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1.
The main patterns of expression of the morphological polyvariance of plants are discussed: diversity of biomorphs, pathways of ontogeny, and disturbances of morphogenesis. The diversity of biomorphs of tap root plants in different ecological conditions has been analyzed in detail. Promising directions of future studies have been formulated. 相似文献
2.
Abstract: Many articulated brachiopods experience marked life habit variations during ontogeny because they experience their fluid environment at successively higher Reynolds numbers, and they can change the configuration of their inhalant and exhalant flows as body size increases. We show that the extant brachiopod Terebratalia transversa undergoes a substantial ontogenetic change in reorientation governed by rotation around the pedicle. T. transversa′s reorientation angle (maximum ability to rotate on the pedicle) decreases during ontogeny, from 180 degrees in juveniles to 10–20 degrees in individuals exceeding 5 mm, to complete cessation of rotation in individuals larger than 10 mm. Rotation ability is substantially reduced after T. transversa achieves the adult lophophore configuration and preferred orientation with respect to ambient water currents at a length of 2.5–5 mm. We hypothesize that the rotation angle of T. transversa is determined mainly by the position of ventral and dorsal points of attachment of dorsal pedicle muscles relative to the pedicle. T. transversa shows a close correlation between the ontogenetic change in reorientation angle and ontogeny of morphological traits that are related to points of attachment of dorsal pedicle muscles, although other morphological features can also limit rotation in the adult stage. The major morphological change in cardinalia shape and the observed reduction of rotation affect individuals 2.5–10 mm in length. The position of ventral insertions of dorsal pedicle muscles remains constant, but contraction of dorsal pedicle muscles is functionally handicapped because dorsal insertions shift away from the valve midline, rise above the dorsal valve floor, and become limited by a wide cardinal process early in ontogeny (<5 mm). The rate of increase of cardinal process width and of distance between dorsal pedicle muscle scars substantially decreases in the subadult stage (5–10 mm), and most of the cardinalia shell traits grow nearly isometrically in the adult stage (>10 mm). T. transversa attains smaller shell length in crevices than on exposed substrates. The proportion of small‐sized individuals and population density is lower on exposed substrates than in crevices, indicating higher juvenile mortality on substrates prone to grazing and physical disturbance. The loss of reorientation ability can be a consequence of morphological changes that strengthen substrate attachment and maximize protection against biotic or physical disturbance (1) by minimizing torques around the pedicle axis and/or (2) by shifting energy investments into attachment strength at the expense of the cost involved in reorientation. 相似文献
3.
《Annales de Paléontologie》2023,109(2):102612
An old find of a graptolite from the Eifelian Jemelle Formation, combined with the recent discovery of an exceptionally well-preserved specimen from the Upper Frasnian (Matagne Formation), allow documentation of the geologically oldest and youngest dendroid fossils from the Devonian of Belgium. Both are ascribed to the long-ranging genus Callograptus (Acanthograptidae). These records enlarge the sparse knowledge of graptolite diversity and occurrences along the southern margin of Laurussia during the mid-Paleozoic. In Belgium, Devonian dendroids are exclusively known from the Dinant Synclinorium. The Devonian dendroid faunas previously reported from the Eifelian, Givetian and Frasnian of the Ardenne–Rhenish massifs (Belgium and Germany) are of low diversity (one to five species per locality). The specimens, generally inedaquatelly preserved, were ascribed to the genera Callograptus, Dictyonema, Palaeodictyota?, and Ruedemannograptus? In these massifs, Carboniferous dendroids are only known from the Viséan (Moliniacian) Denée Konservat-Lagerstätte. Other but undocumented reports of dendroids (and rhabdopleurids) are from the historical type area of the Tournaisian Stage (Tournai area, Brabant Parautochthon). Rhabdopleuridae from the Viséan–Serpukhovian of the Campine Basin (Turnhout borehole) remain incompletely known in the absence of the previously described material. 相似文献
4.
《Geobios》2019
The fauna and facies of the Hiswah Formation in southern Jordan were investigated. The formation indicates deposition under low energy, open-marine, siliciclastic shelf conditions and reflects the maximum of the first regional post-Cambrian marine transgression on the northern edge of the Arabian-Nubian Shield. This eustatic signal accompanied by a maximum flooding surface can be traced over the entire Arabian Plate and Turkey. The lower part of the Hiswah Fm. consists of pelagic mudstones with only occasional thin beds of rippled, hummocky cross-bedded siltstone and mudstone concretions, and contains a distinct fauna of low diversity. The upper part of the formation is characterized by sandstone-siltstone alternations with some hummocky cross-stratification in its lower portion and trough cross-bedding and ripple marks above indicating somewhat shallower conditions. A very detailed re-investigation of the fauna from the lower part of the Hiswah Fm. and extensive new fossil material indicates that the most remarkable faunal element is the planktonic graptolite Didymograptus murchisoni (Beck in Murchison, 1839in) which, in the past, has been determined erroneously as D. bifidus (Hall in Berry, 1962). Numerous specimens of small obolid brachiopod Palaeoglossa sp. cf. P. attenuata (Sowerby, 1839) represent the most common benthic element within the Hiswah fauna. The exceptional arthropod Hanadirella cf. bramkampi El-Khayal, 1985, is reported for the first time from Jordan, representing its fifth known region of occurrence in addition to Saudi Arabia, Algeria, Spain, and France. For the first time, cephalopod remains are described from the Ordovician of Jordan by the occurrence of ?Dideroceras sp. and a second, indeterminate species. The biostratigraphic age of the Hiswah fauna is revised to late Darriwilian (Dw 3, upper Middle Ordovician) according to our re-investigation of the graptolites. The Hiswah fauna represents the oldest body fossils of the Ordovician marine sedimentation in southern Jordan and is palaeogeographically strongly related to northern and northeastern Saudi-Arabia and to high/mid-latitude Gondwana margins in general. 相似文献
5.
Floréal Solé Jocelyn Falconnet Laurent Yves 《Zoological Journal of the Linnean Society》2014,171(4):878-917
We describe six proviverrine species from the Early Eocene of France. Three species are new: M inimovellentodon russelli sp. nov. from Mutigny [mammal palaeogene (MP)8 + 9], B oritia duffaudi sp. nov. from La Borie (MP8 + 9), and L eonhardtina godinoti sp. nov. from Grauves (MP10). We describe new specimens and propose new generic combinations for three species from MP10: Protoproviverra palaeonictides, Matthodon menui, and Oxyaenoides lindgreni. We also propose a new generic combination for the primitive Eoproviverra eisenmanni (MP7). Matthodon menui was previously considered as a possible oxyaenodontan, but the new fossils clearly support its reference to Hyaenodontida. Leonhardtina godinoti and Ma. menui are the oldest occurrences for these genera, which were previously unknown before the Middle Eocene. Moreover, the discovery of the proviverrine Mi. russelli in Mutigny implies that the Proviverrinae dispersed in Northern Europe between biozone Palaeocene‐Eocene (PE) III (Abbey Wood) and biozone PE IV (Mutigny). This also supports a homogenization of the European faunas during the Early Eocene. The dispersal is concomitant with the disappearance of the oxyaenodontans, arfiines, and sinopines (Hyaenodontida) from Europe. The proviverrines may have filled the ecological niches left vacant by the disappearance of the other carnivorous mammals. With 20 genera and over 30 species, proviverrines were successful in Europe. We performed the first phylogenetic analysis comprising almost all the Proviverrinae. Our analyses indicate that the Proviverrinae diversified greatly during the Early Eocene Climatic Optimum and show a general trend towards specialization throughout the Eocene. © 2014 The Linnean Society of London 相似文献
6.
《Geobios》2020
Devonian and Carboniferous dendroid graptolites from Belgium are evaluated and partly revised. New finds in two different stratigraphic intervals of the ‘Carrière de Lompret’, an active quarry exploiting Frasnian limestones and shales east of Frasnes-lez-Couvin, allow the identification of Callograptus sp. and Dictyonema fraiponti, both belonging to the dendroid family Acanthograptidae. The relatively high diversity of the dendroid graptolite fauna from the Viséan Marbre noir de Denée, one of the few Carboniferous graptolite faunas in the world, can be shown to be based on astogenetic and preservational aspects. Nearly all known specimens can be included in the highly variable Dictyonema fraiponti, a fan-shaped large dendroid species with complex stipes formed from tubular thecae, possessing simple to complex bridges connecting adjacent stipes. Some of the graptolite material is well preserved and provides important information on the tubarium construction of Devonian to Carboniferous dendroid graptolites and, thus, is highly significant for a taxonomic and phylogenetic understanding of the youngest dendroid graptolite faunas worldwide. The genera Callograptus and Ptiograptus are revised based on their type species (Callograptus elegans from Quebec, Canada; Ptiograptus percorrugatus from the Silurian or Devonian of Kentucky, USA) and referred to the Acanthograptidae. 相似文献
7.
David A. T. Harper S. Henry Williams 《Lethaia: An International Journal of Palaeontology and Stratigraphy》2002,35(1):71-78
At Yewdale Beck in northern England, a brachiopod fauna dominated by species of Hindella, Kinnella, Mirorthis, Paromalomena and Plectothyrella , occurs interbedded with a graptolite fauna which includes Akidograptus ascensus , Atavograptus ceryx , Persculptograptus parvulus and Normalograptus spp. (including probable examples of N. normalis , N. angustus and N. medius ) within the lower part of the Skelgill Formation. This suggests that taxa of the terminal Ordovician Hirnantia fauna occur within the lower P. acuminatus Biozone, representing the youngest documented occurrence of the Hirnantia brachiopod fauna. Biostratigraphy, brachiopods, graptolites, Ordovician-Silurian boundary. 相似文献
8.
Late Tremadoc to early Arenig graptolite faunas of southern Bolivia and their implications for a worldwide biozonation 总被引:6,自引:0,他引:6
Jörg Maletz Sven O. Egenhoff 《Lethaia: An International Journal of Palaeontology and Stratigraphy》2001,34(1):47-62
The graptolite faunal succession of the late Tremadoc to early Arenig in southern Bolivia is established from the Cieneguillas and Culpina sections and compared with faunas from other regions. The succession yields important data for the discussion on the proposed GSSP at the base of the Tetragraptus approximatus Zone and the completeness of the successions in eastern North America and Scandinavia. The graptolite faunas of the Aorograptus victoriae, Kiaerograptus supremus, Araneograptus murrayi, Hunnegraptus copiosus and Tetragraptus phyllograptoides zones are reviewed and the faunal associations discussed. The Kiaerograptus supremus Zone is regarded as a local Scandinavian biozone. 相似文献
9.
特马豆克阶是奥陶系底部第一个阶,笔石是特马豆克阶高分辨率地层划分与对比的重要化石类群。江南斜坡带是我国早奥陶世特马豆克期漂浮笔石分异度和丰度最高的相区之一,位于该区的湖南益阳南坝剖面,发育有完整的上特马豆克阶笔石地层,特马豆克阶-弗洛阶界线附近地层连续,上特马豆克阶笔石地层研究已取得较大进展,但下特马豆克阶地层缺乏系统研究。近年来,通过对该剖面笔石标本的不间断采集,新识别出下特马豆克阶笔石带Rhabdinopora flabelliformis parabola带。到目前为止,湖南益阳南坝剖面特马豆克阶可以识别出5个笔石带,自下而上依次为:Rhabdinopora flabelliformis parabola带、Adelograptus tenellus带、Aorograptus victoriae带、Araneograptus murrayi带以及Hunnegraptus copiosus带。基于目前已识别出的笔石带,参考国内外同期笔石地层资料,本文详细讨论华南特马豆克期笔石带序列,并与国内外同期地层进行精确对比。 相似文献
10.
Anna Kozowska-Dawidziuk 《Lethaia: An International Journal of Palaeontology and Stratigraphy》2001,34(1):84-96
A new retiolitid genus Cometograptus, restricted to the lundgreni Biozone (Late Wenlock), is characterized by a combination of primitive features such as strong transverse rods and ancora sleeve lists with seams facing out, with advanced features such as a long sicula, central, free nema and pustulose bandages. Cometograptus links two Subfamilies of the Family Retiolitidae. The Subfamily Retiolitinae is divided into redefined lineages: Pseudoretiolites, Retiolites, Paraplectograptus, whereas the Subfamily Plectograptinae is divisible into the Sokolovograptus (questionable), Gothograptus, Spinograptus, Plectograptus lineages, and Cometograptus. The Sokolovograptus group, the first retiolitids bearing ancora sleeve lists with seams facing out, did not survive the lundgreni crisis. The only survivors were Upper Wenlock retiolitids of the Gothograptus lineage. The new genus Cometograptus includes Cometograptus koreni n.gen. and n.sp., Cometograptus tomczyki n.gen. and n.sp., as well as the previously described Cometograptus nevadensis, Cometograptus marsupium and Cometograptus apoxys. 相似文献
11.
STEVEN M. HOLLAND 《Palaeontology》2010,53(6):1237-1254
Abstract: Using Whittaker’s concepts of alpha, beta, and gamma diversity, Sepkoski asked how global diversity was assembled at scales ranging from the community to the province. In the years since, ecologists have recast diversity in terms of additive partitions where total diversity can be decomposed into sample‐level alpha diversity plus the sum of a series of beta diversity terms that reflect progressively larger spatial scales. Given that marine alpha diversity represents a tiny fraction of global diversity, Phanerozoic global diversity patterns must be dominated by changes in beta diversity at one or more scales. A ballooning ecological literature demonstrates wide variation in beta diversity among ecosystems, regions, and taxa, suggesting that large changes in beta diversity on evolutionary timescales are likely. But the question is which scales are the most important. Several recent palaeontological studies help to constrain beta diversity within sedimentary basins, and the emergence of sample‐based databases puts an answer to Sepkoski’s question within reach. A new method for calculating diversity partitions for richness is introduced, which allows the calculation of each species’ contribution to alpha and beta diversity, as well as the contribution of each sampling unit to beta diversity. 相似文献
12.
对贵州遵义高桥大角寺剖面的奥陶系湄潭组下段和中段笔石的研究结果表明,湄潭组下段从下而上可以识别出Didymograptellus eobifidus带、Corymbograptellus deflexus带和Azygograptus suecicus带;中段含有一层笔石,缺少特征属种,根据其总体面貌,可能相当于Expansograptus hirundo带。文章描述其中11属16种:Pseudophyllograptus angustifolius(Hall),Phyllograptus claviger(Monsen),P.annaHall,Didymograptelluseobifidus(Chen and Xia),D.protoindentus(Monsen),Acrograptus kurcki(T rnquist),Corymbograptus vacil-lans(Tullberg,1880),C.deflexus(Elles and Wood),Expansograptus extensus(Hall),E.nitidus(Hall),Xi-phograptus lofuensis(Lee),X.cf.lofuensis(Lee),Azygograptus suecicusMoberg,Dictyonemasp.,Den-drograptussp.,Inocaulis plumulosusHall。这一笔石动物群分异度较低,大多数为上扬子区的常见属种,代表当时黔中古陆北缘近岸海洋环境中的笔石动物群面貌。 相似文献
13.
Queues, in which individuals inherit resources in a predictable, temporally stable order, are widespread in animal social groups. We develop an analytic model to explore the effect of differential survivorship on the stability of a reproductive queue. We show that unless fighting for dominance is potentially fatal, future direct benefits are not alone sufficient to stabilize a queue of non-relatives under constant (age-independent) mortality rates, regardless of whether a dominant becomes an isolate or remains a dominant on the death of the first subordinate. In the absence of fatal fighting, stabilization of such a queue by future direct benefits alone requires either the dominant or the subordinate to have age-dependent mortality rates. Even when the queue is stabilized by present direct reproduction, however, the shape of the lifespan distribution can make a significant difference to the size of the required incentive. In contrast to non-relatives, queues of relatives can be stable without age-dependent mortality, so long as relatedness exceeds a critical value; however, age-dependent mortality can lower this critical value. 相似文献
14.
The lower levels of the Lipeón Formation, in the Eastern Cordillera, north-west Argentina, yield a marine-dominated palynomorph assemblage, together with graptolites of mid to late mid Llandovery age (Demirastrites convolutus and probably Stimulograptus sedgwickii zones). The palynomorph assemblage is dominated by acritarchs, but also contains algae and terrestrial cryptospores. Crassiangulina variacornuta, considered a potentially good global biostratigraphical marker for the Upper Llandovery is recovered for the first time from the Silurian of Argentina. The occurrence of this species in strata not younger than late Aeronian, and independently dated by graptolites, indicates an early first appearance for Crassiangulina variacornuta, in the Lipeón Formation, below the Aeronian/Telychian boundary. The lower part of the unit corresponds to a quiet marine environment; thus supporting that Crassiangulina variacornuta is a facies-sensitive acritarch. 相似文献
15.
JÖRG MALETZ 《Palaeontology》2010,53(2):415-439
Abstract: The virgellar spine is one of the most consistent features of the graptolite sicula. It is present in a large number of graptoloid groups, but evolved separately and independently in these as it is seen from the presence of the spine in either ventral (Axonophora) or dorsal (Phyllograptus, Xiphograptus) position. The evolution of the virgellar spine in the Pan‐Bireclinata in the Upper Dapingian to Lower Darriwilian time interval is known to follow four main steps, from a simple rutellum, through a lamelliform rutellum and a lanceolate virgella to the true virgellar spine. For the xiphograptids and in Phyllograptus, the origin and early development is less well documented, but appears to follow a similar path. However, the individual stages are condensed, and a true virgellar spine emerges already in the Floian time interval. A virgellar spine was found in Didymograptellus bifidus, necessitating a revision of the diagnosis of the genus Didymograptellus. A number of species of the virgellate genera Xiphograptus, Yutagraptus and Didymograptellus are described from isolated material for the first time. The species are useful for the biostratigraphic correlation of endemic mid‐continent North American faunas with the Pacific Type faunal realm. Xiphograptus artus sp. nov., Didymograptellus primus sp. nov. and Didymograptellus cowheadensis sp. nov. from the Cow Head Group of western Newfoundland are described as new. 相似文献
16.
Bennett Dyke Timothy B. Gage Patricia L. Alford Brent Swenson Sarah Williams-Blangero 《American journal of primatology》1995,37(1):25-37
Mortality statistics from three captive populations of chimpanzees (Pan troglodytes) were combined to generate standard model life tables for each sex in this species. The model is compared to an estimate of survivorship of a group of wild animals, and is applied to an incomplete data set to illustrate how the model may be used to extend estimates of mortality statistics to missing older ages. © 1995 Wiley-Liss, Inc. 相似文献
17.
18.
Abstract. This research tests the hypothesis that Australia and eastern North America met in rotational collision during the Palaeozoic with the corollary that the New England fold belt of Australia, rather than the Reguibat promontory of Africa, collided with the Alleghanian orogen in the central Appalachians. Identical Lancefieldian-stage, zone 1 (Early Ordovician) graptolites of the Anisograptid family are found in identical environments in Newfoundland, Norway, and the Lachlan fold belt of Australia. Palaeozoic granites are consistent with a tectonic model in which the Lachlan fold belt caused mechanical deformation of the Canadian Appalachians. The Lachlan and New England fold belts of Australia and the Alleghanian orogen of North America are tectonically consistent with the east coast hypothesis. Major deformations and magmatic episodes are coeval from Silurian to Permian. The tectonic, palaeontologic, lithologic, and geometric evidence for this position is more abundant and precise than the stratigraphic evidence for a west coast location of Australia relative to North America. 相似文献
19.
Stig M. Bergström Charles E. Mitchell 《Lethaia: An International Journal of Palaeontology and Stratigraphy》1986,19(3):247-266
The North American Upper Ordovician reference standard, the Cincinnatian Series, contains rich shelly microfossil faunas in its type area in the Cincinnati Region but graptolites are uncommon in most of its shallow-water calcareous sediments. Consequently, the graptolite correlation of this key sequence has remained uncertain, in part, even controversial. A review of both previously published recently discovered graptolite Occurrences in the type Cincinnatian, combined with data from the important graptolite successions in Oklahoma New York-Quebec, has not only clarified the graptolite correlation of the Cincinnatian but also added new data on the morphology taxonomy, the vertical horizontal distribution, of several taxa. The information now at hand indicates that the Edenian Stage correlates with the C. spiniferus Zone, the Maysvillian Stage with the C. pygmaeus lower middle P. manitoulinensis Zone, the Richmondian Stage with the upper P. manitoulinensis , the D. complanatus , possibly part of the C. inuiti Zone. Comparison between graptolite conodont biostratigraphic evidence reveals no apparent conflict. Correlations are proposed between Upper Ordovician North American stages, graptolite conodont zones, successions in Texas, Oklahoma, Sweden, European graptolite zones, British series. 相似文献
20.
We report a change in the method of averaging parameters of the Siler mortality model fitted to a series of Old World monkey survivorship schedules. Substitution of geometric for arithmetic means of multipliers of exponential terms avoids incorrect weighting of parameters. These changes have minor effects on the composite standard mortality schedule for these animals originally described in 1988. © 1993 Wiley-Liss, Inc. 相似文献