Foods of the south polar skua at Hop Island,Rauer Group,East Antarctica

Summary Pellets regurgitated by south polar skuas at Hop Island, in the Rauer Group, East Antarctica, Princess Elizabeth Land were collected and analysed. Individual collections were grouped in relation to the nearest breeding colony of seabirds, but irrespective of their source the frequencies of occurrence of the food items identified in pellets indicated that Adélie penguins were of varying, but major (29%–72%), importance to skuas inhabiting the Island. Though the incidence of Antarctic petrel remains (from adult or chick) was generally low in pellets (<7%), the occurrence of remains of southern fulmars was high (>27%) at all sites near the fulmar breeding colonies. Fish remains, and beaks of cephalopods, were present in few pellets. Whilst indicating the range of foods eaten on Hop Island, and perhaps their relative importance, it is considered that the pellets by themselves do not reflect accurately the level of predation of eggs of Antarctic petrels and southern fulmars, nor do the pellets necessarily include the remains of young of these species.     

Late Holocene paleofish populations in Effingham Inlet, British Columbia, Canada

Paleontological studies of fish remains in laminated sediments provide a proxy relationship between fish populations and ocean climate. This study examines climate variability from approximately 500 y B.P. to 4000 y B.P. as recorded in fish remain abundances (primarily scales) collected from laminated sediments within Effingham Inlet on the west coast of Vancouver Island, British Columbia. The study also discusses technical issues involving fish remains from laminated sediments, including sampling resolution, sediment volume, identification of remains and appropriate abundance considerations.The dominant species in the assemblage include Northern anchovy and Pacific herring, with lesser percentage-abundances from rockfish, hake, elasmobranches and surfperch. The data indicate that Northern anchovy experienced a shift in scale deposition abundance at approximately 2800 y B.P. with not only a greater total abundance but also a greater consistency of deposition with time. Pacific herring underwent cyclical deposition that changed little through time. Statistical analysis reveals that none of these species is directly responding to climatic signals as indicated in the lithological and palynological record, but that this could have been partially due to the sampling resolution of the piston core. Analysis indicates a lack of correlation between the two dominant species, as might be expected on the basis of their opposing lifestyle strategies with respect to sea surface temperatures.With appropriate sampling strategies, paleohistorical fish remains from laminated sediments reflect changes in population structure and behaviour for some species, illustrate basic secondary trophic information, and provide potential clues to basin-scale oceanographic/climatological variability.     

An environmental history of two freshwater lakes in the Larsemann Hills,Antarctica

The Larsemann Hills are a series of rocky peninsulas and islands in Prydz Bay at 69° 24 S and 76° 20 E. There is about 2000 km² of ice free land with well over 150 freshwater lakes spread evenly over the granite and gneiss hills. The nearshore islands were ice free by 9500 BP, while the present coastline was exposed by 4500 BP. A relatively steady rate of ice retreat is indicated, around 0.3 ma^–1. The two freshwater lakes studied so far have evolved from oligotrophic, proglacial lagoons to fresh or brackish lakes affected by periodic influxes of salt water from sea spray and surges produced by glacial calving. The diatom assemblages increase in species diversity following marine incursion or influence. The major changes are therefore due to the postglacial recovery of sea level, rather than any intrinsic chemical evolution of the lake waters.     

Deglacial ocean and climate seasonality in laminated diatom sediments, Mac.Robertson Shelf, Antarctica

The palaeoceanography and climate history of the East Antarctic Margin (EAM) are less well understood than those of West Antarctica. Yet, the EAM plays an important role in deep ocean circulation and the global ocean system and has likely done so in the past. Deglacial-age marine sediments from the EAM provide clues about its past role during this critical period of rapid climate change. Several deep basins across the EAM such as Iceberg Alley (∼67°S, 63°E) on the Mac.Robertson Shelf (MRS) accommodate thick marine sequences that archive the deglaciation in the form of diatom-rich, continuously laminated (varved) sediments. These laminated sediments are pristinely preserved and contain seasonal and long-term information on the cryospheric and palaeoceanographic changes associated with the rapid retreat of the glacial ice sheet across the MRS. We present results of microfabric analysis of the lower ∼2 m of deglacial varves from jumbo piston core JPC43B (Iceberg Alley). Backscattered electron imagery (BSEI) of polished thin sections and scanning electron microscope secondary electron imagery (SEI) of lamina-parallel fracture surfaces are used to analyze the varves. One hundred and ninety-two laminations are investigated and their nature and temporal significance are discussed in terms of seasonal deposition and cyclicity of diatom species. Our high-resolution palaeodata record exceptionally high diatom production and silica flux associated with the retreat of the East Antarctic Ice Sheet, and seasonal sea-ice changes along the EAM. This information is invaluable for assessing cryospheric-oceanographic variation and, therefore, the local and regional response to this period of rapid climate change. Varves are made up of lamina couplets comprising (i) thickly laminated to thinly bedded orange/orange-brown very pure diatom ooze dominated by Hyalochaete Chaetoceros spp. vegetative cells and resting spores, and (ii) brown/blue-grey terrigenous angular quartz sand, silt and clay with an abundant mixed diatom flora. The colour variation between these two types of lamination is striking. Using floristic and textural information we interpret the diatom oozes as spring flux and the terrigenous laminae as summer flux. Each couplet pair represents one annual cycle and reflects seasonal changes in nutrient availability and stratification associated with the cyclical advance and retreat of seasonal sea-ice. The diatom oozes can reach up to ∼7.5 cm in thickness indicating enormous silica flux to the sea floor associated with ice sheet retreat.     

Bathymetric distribution of fossil foraminifera within marine sediment cores from the eastern part of Lützow-Holm Bay, East Antarctica, and its paleoceanographic implications

Faunal analysis of fossil foraminifera from marine gravity and piston cores collected by the Japanese Antarctic Research Expeditions (1981 and 1992) is used to estimate the impact of the latest Quaternary paleoceanography on coastal environments of the eastern part of Lützow-Holm Bay, East Antarctica.Accelerator Mass Spectrometry (AMS) carbon-14 ages produced from sedimentary organic carbon were less than 16 ka (non-corrected). Detailed correlation among submarine cores and Holocene elevated marine deposits exposed on the eastern shore of the embayment is difficult due to the indefinite reservoir correction value for marine organic matter and to upward-increasing abnormal ages for some cores.A local carbonate dissolution level can be delineated around the present depth of 300–400 m or shallower in the eastern part of Lützow-Holm Bay during the Holocene, based on distributional trends of arenaceous, calcareous benthic, and planktonic foraminifera recognized within a depth less than 600 m. Downcore recovery of calcareous foraminifera containing Bulimina aculeata from two cores obtained in a drowned glacial trough deeper than 600 m situated far beyond the dissolution depth of CaCO₃ indicates the incursion of warm, high-nutrient, and CaCO₃-saturated Circumpolar Deep Water (CDW) from the offshore area along the trough toward the southeastern coast of Lützow-Holm Bay during the Holocene. The intrusion of CDW impacted on the marine environments of the southeastern coast, thereby contributing to peripheral retreat of the ice sheet as well as increasing calcareous benthic foraminiferal productivity along the southeastern coast of Lützow-Holm Bay.     

Holocene vegetation and climate history of the northern Golan heights (Near East)

In this paper we present and discuss palynological results based on a composite profile from Birkat Ram crater lake (Northern Golan, Near East) in order to reconstruct the environmental history, including human impact, of the last 6500 years. Furthermore we apply a newly-developed botanical climatological transfer function to reconstruct climate variations in the northern Golan Heights based on this pollen data-set. The Birkat Ram record is strongly influenced by anthropogenic indicators in the pollen diagram with high quantities during the Chalcolithic period/Early Bronze Age, during the Hellenistic-Roman-Byzantine periods, during the Crusader period and finally during modern times. The palaeoclimate reconstruction method used is based on a Bayesian approach and is robust in avoiding the influence of these strong anthropogenic signals on the reconstruction results. The area has always had Mediterranean climate conditions and no distinctive climate changes can be identified during the past 6500 years. Because of the orography of the Mt. Hermon region the particular geographical position of the northern Golan Heights is obviously capable of buffering large-scale fluctuations in precipitation, which have otherwise been documented for several regions in the Near East. Electronic Supplementary Material Supplementary material is available for this article at An erratum to this article can be found at     

Holocene vegetation history with implications of human impact in the Lake Chaohu area, Anhui Province, East China

Palynological analyses of Holocene deposits located about 2 km to the southwest of the Lake Chaohu, Anhui Province, documented well the local vegetation history, its inferred environment and human impacts for the first time. An evergreen and deciduous mixed broad-leaved forest dominated by Cyclobalanopsis and Quercus existed from ca. 10,500 cal b.p. and became fully developed between ca. 8,250 and 7,550 cal b.p. Notable fluctuations occurred in the main components of the flora indicated by the decline in Cyclobalanopsis and other arboreal plants (AP), and an increase in terrestrial herbs between ca. 7,550 and 3,750 cal b.p., inferring the progressive opening of the forest under considerable human interference, which largely agrees with the archaeological evidence. After ca. 3,750 cal b.p., the broad-leaved forest largely gave way to terrestrial herbs, and never again recovered. Pinus continued to rise alongside the majority of herbs between ca. 3,750 and 2,000 cal b.p., then also declined after ca. 2,000 cal b.p. Human influence on the natural vegetation displayed in the pollen diagram seems to increase greatly up the core. The disappearance of broad-leaved forest indicates significant human impact after ca. 3,750 cal b.p., which is consistent with both the archaeological evidence and historical records. From that time the natural environment in the study area was subjected to long-standing pressure from increasing farming and population.     

Devonian Trace Fossils of the Horlick Formation,Ohio Range,Antarctica: Systematic Description and Palaeoenvironmental Interpretation

The clastic Horlick Formation contains an ichnofauna of 28 ichnotaxa dominated by burrowers. These are a mixture of simple vertical forms (Skolithos linearis, S. magnus, Bergaueria cf. langi, Rosselia socialis, Monocraterion isp.), U or arc-like forms (Diplocraterion parallelum, Arenicolites types A and B, Catenarichnus antarcticus, C. isp., aff. Lanicoidichna isp.), and complex, vertical spiral structures (Asterosoma isp., Spirophyton isp.). Horizontal burrows include Ancorichnus cf. capronus, Palaeophycus tubularis, and Psammichnites devonicus isp. nov. Surface traces comprise Haplotichnus isp., Cruziana problematica, C. rhenana, Rusophycus aff. carbonarius, R. isp., Protovirgularia rugosa, Lockeia ornata and cubichnia indet., while trackways include Diplichnites gouldi, D. isp., Maculichna? isp. and large imprints. The Horlick Formation (maximum 56 m) records an early Devonian transgression onto a deeply weathered land area that lay in the direction of Marie Byrd Land, spreading from the South Africa sector of Gondwana. Analysis of the trace fossils confirms their formation in near-shore to intertidal environments, with some ichnotaxa living close to the marine/fluvial boundary (e.g., Spirophyton, aff. Lanicoidichna, Cruziana problematica, Rusophycus aff. carbonarius, Maculichna?). The ichnofauna probably comprised a variety of marine suspension and deposit-feeding worms, shallow burrowing molluscs (bivalves and possibly bellerophontids) and several different types of arthropods, including trilobites.     

Late glacial and Holocene vegetation,climate and fire history inferred from Lagoa Nova in the southeastern Brazilian lowland

The pollen and charcoal record of a 1,500-cm-long lake sediment core from Lagoa Nova allows a reconstruction of vegetation, climate and fire history in the Atlantic lowland of southeastern Brazil. Today the potential vegetation of the study region would be dense and tall semi-deciduous forest, related to a dry season of 4 months and 1,250 mm precipitation per year. The lowermost core section of Lagoa Nova is probably of late Glacial age and contains only few, poorly preserved pollen grains, indicating dry environmental conditions. Pollen preservation was good above 1,005-cm core depth. Extrapolating from four radiocarbon dates, this interval represents about the last 10,000 years. During the early Holocene (until about 8,500 b.p.), the landscape was dominated by savanna of the campo cerrado type (open shrub woodland), with frequent trees of Curatella americana and small areas of gallery forests along the drainage system. Savanna fires, probably natural, were frequent at that time. The palaeovegetation pattern is consistent with a long dry season of about 6 months and annual precipitation lower than today. Then, up to about 7,560 b.p., gallery forests expanded in the valleys, reflecting a shorter dry season of about 5 months and somewhat higher annual rainfall. Fire was less frequent during this wetter period. Between about 7,560 and 6,060 b.p., savanna expanded and gallery forests retreated, indicating a return to drier climatic conditions of probably between 5 and 6 months dry season and lower precipitation. Fire was again frequent, but not as frequent as during the early Holocene. During the mid to late Holocene period between about 6,060 and 2,180 b.p., the valleys were covered by semi-deciduous forest, but on the hills cerrado vegetation continued to grow. The dry season was probably around 5 months and rainfall was higher than in the preceding period. During the following period between about 2,810 and 600 b.p., the open cerrado on the hills changed to closed cerrado, reflecting wetter conditions with a shorter dry season than in the previous period. It was only after about 600 b.p. that dense semi-deciduous forests expanded throughout the study region, indicating the start of modern, wet climatic conditions with an annual dry season of about 4 months. Fire was nearly absent during this period. The reconstructed palaeoenvironment from the Lagoa Nova record is similar to that of Lago do Pires, indicating that changes in past vegetation, climate and fire frequency are regionally consistent in southeastern Brazil.     

Genetic diversity and dispersal of the moss Sarconeurum glaciale on Ross Island, East Antarctica

The extent of genetic variation and dispersal mechanisms were investigated over short distances of 1–100 m, and up to 3 km, by the random amplified polymorphic DNA (RAPD) technique, for the moss Sarconeurum glaciale , at three locations on Ross Island, Antarctica. At Arrival Heights, genetic variation occurred within single colonies, and the relationships between clumps indicated that they were dispersed down small, meltwater drainage channels by water. The genetic similarities between the colonies from Arrival Heights and others from Scott Base and Crater Hill, a few km away, together with the prevailing wind direction and absence of this moss in the intervening snow-covered area, suggested longer-distance dispersal by wind. Overall, the Ross Island samples appeared to form a single, polymorphic population that was distinct from another population at Canada Glacier, 110 km distant. Somatic mutation, rather than immigration of genetically different propagules from elsewhere, appeared to be the most probable cause of genetic variability in these haploid, vegetatively reproducing Antarctic moss populations. Initiation of recolonization of S. glaciale across a dirt track at Arrival Heights was also investigated by RAPDs, to investigate how regrowth of mosses in disturbed areas occurred in the extreme environment of Antarctica.     

Late Quaternary vegetation and climatic history of eastern Nepal

The late Quaternary vegetation and climatic history at high altitude in eastern Nepal was studied through stratigraphy, radiocarbon dating and pollen analysis. The Thulo Pokhari lake (27°41′ N, 87°43′ E, ca. 3980 m a.s.l.) is just above the subalpine Abies spectabilis–Betula utilis forest zone and surrounded by Rhododendron scrub. The climate in the studied area is very humid under the influence of the monsoon. Silty sediments suggesting lake conditions prevailed at the lower part of the sediment samples, whereas the upper part consisted mainly of herbaceous peat indicating a mire condition. The pollen record extends back to ca. 11000 yr bp . Although the absence of major changes between the pollen assemblages of the Pleistocene and those in the Holocene was attributed to the situation of the sampling plot above the forest limit, the results suggest the following history. (1) The abundance of Pinus pollen around 11000 yr bp represents expansion of the area unoccupied by late successional trees and/or a drier condition. Significant occurrence of Picea pollen is noteworthy, since this genus is absent in eastern Nepal at present in spite of its wide distribution throughout the neighbouring areas of the Himalayas. (2) The increase in abundance of Quercus pollen and the decline in Pinus pollen in the latest Pleistocene represent climatic amelioration favourable to late successional mesophytes. (3) The climate was most moist and mild in the SP-III, although the absolute age of the pollen zone is not distinct. The percentages of broadleaved trees such as Alnus, Betula, Carpinus and Corylus were increased at the expense of Quercus and coniferous pollen types. Sedimentation was interrupted by lake level decline and/or glacial advance during the period from ca. 11000–1600 yr bp . (4) The increase in abundance of coniferous pollen types such as Pinus, Abies and Tsuga indicates that the climate became drier and cooler, and the present subalpine forest zone composed of A. spectabilis, T. dumosa and Betula utilis was established around 1600 yr bp . The formation of bog or alpine meadow vegetation during the period was shown by the ¹⁴C dates of the peat and the increase in Rosaceae and herbaceous pollen. (5) The increase in abundance of Ericaceae and Alnus pollen in the SP-V indicates that the present alpine Rhododendron scrub zone has been established since ca. 940 yr bp as a result of human influence on the original vegetation. This interpretation is supported by the decline in the proportion of arboreal pollen. Since the age of the initiation of human influence varies with the altitude and region, further study will be necessary to determine the accurate age.     

Crossvallia unienwillia, a new Spheniscidae (Sphenisciformes, Aves) from the Late Paleocene of Antarctica

Modern penguins are typically small to medium-sized birds, and nearly all known modern specimens are smaller than the most archaic Paleogene relatives. Here we report an incomplete humerus, associated femur and tibiatarsus of a new spheniscid, Crossvallia unienwillia nov. gen. and sp., from the Late Paleocene (~55 million years, Myr) of Antarctica, extending the record of spheniscids by about 15 Myr. Its large size (length ~140 cm) supports the idea that large body size in penguins was acquired independently at different times (Late Paleocene and Late Eocene) under dramatically different environmental conditions. Comparison of its osteological anatomy suggests that the new taxon is closely related to the extinct Anthropornithinae rather than to other penguin lineages.     

The deep population history of northern East Asia from the Late Pleistocene to the Holocene

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Late Quaternary dynamics of pinewoods in the Iberian Mountains

The role of pinewoods as native forests in the Iberian Mountains, and their long-term dynamics during the last interglacial, has been intensely debated among geobotanists. This debate has resulted in several different hypotheses regarding the interpretation of the present day landscape.Recent research on Late Quaternary palaeoecological records has enabled long-term patterns of forest dynamics in the main ranges of the Iberian Peninsula to be tested, and the parallel analysis of micro- (mainly pollen) and macroremains (woods, charcoals, fruits, seeds and leaves) has enhanced the resolution of these palaeobiological records. Additional historical information has allowed a complete overview of the past vegetal landscapes to be obtained.This work summarises the available data for Iberian mountain pines during the Lateglacial and the Holocene, focussing on the mountain regions in scenarios of absence or dominance. Based on this overview, life history traits and patterns of plant dynamics during the Holocene are discussed and compared with present landscapes.In the long term, Pinus does not always respond as a serial element in succession dynamics, not even in very favourable environments for hardwoods. Processes such as incumbency or resilience, and features such as frugality and their ability to disperse, affect the persistence of pines to the present day. This is exemplified for continental mountain areas, in which the absence of anthropogenic activities could have determined the wider coverage of pinewoods seen today.     

Review of dated Late Quaternary palaeolimnological records in the Carpathian Region,east-central Europe

The Carpathian Region (including mountains and plains) has for a long time been lacking good palaeoenvironmental and especially palaeolimnological records, particularly for the Late Quaternary. In the last two decades, many new sedimentary sequences were obtained and studied using a wide range of palaeoproxies. This article reviews results from 123 sequences in the Carpathian Region, all dated by radiometric methods. Our aim was to pay attention to the existence of these data; many of them published in national periodicals and journals. Palaeoenvironmental records with at least two proxies and with palaeolimnological interpretation were compiled in both tabular form and on maps. Inspite of the density of examined sites, an assessment of the dataset led us to the following conclusions: (1) very few provide firm hydrological–limnological interpretation, such as lake level and mire water-depth fluctuation, lake productivity changes and pH changes; (2) only 47 of them are real multi-proxy studies (have at least two proxies employed on the same sediment core); (3) glacial lakes in Slovakia and Romania as well as in Ukraine are seriously under-investigated although they would be ideal objects of palaeolimnological works with the many proxies applicable on them; (4) the Hungarian lowland areas are dominated by shallow tectonic lakes or palaeochannels, often with unsatisfactory preservation of certain biological proxies (e.g. diatoms, chironomids, cladocerans). Consequently, palaeolimnological studies from this region have to apply a different combination of proxies and approach than mountain lake studies.

High- and low-latitude forcing of Plio-Pleistocene East African climate and human evolution

The late Cenozoic climate of East Africa is punctuated by episodes of short, alternating periods of extreme wetness and aridity, superimposed on a regime of subdued moisture availability exhibiting a long-term drying trend. These periods of extreme climate variability appear to correlate with maxima in the 400-thousand-year (kyr) component of the Earth's eccentricity cycle. Prior to 2.7 Ma the wet phases appear every 400 kyrs, whereas after 2.7 Ma, the wet phases appear every 800 kyrs, with periods of precessional-forced extreme climate variability at 2.7-2.5 Ma, 1.9-1.7 Ma, and 1.1-0.9 Ma before present. The last three major lake phases occur at the times of major global climatic transitions, such as the onset of Northern Hemisphere Glaciation (2.7-2.5 Ma), intensification of the Walker Circulation (1.9-1.7 Ma), and the Mid-Pleistocene Revolution (1.0-0.7 Ma). High-latitude forcing is required to compress the Intertropical Convergence Zone so that East Africa becomes locally sensitive to precessional forcing, resulting in rapid shifts from wet to dry conditions. These periods of extreme climate variability may have provided a catalyst for evolutionary change and driven key speciation and dispersal events amongst mammals and hominins in East Africa.