.8,后0.74),标准差RMSE(前:1.50,后1.05),平均偏差MBE(前:0.5,后0.35),误差减小;两个生长季中,土壤蒸发占冠层上方总蒸散的比例随生育进程而变化,全生育期发散比平均值分别为34%和29%;BEPS模拟的0~40 cm土层深度土壤水分随时间变化趋势与实测值基本一致,可以解释78%以上的土壤水分实测值变化,并能快速地响应降水变化。本研究表明,生态模型可以用于模拟冬小麦农田蒸散和土壤水分变化,并有助于厘定农田冠层中难以区分的植被蒸腾和土壤蒸发的比例关系,可为进一步开展气候变化背景下的区域蒸散发评估及与之相联系的农田节水管理奠定基础。     

玉米农田蒸散过程及其对气候变化的响应模拟

应用基于生理生态学过程的EALCO模型,对玉米农田生态系统的蒸散(ET)过程进行了模拟,在模型检验基础上,使用该模型模拟了玉米农田生态系统ET过程对未来气候变化的响应。结果表明,EALCO模型中能量与水过程的动态耦合机制使模型能够较好地模拟农田蒸散过程,基于涡度相关法的观测值与模型模拟值在小时、日尺度上均吻合较好,模型可以解释67%的日蒸散的变化特征。对土壤蒸发与冠层蒸腾的分别模拟显示,生长季土壤蒸发约占ET的36%。温度的升高会引起ET与冠层蒸腾的增加,同时土壤蒸发减少;ET对降水减少的响应较为敏感,主要表现在土壤蒸发的下降。大气CO2浓度升高对冠层蒸腾影响显著,该情景下冠层蒸腾下降幅度最大。研究所假设的2100年气候情景下,该农田生态系统生长季蒸散将减少,然而相对于降水的减少而言,蒸散的减少量较小,即水分支出项相对增加,因此,发生土壤水分匮乏的可能性加大,这可能会加剧该地区的暖干化趋势,给作物产量及生态环境带来威胁。     

杉木人工林单叶至冠层光合作用的扩展与模拟研究

根据野外条件下杉木（Ｃｕｎｎｉｎｇｈａｍｉａ ｌａｎｃｅｏｌａｔａ（Ｌａｍｂ．）Ｈｏｏｋ．）针叶光合作用的测定结果。考虑到光合作用对光的非线性响应特性,及其与叶所处的实际冠层环境变量和冠层的空气动力学特性的相关,以筒化的林冠辐射传输模型为基础,结合不同部位和年龄针叶的光响应曲线,实现了叶室测量结果向冠层环境的调整,并进行了冠层光合作用模拟的初步研究。经过调整后的冠层光合作用平均比未经过调整的值高１     

温带阔叶红松林生态系统潜热通量模拟——气孔导度组合模型在Shuttleworth-Wallace双源模型中的应用

叶片水平的气孔导度组合模型已被成功扩展到冠层水平,并被应用于冬小麦生态系统潜热通量的模拟研究,但该研究仅基于1a的数据,有必要研究模型在更长时间尺度和其它生态系统类型的适用性.以长白山阔叶红松林(CBS)为研究对象,将组合模型进一步应用于Shuttleworth-Wallace双源模型,模拟了CBS 3a生长季内的潜热通量,利用涡度相关系统观测的潜热通量数据对模型进行验证,并对比了双源模型与单源模型的模拟结果.结果显示,双源模型较单源模型能取得更高的模拟精度,生长季不同时期的潜热通量模拟值和实测值的日变化较一致.对双源模型模拟值和实测潜热通量的相关分析显示,二者直线回归斜率和R2分别为0.96和0.72.对长白山阔叶红松林生态系统的蒸散和植被蒸腾的季节和年际变异分析发现,影响冠层蒸散和植被蒸腾季节动态的主要因素是饱和差和辐射,而影响它们年际动态的主要因素则是饱和差和温度.     

芦苇湿地多时空尺度蒸散模拟研究进展

湿地蒸散是湿地水分损失的主要途径,由于其对气候变化的高度敏感性和重要的反馈作用而倍受关注。湿地蒸散过程可以分为叶片、植株、冠层、景观至区域等多种尺度,如何实现不同时空尺度的过程和参数耦合成为湿地蒸散研究的重点和难点。本文回顾了湿地蒸散模拟研究的发展历程,着重总结了芦苇湿地蒸散监测及模拟方面的技术和方法,并从不同时间尺度(日、月、年)和不同空间尺度(叶片、植株、冠层、区域)以及实现不同时空尺度拓展的关键技术研究等方面进行归纳和总结,探讨了今后的研究重点。实现不同时空尺度的芦苇湿地蒸散过程的准确模拟,关键在于大气-植被-土壤界面的参数耦合,而目前针对不同尺度的芦苇湿地蒸散参数化研究很少,因此,有必要探讨适合芦苇湿地多尺度蒸散模拟及参数化方法,为芦苇湿地蒸散评估模型的选择提供理论支持。     

用光合-蒸散耦合模型模拟冬小麦水热通量的日变化

从SPAC理论出发,建立了一个冬小麦光合和蒸散的耦合模型．感热通量和潜热通量采用Shuttleworth-Wallace的双层模型计算,并通过冠层阻力的参数化,将光合作用与蒸腾作用耦合起来．用涡度相关方法,观测了感热通量和潜热通量,对模型进行了验证．结果表明,模拟值与观测值比较一致,模型可以很好地模拟感热通量和潜热通量的日变化过程．对模型的敏感性分析发现,冬小麦蒸腾比较敏感的参数有凋萎点、气孔导度参数、叶对红外辐射的反射率和光响应曲线凸度;土壤蒸发只对土壤阻力参数的敏感性较强．本模型对水热通量与环境因子作用过程的理论研究和指导农田的灌溉制度等有一定的意义．     

华北平原冬小麦农田蒸散量

以华北平原冬小麦农田为研究对象,采用涡度相关技术和热红外遥感技术,研究了不同环境条件下土壤含水量与农田蒸散量及作物冠层温度的关系.结果表明,冬小麦在农田郁闭(LAI≥3)、晴天和土壤相对含水量低于田间持水量65%的情况下,蒸发比值日变化正午前后出现相对较低且平稳的变化趋势.在晴天情况下,农田潜热通量与作物冠层温度日变化和季节变化均呈极显著的非线性相关关系,而冠气温差、农田相对蒸散量则与0～100 cm土层的土壤相对含水量密切相关.以13:30～14:00的平均冠层温度值T_c、日最高气温T_{a max}和日净辐射总量Rn_d为统计数据,确立了冬小麦农田日蒸散量ET_d （mm）估算简化模式参数.     

土壤-植物-大气连续体水热动态模拟的研究

本文从能量平衡原理和质量守恒定律出发,描述了土壤-植物-大气连续体中的热量转换和水分输送,模拟了系统中水分和热量的动态变化过程,并用所建立的模拟模型计算了冬小麦群落的冠层温度、叶水势及系统的潜热与显热变化关系,结果表明该模型有一定的可靠性。     

Canopy Photosynthesis and Crop Growth Rate of Eight Temperate Forage Grasses

The rates of canopy and individual leaf photosynthesis, ratesof growth of shoots and roots, and the extinction coefficientfor light of eight temperate forage grasses were determinedin the field during early autumn. Canopy gross photosynthesiswas calculated as net photosynthesis plus dark respiration adjustedfor temperature using a Q₁₀ = 2. The relationships between canopygross photosynthesis and light intensity were hyperbolic, andthe initial slopes of these curves indicated that light wasbeing utilized efficiently at low light intensities. The initialslope depended on the distribution of light in the canopy andthe quantum efficiency of the individual leaves. The maximumrate of canopy gross photosynthesis reflected the maximum rateof individual leaf photosynthesis. Although the maximum rateof canopy gross photosynthesis was correlated with crop growthrate, there was no significant relationship between daily grossphotosynthesis and crop growth rate. Indeed, daily gross photosynthesisvaried by only 22 per cent, whereas the daily growth of shootsand roots varied by 120 per cent. This poor correlation is influencedby a negative correlation (P < 0.01) between the maximumrate of canopy gross photosynthesis and the initial slope ofthe curve relating canopy gross photosynthesis and light intensity.Difficulties in the interpretation of measurements of dark respirationappeared to confound attempts to relate daily net photosynthesisto crop growth rate, individual leaf photosynthesis, and theextinction coefficient for light.     

Carbon balance in a monospecific stand of an annual herb <Emphasis Type="Italic">Chenopodium album</Emphasis> at an elevated CO<Subscript>2</Subscript> concentration

Elevated CO₂ enhances carbon uptake of a plant stand, but the magnitude of the increase varies among growth stages. We studied the relative contribution of structural and physiological factors to the CO₂ effect on the carbon balance during stand development. Stands of an annual herb Chenopodium album were established in open-top chambers at ambient and elevated CO₂ concentrations (370 and 700 μmol mol⁻¹). Plant biomass growth, canopy structural traits (leaf area, leaf nitrogen distribution, and light gradient in the canopy), and physiological characteristics (leaf photosynthesis and respiration of organs) were studied through the growing season. CO₂ exchange of the stand was estimated with a canopy photosynthesis model. Rates of light-saturated photosynthesis and dark respiration of leaves as related with nitrogen content per unit leaf area and time-dependent reduction in specific respiration rates of stems and roots were incorporated into the model. Daily canopy carbon balance, calculated as an integration of leaf photosynthesis minus stem and root respiration, well explained biomass growth determined by harvests (r ² = 0.98). The increase of canopy photosynthesis with elevated CO₂ was 80% at an early stage and decreased to 55% at flowering. Sensitivity analyses suggested that an alteration in leaf photosynthetic traits enhanced canopy photosynthesis by 40–60% throughout the experiment period, whereas altered canopy structure contributed to the increase at the early stage only. Thus, both physiological and structural factors are involved in the increase of carbon balance and growth rate of C. album stands at elevated CO₂. However, their contributions were not constant, but changed with stand development.     

The effect of ryegrass mosaic virus on carbon assimilation and growth of ryegrasses

The growth of simulated swards of Italian and perennial ryegrass, artificially infected with ryegrass mosaic virus (RMV) or RMV-free, was investigated. During the build up of RMV infection, dry-matter yield was usually decreased in infected swards, and the effect was confirmed when single vegetative regrowths of swards were investigated in detail. The primary cause of lower yields in RMV-infected swards appears to be a decrease in net canopy photosynthesis (maximum decrease about 50%) and an associated increase in dark respiration (maximum increase about 50%). The decrease in net canopy photosynthesis is due primarily to a lowered rate of net photosynthesis of constituent leaves within the canopy. A secondary effect of RMV infection is to decrease tillering (maximum decrease about 30%) which results in a change in canopy structure and in particular a lower leaf area index. In the present experiments, lower light utilization by the swards is less important than the decrease in leaf photo-synthetic efficiency in lowering yield.     

Canopy photosynthetic production in a Japanese larch forest. II. Estimation of the canopy photosynthetic production

Seasonal changes and yearly gross canopy photosynthetic production were estimated for an 18 year old Japanese larch (Larix leptolepis) forest between 1982 and 1984. A canopy photosynthesis model was applied for the estimation, which took into account the effect of light interception by the non-photosynthetic organs. Seasonal changes in photosynthetic ability, amount of canopy leaf area and light environment within the canopy were also taken into account. Amount of leaf area was estimated by the leaf area growth of a single leaf. The change of light environment within the canopy during the growing season was estimated with a light penetration model and the leaf increment within the canopy. Canopy respiration and surplus production were calculated as seasonal and yearly values for the three years studied. Mean yearly estimates of canopy photosynthesis, canopy respiration and surplus production were 37, 13 and 23 tCO₂ ha⁻¹ year⁻¹, respectively. Vertical trend, seasonal changes and yearly values of the estimates were analyzed in relation to environmental and stand factors.     

Growth Response of a Chrysanthemum Crop to the Environment. II. A Mathematical Analysis Relating Photosynthesis and Growth

Vegetative crops of chrysanthemum were grown for 5 weeks inthree replicate daylit assimilation chambers. Weekly harvestswere made from each crop for growth analysis, and on seven occasionsduring the 5-week period continuous measurements of the netCO₂ exchange rate of each crop were made over a 24 h period.A semi-empirical model for canopy photosynthesis was fittedto these data. The photosynthesis model was then incorporatedinto a simple, dynamic growth model. Using fitted values ofthe canopy photosynthesis parameters, the daily total radiationintegrals, and the experimentally observed relationship betweenthe leaf area index and crop dry matter per unit ground area,the crop growth model was used to simulate growth over the 5-weekperiod. The predicted and measured crop dry weights were inclose agreement over the whole period.     

温室甜椒叶面积指数形成模拟模型

叶面积指数是光合作用驱动的作物生长模型以及冠层蒸腾模型所需的重要作物参数,温度和辐射是影响叶片生长的重要环境因子.通过不同定植期、不同品种、不同地点的试 验,定量分析了温室甜椒出叶数、叶片长度和叶面积指数与温度和辐射的关系,构建了温室 甜椒叶面积模型,并利用独立的试验资料对模型进行了检验.结果表明：甜椒出叶数与出苗 后累积辐热积呈指数函数关系;叶片长度与出叶后累积辐热积呈负指数函数关系;甜椒出叶 数、叶片长度和叶面积指数的模拟结果与实测值之间的决定系数R²分别为0.94、0.89、0.93,其回归估计标准误RMSE分别为3.4、2.15 cm、0.15.该模型能够利用气温、辐射、 种植密度和出苗日期准确地预测温室甜椒叶面积指数动态,且模型参数少、实用性强,可以为温室甜椒生长模型和蒸腾模型提供必需的叶面积指数动态信息.     

Modelling the effects of elevated atmospheric CO2 on crown development, light interception and photosynthesis of poplar in open top chambers

An open-top chamber experiment was carried out to examine the likely effects of elevated atmospheric [CO₂] on architectural as well as on physiological characteristics of two poplar clones ( Populus trichocarpa × P. deltoides clone Beaupré and P. deltoides × P. nigra clone Robusta). Crown architectural parameters required as input parameters for a three-dimensional (3D) model of poplar structure, such as branching frequency and position, branch angle, internode length and its distribution pattern, leaf size and orientation, were measured following growth in ambient and elevated [CO₂ ] (ambient + 350 μmol mol^–1) treated open-top chambers. Based on this information, the light interception and photosynthesis of poplar canopies in different [CO₂] treatments were simulated using the 3D poplar tree model and a 3D radiative transfer model at various stages of the growing season. The first year experiments and modelling results showed that the [CO₂] enrichment had effects on light intercepting canopy structure as well as on leaf photosynthesis properties. The elevated [CO₂] treatment resulted in an increase of leaf area, canopy photosynthetic rate and above-ground biomass production of the two poplar clones studied. However, the structural components responded less than the process components to the [CO₂] enrichment. Among the structural components, the increase of LAI contributed the most to the canopy light interception and canopy photosynthesis; the change of other structural aspects as a whole caused by the [CO₂] enrichment had little effect on daily canopy light interception and photosynthesis.     

Enhancement of rice canopy carbon gain by elevated CO(2) is sensitive to growth stage and leaf nitrogen concentration

Increasing our understanding of the factors regulating seasonal changes in rice canopy carbon gain (C(gain): daily net photosynthesis -- night respiration) under elevated CO(2) concentrations ([CO(2)]) will reduce our uncertainty in predicting future rice yields and assist in the development of adaptation strategies. In this study we measured CO(2) exchange from rice (Oryza sativa) canopies grown at c. 360 and 690 micromol mol(-1)[CO(2)] in growth chambers continuously over three growing seasons. Stimulation of C(gain) by elevated [CO(2)] was 22-79% during vegetative growth, but decreased to between -12 and 5% after the grain-filling stage, resulting in a 7-22% net enhancement for the whole season. The decreased stimulation of C(gain) resulted mainly from decreased canopy net photosynthesis and partially from increased respiration. A decrease in canopy photosynthetic capacity was noted where leaf nitrogen (N) decreased. The effect of elevated [CO(2)] on leaf area was generally small, but most dramatic under ample N conditions; this increased the stimulation of whole-season C(gain). These results suggest that a decrease in C(gain) enhancement following elevated CO(2) levels is difficult to avoid, but that careful management of nitrogen levels can alter the whole-season C(gain) enhancement.