A new ground beetle (Carabidae, Protorabinae) from the Lower Cretaceous of Inner Mongolia, China

Cretorabus rasnitsynisp. n., belonging to the extinct subfamily Protorabinae of Carabidae, was described based on a well-preserved specimen from the Lower Cretaceous Yixian Formation of Yangshuwanzi, Inner Mongolia. The diagnostic characters forCretorabus are revised, and the key to species of the genus was presented. The fossil record of Protorabinae was summarized. Sinocarabus Hong, 1982 and Obesofemoria Hong, 1982 cannot be attributed to Protorabinae.     

A new species of Tingidae (Insecta: Hemiptera: Heteroptera) from the Lower Cretaceous of Transbaikalia

A new species of lace-bug Sinaldocader ponomarenkoi sp. nov. (Tingidae: Phatnomatini) is described from the Lower Cretaceous Transbaikalian locality Baissa. The new species differs from the type species S. drakei Popov, 1989 from the Lower Cretaceous of Mongolia in the well-developed lateral carinae of pronotal disc and in the details of hemelytral venation.     

Foraminifera diversity changes and paleoenvironmental analysis: the Lower Cretaceous shallow-water carbonates of San Lorenzello, Campanian Apennines, southern Italy

A high-resolution stratigraphic study, carried out on the carbonate platform strata of the San Lorenzello section (Matese Mountains, southern Italy), Valanginian–Hauterivian in age, has allowed to: recognise lithofacies and their associations; assign the lithofacies associations to specific environments and individuate early meteoric diagenetic modifications, recurring at specific horizons. In this frame the vertical variation of benthic foram diversity has been analysed. On the whole, foraminiferal genera diversity decreases from open to restricted marine environments. Moreover, a climatic control on carbonate sedimentation is suggested by a Milankovitch cyclicity organised in elementary cycles, bundles and superbundles as well as by diagenetic characteristics testifying that humid and arid conditions alternated during the Early Cretaceous times. The orbital cyclicity is also documented by foraminiferal diversity changes, even if some discrepancies between the lithofacies and the diversity locally occur. Therefore, the above diversity changes do not appear to provide sufficient information for the sequence-stratigraphic interpretation of shallow-water carbonates.     

A new subfamily of ichneumonids from the Lower Cretaceous of Transbaikalia and Mongolia (Insecta: Hymenoptera: Ichneumonidae)

A new subfamily of ichneumonids, Palaeoichneumoninae, is described from the Lower Cretaceous of Transbaikalia (Baisa locality) and Mongolia (Bon Tsagan and Kholbotu Gol localities). The new subfamily is intermediate between the archaic subfamily Tanychorinae and the Recent Ichneumonidae. It includes 12 new species, described in three new genera: Palaeoichneumon freja gen. et sp. nov., P. ornatus sp. nov., P. danu sp. nov., P. micron sp. nov., P. mirabilis sp. nov., P. tenebrosus sp. nov., P. townesi sp. nov., Rudimentifera mora gen. et sp. nov., R. suspecta sp. nov., Dischysma maculata gen. et sp. nov., D. similis sp. nov., and D. ramulata sp. nov.     

Non-marine biota from the Lower Permian of the central Southern Alps (Orobic and Collio basins, N Italy): a key to the paleoenvironment

The Lower Permian in the central Southern Alps yields an important low-diversity fossil assemblage which was deposited in a varied continental setting, showing mainly alluvial fan to lacustrine and, locally, playa-like floodplain environments. The present study is not taxonomical and its objective is to make a first report of new invertebrate organisms and trackways discovered in the Orobic and Collio basins. Such a fossil record, which comprises freshwater jellyfishes, arthropod tracks, stromatolites, algae and other organisms, will further our knowledge about the local and regional geological history of the central-western Southern Alps and improve our understanding of Early Permian palaeoenvironments. On the whole, in both intramontane basins the ichnodiversity and fossil content decreases from a stratigraphic lower portion, mainly lacustrine and alluvial, towards an upper one, characterised by coarse alluvial deposits to floodplain fines. On the basis of both the already known palaeontological data from the two basins, mainly macroflora and tetrapod footprint associations, and the recently discovered taxa, we tried to make reliable palaeoenvironmental inferences and, possibly, hypothesise on a climatic change, which could have occurred during the Early Permian.     

Basal titanosauriform (Dinosauria, Sauropoda) teeth from the Lower Cretaceous Yixian Formation of Liaoning Province, China

The Yixian Formation of Liaoning Province, People's Republic of China has yielded a diverse fauna of non-avian dinosaurs, but is dominated by small-bodied taxa. Here, we describe a series of isolated teeth from the Lujiatun Beds of the formation that are referable to a basal titanosauriform sauropod. Some of the teeth possess a distinctive circular boss on the lingual surface, which suggests that they are referable to cf. Euhelopus sp. This identification provides some additional support for biostratigraphical correlations between the Jehol Group and the Mengyin Formation of Shandong Province that suggest an Early Cretaceous age for the latter unit. Moreover, the titanosauriform affinities of the teeth provide further evidence for the dominance of this sauropod clade in eastern Asia during the Cretaceous.     

A new family of aphids (Hemiptera, Sternorrhyncha) from the Lower Cretaceous of Baissa, Transbaikalia

The family Rasnitsynaphididaefam. n. has a unique combination of characters: 9-segmented antennae; rhinaria arranged in many transverse rows, surrounding the antennal segments; segment IX narrower than other segments of flagellum, always without rhinaria; cubitus branches separated; ovipositor present; siphuncular pores absent. The new family comprises the genus Rasnitsynaphisgen. n. with three species, Rasnitsynaphis ennearticulatasp. n., Rasnitsynaphis coniunctasp. n., and Rasnitsynaphis quadratasp. n., all from the Lower Cretaceous of Transbaikalia.     

Representatives of the family Bochianitidae (Ammonoidea) from the Lower Cretaceous of the Crimean Mountains

The heteromorph ammonites of the family Bochianitidae from the Lower Cretaceous of the Crimean Mountains are revised. The validity of the genus Janenschites, separated from the genus Bochianites is confirmed. The species Bochianites neocomiensis (d’Orbigny), B. goubechensis Mandov, B. levis sp. nov. and B. crymensis sp. nov. are described from the Berriasian and the species Janenschites oosteri (Sarasin et Schöndelmayer) and J. incisus sp. nov. are described from the Lower Barremian. The family Bochianitidae first appeared at the beginning of the Berriasian in the southern regions (Africa and the Crimea), and spread to the northern regions of western Europe in the Valanginian-Hauterivian.     

The oldest psyllipsocid booklice, in Lower Cretaceous amber from Lebanon (Psocodea, Trogiomorpha, Psocathropetae, Psyllipsocidae)

Libanopsyllipsocus alexanderasnitsynigen. et sp. n., of Psyllipsocidae is described and figured from the Lower Cretaceous amber of Lebanon. The position of the new taxon is discussed and the fossil is compared to other psyllipsocids. The species represents the earliest record of the family Psyllipsocidae.     

First record ofColoborhynchus (Pterosauria) from the Santana Formation (Lower Cretaceous) of the Chapada do Araripe, Brazil

The anterior tips of associated upper and lower jaws of a pterosaur from the Lower Cretaceous of Brazil are described and assigned to the taxonColoborhynchus in the family Ornithocheiridae. It is characterized by the shape and position of the sagittal crest on the upper and lower jaw, the arrangement and length of the teeth and the spoon-like lateral expansion of the anterior parts of the jaws. It closely resemblesColoborhynchus wadleighi from North America andColoborhynchus clavirostris from England. Diagnostic anatomical characteristics permit a revision of the genusTropeognathus, which is shown here to be a junior synonym of other described taxa.Tropeognathus mesembrinus is referred toCriorhynchus andT. robustus toColoboryhnchus. Consistent anatomical features enable the new jaw fragment to be assigned toColoborhynchus robustus.      

Two new dinosaur tracksites from the Lower Cretaceous Jiaguan Formation of Sichuan Basin,China: specific preservation and ichnotaxonomy

Two new dinosaur tracksites are reported from the Lower Cretaceous Jiaguan Formation in the Sichuan Basin, Qijiang District of Chongqing. These are the Gaoqing-Yongsheng and the Huibu tracksites, which represent the 13th and 14th reports from this formation. The Gaoqing-Yongsheng tracksite reveals the trackway of a large biped (ornithopod) in association with isolated sauropod tracks and large indeterminate undertracks with radial cracks. These features are preserved as natural casts with pebble infillings in a coarse, cross bedded and very thick bedded sandstone sequence. The Huibu tracksite reveals isolated theropod tracks and ornithopod tracks, the latter having a quadripartite, Caririchnium-like morphology, preserved in a thin bedded sandstone sequence with intercalated mudstone.     

Unusual deinonychosaurian track morphology (Velociraptorichnus zhangi n. ichnosp.) from the Lower Cretaceous Xiaoba Formation,Sichuan Province,China

A new Lower Cretaceous dinosaur footprint locality named the Mujiaowu tracksite in the Xiaoba Formation, Sichuan Province, has yielded a new assemblage containing the didactyl deinonychosaurian ichnogenus Velociraptorichnus. This is the eleventh report in the global record, the seventh from Asia and the fifth from China. All, except for an isolated report from Europe, occur in Lower Cretaceous deposits. Unlike previous reports of the ichnogenus, the Mujiaowu site has yielded both didactyl Velociraptorichnus tracks and tridactyl tracks which we interpret as a different expression of this same ichnogenus, caused by registration of digit II, either due to special substrate conditions or to less claw retraction. These tridactyl Velociraptorichnus footprints are assigned to the new ichnospecies Velociraptorichnus zhangi. Such tridactyl deinonychosaurid footprint morphology is predictable, but based on current ichnological evidence appears to be the exception rather than the rule.     

Relationships between plant communities and late snow melting on Mount Prado (Northern Apennines,Italy)

Relationships between the spatial and temporal distribution of long-lasting snow-cover and the spatial distribution of plant communities above timberline were studied on Mount Prado (2054 m), a representative sample area of the summit vegetation of the Northern Apennines (Italy). The spatial analysis was carried out by creating a geographic information system. Vegetation map and two snow-cover maps were both georeferenced to a scale 1 : 2000. The vegetation map is formed by 28 plant communities (including 7 combinations of communities, or vegetation mosaics) distributed into 277 map polygons. The maps were compared by an overlay procedure. The results show that the six plant communities (including three mosaics) which have a June snow-cover higher than 60% are diversified by a snow-melting gradient from early June to mid-July. The slowest snow melt corresponds to a snow-bed community (Salicetum herbaceae) and to a mesophytic grassland (Trifolium thalii-Festuca puccinellii community, Luzula alpino-pilosa variant).     

Lower Cretaceous dinoflagellate cyst and acritarch stratigraphy of the Cismon APTICORE (Southern Alps, Italy)

A pelagic sedimentary succession, virtually complete from the Upper Hauterivian to the Upper Aptian and unconformably overlain by the Middle-Upper Albian p.p., was continuously cored in the Belluno Basin (southern Alps, NE Italy) as part of the APTICORE Program. APTICORE at Cismon Valley penetrated 131.8m of limestones, marls and black shales, with 100% recovery of good quality cored material.One hundred and forty-six samples recovered from the marl and shale beds of the Cismon core were processed and analyzed for palynomorphs. Most of them yielded relatively rich and fairly well preserved assemblages of marine and terrestrially-derived palynomorphs.The results of a qualitative study of dinoflagellate cysts and acritarchs are presented and discussed. The distributions of 150 taxa are tabulated against the chronostratigraphy independently established on the basis of original litho-, bio-, chemo-, magnetostratigraphic investigations and of correlations with extensively studied sections outcropping in the vicinity of the Cismon drill site.The acritarch Pinocchiodinium erbae gen. et sp. nov. is described. Due to its distinctive morphology and extremely constant occurrence also in the black shales of the Selli Level, it is proposed as a marker species for the Aptian sediments of the Tethys.The dinoflagellate cysts Kallosphaeridium dolomiticum sp. nov. and Nexosispinum hesperus brevispinosum subsp. nov. are described from the Upper Hauterivian. Additional taxonomic remarks are made about other dinoflagellate cyst species, including the emendations of Tanyosphaeridium magneticum Davies 1983 and Bourkidinium granulatum Morgan 1975.The biostratigraphic value of selected taxa is discussed and compared with data known both from the Tethyan and Boreal realms. In particular, the extinction of Bourkidinium granulatum emend. is proposed as the best dinoflagellate cyst event for the delimitation of the Hauterivian-Barremian boundary in the Northern Hemisphere. The first appearance datums of Prolixosphaeridium parvispinum and Odontochitina operculata, and the slightly younger last appearance datum of Nexosispinum vetusculum are confirmed as useful biohorizons for recognition of the lower part of the Upper Barremian and hence for the approximation of the Lower-Upper Barremian boundary. The last occurrences of Rhynchodiniopsis aptiana and Phoberocysta neocomica are calibrated in the basal Aptian.     

The oldest members of the families Aeolothripidae and Thripidae (Insecta: Thysanoptera) from the Lower Cretaceous of Transbaikalia

Fusithrips crassipes Shmakov, gen. et sp. nov. (Aeolothripidae) and Convexithrips robustus Shmakov, gen. sp. nov. (Thripidae), the oldest members of these families, are described from the Lower Cretaceous of the Baissa locality. This allowed the improvement of the time of emergence of the Recent Thysanoptera assemblage, which includes phloeothripids along with aeolothripids and thripids.     

Biostratigraphy of Lower Cretaceous microfossils from the Araripe basin, northeastern Brazil

A biostratigraphic study was carried out in the Lower Cretaceous Araripe basin, northeastern Brazil, allowing the recognition of several chronostratigraphic units: the Dom João (Jurassic?-Lower Cretaceous?), the Rio da Serra (Neocomian) and the Alagoas (Aptian/Albian) local stages. For the first time a large hiatus between the Rio da Serra and Alagoas local stages is carefully documented. The palynomorphs and the ostracode associations throughout the Jurassic?-Aptian/Albian sequence allow the interpretation of the paleoenvironmental evolution of the Araripe basin which otherwise confirms that a polycyclical sedimentation occurred in the basin, being one of the controlling factors on the distribution of ostracodes and palynomorphes.     

The palaeoecology of the dinosaurs of the Wessex Formation (Wealden Group, Early Cretaceous), Isle of Wight, Southern England

The Wessex Formation on the Isle of Wight yields an Early Cretaceous dinosaur fauna. Sedimentological evidence shows that this represents a mosaic of fluvial, floodplain and lacustrine environments within a relatively narrow east-west oriented valley. The vegetational cover on the alluvial plain had a savannah-or chaparral-like aspect, probably of low productivity. The relative scarcity of small aquatic vertebrates, absence of coals, abundance of oxidixed sediments and the presence of immature calcretes indicate seasonal water supply. The dinosaur taxa compising the Wessex Formation faunal assemblage represent a single palaeocommunity which inhabited the local alluvial plain, although some species may have been transient. The fauna had a relatively low diversity and this is attributed to the low productivity of the local vegetation. Iguanodontids and Hypsilophodon were the dominant elements in the fauna. In contrast to Late Jurassic dinosaur faunas, sauropods are less abundant in the Wessex Formation, although they remain taxonomically diverse. It is concluded that climatic changes which took place in the Late Jurassic or Early Cretaceous resulted in the appearance of low productivity vegetation and that this was incapable of supporting large sauropod populations.     

Asperula calabra (Rubiaceae) and allied taxa in southern Apennines,Italy

The present work is aimed at studying several Asperula (Rubiaceae) sect. Cynanchicae populations in southern Apennines, Italy, with particular reference to those referred to A. calabra, by employing biometrical methods on macromorphological data. Among other historical misapplications, A. cynanchica subsp. cynanchica seems to be very rare or missing in southern Italy. The enigmatic Asperula calabra, confirmed to be limited to a single mountain population in Calabria, appears to be strictly related to A. aristata subsp. scabra, so we suggest to treat it as a further subspecies of A. aristata. This latter species is otherwise distributed throughout southern Italy with subsp. scabra and subsp. aristata, characterised by slight morphological differences, which are correlated to altitudinal ranges.