Brachiopods of the suborder Delthyridina Ivanova (order Spiriferida) from the Lower and Middle Devonian of northeastern Russia

New brachiopods of the suborder Delthyridina Ivanova (order Spiriferida) from the Lower and Middle Devonian of northeastern Russia, Ivanothyris costatus sp. nov. (subfamily Delthyridinae), Eoreticularia lata sp. nov., Yania gen. nov. with the type species Y. tumida sp. nov. (subfamily Eoreticulariinae), Reticulariopsis equitans Modzalevskaya, and R. (?) borealis sp. nov. (subfamily Reticulariopsinae) are described.     

New archaeocyatha from the Atdabanian Stage of the Siberian Platform

New Archaeocyatha genera and species, Turgorocyathus elegans gen. et sp. nov., Angustocyathus porus gen. et sp. nov., Flossocyathus squamosus gen. et sp. nov., Rotundocyathus oimuranicus sp. nov., and Carinacyathus inopinatus sp. nov., from the type sections of the Atdabanian Stage of the Lower Cambrian of the Siberian Platform are described.     

New Devonian and Carboniferous bryozoans of the Holy Cross Mountains (central Poland)

New bryozoans from the Middle-Upper Devonian (Eifelian-Frasnian) and the Lower Carboniferous (Tournaisian) of the Holy Cross Mountains in central Poland are described: Bigeyella sparsa gen. et sp. nov., B. separata gen. et sp. nov., Eridopora singula sp. nov., Leptotrypa pulchra sp. nov., Kysylschinipora klarae sp. nov., Coelotubulipora rara sp. nov., Alternifenestella genuina sp. nov., Exfenestella polonica sp. nov., and Rectifenestella localis sp. nov. Some paleogeographic and stratigraphic aspects of the Paleozoic deposits of this region are discussed and main bryozoan localities are described.     

A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida

Background

Lacewings (insect order Neuroptera), known in the fossil record since the Early Permian, were most diverse in the Mesozoic. A dramatic variety of forms ranged in that time from large butterfly-like Kalligrammatidae to minute two-winged Dipteromantispidae.

Principal Findings

We describe the intriguing new neuropteran family Parakseneuridae fam. nov. with three new genera and 15 new species from the Middle Jurassic of Daohugou (Inner Mongolia, China) and the Early/Middle Jurassic of Sai-Sagul (Kyrgyzstan): Parakseneura undula gen. et sp. nov., P. albomacula gen. et sp. nov., P. curvivenis gen. et sp. nov., P. nigromacula gen. et sp. nov., P. nigrolinea gen. et sp. nov., P. albadelta gen. et sp. nov., P. cavomaculata gen. et sp. nov., P. inflata gen. et sp. nov., P. metallica gen. et sp. nov., P. emarginata gen. et sp. nov., P. directa gen. et sp. nov., Pseudorapisma jurassicum gen. et sp. nov., P. angustipenne gen. et sp. nov., P. maculatum gen. et sp. nov. (Daohugou); Shuraboneura ovata gen. et sp. nov. (Sai-Sagul). The family comprises large neuropterans with most primitive wing venation in the order indicated by the presence of ScA and AA1+2, and the dichotomous branching of MP, CuA, CuP, AA3+4, AP1+2. The phylogenetic position of Parakseneuridae was investigated using a phylogenetic analysis of morphological scoring for 33 families of extinct and extant Neuropterida combined with DNA sequence data for representatives of all extant families. Parakseneuridae were recovered in a clade with Osmylopsychopidae, Prohemerobiidae, and Ithonidae.

Conclusions/Significance

The presence of the presumed AA1+2 in wings of Parakseneuridae is a unique plesiomorphic condition hitherto unknown in Neuropterida, the clade comprising Neuroptera, Megaloptera, Raphidioptera. The relative uncertainty of phylogenetic position of Parakseneuridae and the majority of other families of Neuroptera reflects deficient paleontological data, especially from critical important periods for the order, earliest Triassic and latest Triassic/earliest Jurassic.     

New and little-known Eoblattida (Insecta) from the Paleozoic of Russia

New taxa of the insect order Eoblattida from the Upper Carboniferous of Russia are described, including Narkemina kata sp. nov., Narkeminopsis inversa sp. nov., Carbonokata storozhenkoi gen. et sp. nov., Tshunoptera ampla gen. et sp. nov., and Evenkiophlebia collucata gen. et sp. nov. from the Chunya locality (Krasnoyarsk Region), Narkemulla sibirica gen. et sp. nov. from the Chunya and Izykhskie Kopi localities, Khakassia (all Cnemidolestidae), and Izykhia tridentis gen. et sp. nov. from Izykhskie Kopi (?Spanioderidae). Narkeminuta permiana gen. et sp. nov. (Cnemidolestidae) is described from the Kedrovka locality (Kemerovo Region; Lower Permian), and Issadische maximum gen. et sp. nov. (Eoblattida incertae familiae) is descriobed from the Isady locality (Vologda Region; Upper Permian). Permeoblatta borealis Rasnitsyn et Aristov, 2010 (Idelinellidae) from Isady is redescribed.     

New beetle species of the genus Lasiosyne (Coleoptera, Lasiosynidae) from the Late Jurassic and Early Cretaceous of Russia and Mongolia

Six new species of the genus Lasiosyne Tan et al., 2007, the type genus of the family Lasiosynidae Kirejtshuk et al., 2010, are described from Jurassic and Lower Cretaceous deposits of Mongolia and East Siberia: L. shartegiensis sp. nov., L. insculpta sp. nov., L. longitarsa sp. nov., L. cataphracta sp. nov., L. punctata sp. nov., and L. decora sp. nov. A modified diagnosis of the genus is proposed.     

Triassic radiolarians of Kotel’nyi Island (New Siberian Islands,Arctic)

Triassic radiolarians from Kotel’nyi Island (New Siberian Islands, Arctic Region) are studied. Radiolarians occur in the Middle and Upper Triassic beds, which are well dated by ammonoids, nautiloids, and bivalves. In the Middle Triassic beds, which are composed of dark gray and black clays and claystones with interbeds of combustible schist, siltstone, clayey limestone, and many phosphatic concretions, the following two assemblages are recognized: (1) a Late Anisian assemblage, including Glomeropyle clavatum Bragin, sp. nov., G. boreale Bragin, G. manihepuaensis Aita, G. insulanum Bragin, sp. nov., Triassospongosphaera multispinosa (Kozur et Mostler), Tetraspongodiscus hibernus Bragin, sp. nov., T. borealis Bragin, sp. nov., Eptingium abditum Bragin, sp. nov., E. manfredi Dumitrica, Spongostephanidium japonicum (Nakaseko et Nishimura), and Ladinocampe vicentinensis Kozur et Mostler; (2) Late Ladinian assemblage, with Muelleritortis firma (Gorican), M. kotelnyensis Bragin, sp. nov., Tiborella nivea Bragin, sp. nov., Pseudostylosphaera goestlingensis (Kozur et Mostler), P. omolonica Bragin, Triassospongosphaera multispinosa (Kozur et Mostler), Sarla cincinnata Bragin, sp. nov., S. obscura Bragin, sp. nov., S. prava Bragin, sp. nov., and Eonapora stiriaca Bragin, sp. nov. The Upper Triassic beds, composed of gray and black clays and claystones with lenses of clayey limestones, with sideritic and phosphatic-calciferous concretions have yielded (1) an Early Carnian assemblage, with Pentactinocarpus colum Bragin, sp. nov., Glomeropyle cuneum Bragin, sp. nov., G. algidum Bragin, sp. nov., G. aculeatum Bragin, sp. nov., Kahlerosphaera isopleura Bragin, sp. nov., Stauracanthocircus folium Bragin, sp. nov., Tetraspongodiscus uncatus Bragin, sp. nov., Poulpus costatus (Kozur et Mostler), Eonapora robusta Kozur et Mostler, Planispinocyrtis kotelnyensis Bragin, sp. nov., Annulotriassocampe baldii (Kozur), and Pseudoeucyrtis annosus Bragin, sp. nov.; (2) a Middle-Late Carnian assemblage with Pseudostylosphaera glabella Bragin, sp. nov., P. gracilis Kozur et Mock, P. voluta Bragin, sp. nov., P. gelida Bragin, sp. nov., Kahlerosphaera unca Bragin, sp. nov., K. aspinosa Kozur et Mock, K. fuscinula Bragin, sp. nov., K. acris Bragin, sp. nov., Capnuchosphaera kuzmichevi Bragin, sp. nov., C. triassica De Wever, C. angusta Bragin, sp. nov., Sarla intorta Bragin, sp. nov., S. compressa Bragin, sp. nov., S. aequipeda Bragin, sp. nov., Betraccium irregulare Bragin, B. kotelnyensis Bragin, sp. nov., Spongotortilispinus carnicus Kozur et Mostler, S. subtilis Bragin, sp. nov., Dumitricasphaera simplex Tekin, D. aberrata Bragin, sp. nov., D. arbustiva Bragin, sp. nov., Zhamojdasphaera epipeda Bragin, sp. nov., Z. proceruspinosa Kozur et Mostler, Vinassaspongus subsphaericus Kozur et Mostler, Palaeosaturnalis triassicus Kozur et Mostler, Paronaella concreta Bragin, sp. nov., P. aquilonia Bragin, sp. nov., Tetraspongodiscus cincinnalis Bragin, sp. nov., Annulotriassocampe baldii (Kozur), Canoptum zetangense Wang et Yang, Whalenella speciosa (Blome), Syringocapsa turgida Blome, Droltus gelidus Bragin, sp. nov., and D. niveus Bragin, sp. nov.; (3) an Early Norian assemblage with Pseudostylosphaera glabella Bragin, sp. nov., P. gelida Bragin, sp. nov., P. voluta Bragin, sp. nov., Kahlerosphaera retunsa Bragin, sp. nov., Capnuchosphaera deweveri Kozur et Mostler, Sarla globosa Bragin, sp. nov., Palaeosaturnalis mocki Kozur et Mostler, Paronaella aquilonia Bragin, sp. nov., Syringocapsa turgida Blome, and Droltus gelidus Bragin, sp. nov.; (4) a Middle Norian assemblage with Sarla globosa Bragin, sp. nov., Sarla sp., Syringocapsa turgida Blome, Canoptum sp., and Laxtorum? sp. The fact that these assemblages include taxa that are recorded more southerly, including paleotropical localities, and support dating based on mollusks, enables the use of these data for Boreal-Tethyan correlation. The greatest similarity to Tethyan associations is observed in the Late Carnian and Early Norian, that is, the periods of the greatest penetration of thermophilic mollusks into this basin. A total of 69 radiolarian species of 3 orders, 18 families, and 29 genera are described; 44 species are newly described; the stratigraphic and geographical ranges of the majority of taxa are improved considerably.     

New gall midges of the subfamily Lestremiinae (Diptera, Cecidomyiidae) from the Rovno amber

Two new genera and eight new species of the gall midges subfamily Lestremiinae, viz. Berestella insuperabilis gen. et sp. nov., Monardia impellucida sp. nov., Xylopriona aristata sp. nov. (Micromyini), Aprionus improvisus sp. nov., A.? vlaskini sp. nov. (Aprionini), Vicemyia immediata gen. et sp. nov. (Peromyiini), Cordylomyia cauta sp. nov. (Campylomyzini) and Bryomyia necessaria sp. nov. (Bryomyiini) from the Late Eocene Rovno amber are described. New combinations, i.e., Cordylomyia declinata (Fedotova, 2004), comb. nov. and Cordylomyia magnifica (Nel et Prokop, 2006), comb. nov. (both are transferred from Neurolyga) are proposed. No species in common with the fauna from the Baltic amber are recorded.     

A new subfamily of ichneumonids from the Lower Cretaceous of Transbaikalia and Mongolia (Insecta: Hymenoptera: Ichneumonidae)

A new subfamily of ichneumonids, Palaeoichneumoninae, is described from the Lower Cretaceous of Transbaikalia (Baisa locality) and Mongolia (Bon Tsagan and Kholbotu Gol localities). The new subfamily is intermediate between the archaic subfamily Tanychorinae and the Recent Ichneumonidae. It includes 12 new species, described in three new genera: Palaeoichneumon freja gen. et sp. nov., P. ornatus sp. nov., P. danu sp. nov., P. micron sp. nov., P. mirabilis sp. nov., P. tenebrosus sp. nov., P. townesi sp. nov., Rudimentifera mora gen. et sp. nov., R. suspecta sp. nov., Dischysma maculata gen. et sp. nov., D. similis sp. nov., and D. ramulata sp. nov.     

New click beetles (Coleoptera: Elateridae) from the cretaceous of Russia and Kazakhstan

A new tribe, two new genera, and seven new species of click beetles are described: Pollostelaterini, trib. nov., Pollostelater baissensis, gen. et sp. nov. from the Lower Cretaceous of Transbaikalia (Baisa locality), five new species in the genus Cryptocoelus Dolin et Nel C. sinitshenkovae, sp. nov. (Romanovka locality), C. shcherbakovi, sp. nov., C. baissensis, sp. nov., C. lukashevichae, sp. nov., C. dolini, sp. nov. (all from Baisa locality), and Turonelater giganteus, gen. et sp. nov. from the Turonian of Southern Kazakhstan (Kzyl-Dzhar locality).     

Larvae of caddisflies (Insecta: Trichoptera) from the Mesozoic of Siberia

Body fossils of caddisfly larvae are described for the first time based on material from the Lower Cretaceous of Siberia (Baissa locality, Neocomian). The material includes a fully grown larva of Baissoplectrum separatum gen. et sp. nov. (Brachycentridae), Creterotesis coprolithica gen. et sp. nov. (Leptoceridae), and Cretolype minuta gen. et sp. nov., a tiny larva probably belonging to an uncertain family of the suborder Annulipalpia (Hydropsychina). C. coprolithica built unusually soft larval cases of pellets; these cases are described as a separate new indusigenus and indusispecies Coprindusia pallida gen. et sp. nov. Bionomics, taxonomy, and evolution of the Mesozoic larvae of Trichoptera are discussed.     

New gall midge taxa of the subfamilies porricondylinae and lasiopterinae (Diptera,Cecidomyiidae) from the Rovno amber

Five new gall midge taxa of the subfamilies Porricondylinae and Lasiopterinae are described from the Late Eocene Rovno amber: Adsumyia integra gen. et sp. nov. (Dicerurini), Gratomyia inexigentis gen. et sp. nov. (Holoneurini), Winnertzia recusata sp. nov. (Winnertziini), Kovaleviola injusta gen. et sp. nov., and Spungisiola insuperabilis gen. et sp. nov. (Brachineurini).     

New seeds of gymnosperms from the Lower Permian of the Ural mountains

New taxa of detached seeds are described from the Kungurian (Lower Permian) of the Middle Cis-Urals: Laevigatospermum compressum gen. et sp. nov., Hirsutospermum paniculatum gen. et sp. nov., Rugosospermum callosum gen. et sp. nov., and Craspedosperma filiferum sp. nov. The systematic position of the new taxa is discussed.     

First gall midges (Diptera,Cecidomyioidea) from Late Cretaceous amber of the Taimyr Peninsula

Gall midges from Santonian amber of Yantardakh (Taimyr Peninsula) are investigated for the first time. Eight new genera and eight new species belonging to five tribes of the subfamily Micromyinae (Lestremiidae) are described. These are Caputmunda yantardakhica gen. et sp. nov. (Catochini), Cretoperomyia taimyrica gen. et sp. nov. (Peromyiini), Palaeostrobliella dmitrii gen. et sp. nov., Yantardakhiella pusilla gen. et sp. nov., and Zherikhiniella pedicellata gen. et sp. nov. (Strobliellini), Menssana norilsk gen. et sp. nov. (Micromyini), and Cretomycophila ekaterinae gen. et sp. nov., and Corporesana khatanga gen. et sp. nov. (Aprionini). A representative of Porricondylinae (Cecidomyiidae, Porricondylinae, Holoneurini) is recorded. The diagnoses based on morphometric parameters of tribes and previously established genera are emended. The species composition of gall midges from three Cretaceous amber faunas of different ages are analyzed.     

New and little known orthopteroid insects (polyneoptera) from fossil resins: Communication 2

New taxa of the suborder Blattina (order Dictyoptera), possibly belonging to the family Corydiidae (Erucoblatta semicaeca gen. et sp. nov., Miocene; Proholocompsa gen. nov., Eocene; and Holocompsa nigra sp. nov. and H. abbreviata sp. nov., Miocene) and belonging to the family Ectobiidae (Plectoptera electrina sp. nov., Miocene; Agrabtoblatta symmetrica gen. et sp. nov. and ?Symploce rete sp. nov., Pleistocene) are described. The taxonomic position of the enigmatic genus Raphidiomimula Grimaldi et Ross from the Upper Cretaceous is discussed.     

New Emsian foraminifers from of the Karpinsky Horizon limestones Severouralsk Bauxite Mine

Four new foraminiferal species, Parathuramminites mutilatus sp. nov., P. stelliformis sp. nov., P. subrus sp. nov., and P. minutus sp. nov., and a new genus and species, Algaeformis porosus gen. et sp. nov. from clayey limestones and marls of the basal Karpinsky Horizon (Emsian Stage, Lower Devonian) of the Severouralsk Bauxite Mine are described.     

New and little known orthopteroid insects (Polyneoptera) from fossil resins: Communication 3

New taxa of Orthoptera Ensifera are described in the families Mogoplistidae [Protomogoplistes asquamosus gen. et sp. nov. (Upper Cretaceous) in the subfamily Protomogoplistinae subfam. nov. and Archornebius balticus gen. et sp. nov. (Eocene), Pseudarachnocephalus gen. nov., P. dominicanus sp. nov., and P. latiusculus sp. nov. (all Miocene) in Mogoplistinae] and Gryllidae [Eopentacentrus borealis gen. et sp. nov. (Eocene), ?Grossoxipha feminea sp. nov. (Miocene), and Apentacentrus copalicus sp. nov. in the subfamily Pentacentrinae, ?Cyrtoxipha electrina sp. nov. and ?Cyrtoxipha illegibilis sp. nov. (both Miocene) in Trigonidiinae, and Baltonemobius fossilis gen. et sp. nov. (Eocene) in Nemobiinae]. The Miocene genera Proanaxipha Vickery et Poinar and Grossoxipha Vickery et Poinar are transferred from the subfamily Trigonidiinae to Pentacentrinae. P. latoca Vickery et Poinar and Abanaxipha longispina Vickery et Poinar are redescribed; the male of the latter species is described for the first time.     

New radiolarian genera and species from the Lower Permian of the Southern Urals and Northern Mugodzhary

Two new genera and five new species of Lower Permian radiolarians, Arcoclathrata alekseevi gen. et sp. nov., Entactinia pinrasensis sp. nov., Microporosa rozhnovi gen. et sp. nov., M. aktastensis sp. nov., and Somphoentactinia saecularis sp. nov., from the Southern Urals and Northern Mugodzhary are described. The new spherical radiolarian taxa belong to three orders: Entactiniata, Cancelliata, and Spongiata of two classes, Sphaerellaria and Spumellaria.     

New jewel beetles (Coleoptera, Buprestidae) from Cretaceous deposits of Russia

A new genus and two new species of jewel beetles, Cretalbiana sukatshevae gen. et sp. nov. and C. major sp. nov., are described from the Middle Albian (Lower Cretaceous) of Khetana, Khabarovsk Region, Russia.