Cardiolinka bohemica (Barrande, 1881) – A first representative of the Late Silurian Bohemian type Bivalvia fauna from the northern Arabian Plate,Southeast Turkey

The Rheic Ocean was a major oceanic domain between Baltica, Laurentia, Perunica and Gondwana in Ordovician-Silurian times. The cosmopolitan nepiomorphian bivalves Praecardioidei Newell, 1965 and Antipleuroidei Kříž, 2007 are characteristic of the Silurian of Perunica, peri-Gondwana, and Baltica, and occur also in Laurentia and Siberia. The Bohemian-type bivalve Cardiolinka Kříž, 1981 (Nepiomorphia Kříž, 2007, Cardiolidae Hoernes, 1884), from the Late Silurian of the Bahar-1 well core, has been found for the first time in southeastern Turkey. The strata containing the species Cardiolinka bohemica (Barrande, 1881) occur in the middle part of the Dadaş Formation in the interior Petroleum District X-Siirt of the northern parts of the Arabian Plate. The cosmopolitan species C. bohemica was until now known from the Latest Ludlow to Pridoli of the Prague Basin, France, Carnic Alps, Sardinia, East European Platform (Poland), eastern Serbia, Moesian Platform, and Arctic Canada. The new surprising subsurface data on C. bohemica in Diyarbakır-Bismil area (southeastern Turkey) therefore represent another piece of evidence in favour of strong faunistic affinity between Perunica, peri-Gondwanan Europe and the northern Gondwana margin.     

Kubergandian (Roadian,Middle Permian) of the Lycian and Aladağ Nappes (Southern Turkey)

The Ni?angahtepe Formation (Teke Dere unit) in the Lycian Nappes, and reworked pebbles and blocks from the Late Triassic Gevne Formation in the Alada? unit (both in Southern Turkey), present a well-developed Kubergandian microfauna and microflora found for the first time in this area. The following cyanobacterial and algal taxa are newly established: Clinortonella nov. gen., Ramovsia lyciana nov. sp., and Kantia? guevenci nov. sp.; some Cancellina and Armenina are revised. These Kubergandian faunas and floras present strong palaeobiogeographical affinities with the Eurasian southern margin (northern Palaeotethyan border). The Teke Dere unit represents an accretionary prism/fore-arc sequence containing different parts of the Palaeotethyan succession. The Late Triassic Gevne Formation marks the Eo-Cimmerian unconformity related to the closure of the Palaeotethys between the Taurus and Anatolian terranes.     

Late Pennsylvanian to Middle Permian revised algal and foraminiferan biostratigraphy and palaeobiogeography of the Lycian Nappes (SW Turkey): Palaeogeographic implications

Shallow-marine limestones associated to a Palaeotethyan seamount in the Teke Dere unit of the Tavas Nappe (Lycian Nappes, SW Turkey) are essentially latest Moscovian-Kasimovian in age. The wide range of microfauna and -flora of the series show biogeographic affinities comparable to those from the northern Palaeotethyan borders (especially to assemblages from the Carnic Alps, Urals, Donbass and Darvaz). These biogeographic affinities seem to persist until the end of the Early Permian (Artinskian). The Middle Permian fauna is represented by the typical warm, tropical assemblages known at the same time in the Palaeotethys (NW Caucasus, Darvaz, south China, Primorie and Japan), and in the Neotethys (Transcaucasia, central Iran, southern Afghanistan and Sibumasu). The new Kasimovian algae and incertae sedis Novantiellopsis elliottii n. gen. n. sp., Uvanellopsis fluegelii n. gen. n. sp., Tubiphytes rauzerae n. sp. and Asselodiscus davydovi n. sp. are described.     

Lower Cambrian Small Shelly Fossils of northern Sichuan and southern Shaanxi (China), and their biostratigraphic importance

Extensive information has been acquired on the occurrence of Lower Cambrian Small Shelly Fossils (SSF) in the shallow water sediments of the western fringe of the Yangtze Platform (eastern Yunnan and central Sichuan Provinces) during the last two decades. However, less is known on the temporal and spatial distribution of SSFs on the northern margin of the Yangtze Platform. Here we report SSF associations from Kuanchuanpu (Ningqiang County, southern Shaanxi Province), Shatan, and Xinli (both Nanjiang County, northern Sichuan Province) representing the Southwest Shaanxi region and from Sanlangpu, and Xiaowan (Xixiang County, southern Shaanxi Province) of the Southeast Shaanxi - Northwest Hubei region. The thickness of the Lower Cambrian SSF-bearing strata strongly decreases in this region from West to East and the stratigraphic hiatus increases. An earliest Cambrian hiatus is widely distributed on the northern margin of the Yangtze Platform. At the same time, a syngenetic brecciation occurred in wide shallow shelf settings. A tectonic mobilization, possibly related to the assembly of Gondwana following the breakup of Rodinia, may be responsible for this. In sections near Kuanchuanpu (Ningqiang County), the Meishucunian (equivalent to the Nemakit-Daldynian to Tommotian of Siberia) succession comprises ca. 55 m of dark phosphatic carbonates, siltstones, and cherts. The SSF associations are similar to those of East Yunnan and, in ascending order, can be discriminated into the Anabarites trisulcatus - Protohertzina anabarica Zone, the Watsonella yunnanensis Zone (previously mentioned as Heraultipegma yunnanensis Zone), and the Sinosachites flabelliformis - Tannuolina zhangwentangi Zone. The Siphogonuchites triangularis - Paragloborilus subglobosus Zone cannot be unambiguously detected in the Southwest Shaanxi region. The occurrence of embryos and eggs within the Anabarites trisulcatus - Protohertzina anabarica Zone, which has been reported earlier, is remarkable. The previously reconstructed life cycle and taxonomic treatment of Olivooides / Punctatus are re-discussed. The earliest SSF associations (Anabarites trisulcatus - Protohertzina anabarica Zone to Watsonella yunnanesis Zone) are missing in the Southeast Shaanxi - Northwest Hubei region and the Ninella tarimensis - Cambroclavus fangxianensis Assemblage Zone and Rhombocorniculum cancellatum Taxon Range Zone are introduced herein.     

Triassic in Iran

Summary Except of the Nakhlak and Aghdarband regions, Lower and Middle Triassic strata in Iran consist almost exclusively of carbonate rocks built on vast platforms along the shelves of the Paleo- and Neotethys. The depositional environments varied from shallow shelf sea to lagoonal and near-shore tidal flats, becoming even evaporitic towards the coastal regions of the Persian Gulf. During the early Late Triassic the formerly dominating platform carbonates were strongly reduced, being restricted for the rest of the Early Mesozoic to Southwest Iran which remained on the northern passive margin of Gondwana (later Arabian Plate). On the Iran Plate otherwise, as a result of the closure of the Paleotethys and the continent-continent collision with the southern active margin of Eurasia (Turan Plate), there was a change to thick siliciclastic and molassic sediments with only minor carbonate content durig the Upper Triassic and Lower Jurassic. Geographically restricted and completely different sequences of thick volcanoclastic and basinal Triassic sediments are known from Nakhlak in Central and from Aghdarband in Northeast Iran, both interpreted as remanents of the northern Paleotethys margin (Turan Plate).     

Signification paléoécologique et paléogéographique des bivalves du Carbonifère inférieur du bassin d’Ancenis (Massif armoricain)

Palaeoecological and palaeogeographical inferences from Early Carboniferous bivalves of the Ancenis Basin (Variscan belt, France). In the basal part of the Ancenis Formation, of Dinantian age, greenish to purple mudstones display a few bivalves, assigned to the genera Lithophaga, Modiolus, and Naiadites. Because Naiadites is considered to be a non-marine bivalve, and is only previously reported in Scotland in Dinantian rocks, a brackish environment can be assumed for the Ancenis Basin during the Early Carboniferous. To cite this article: M. Ballèvre, H. Lardeux, C. R. Palevol 4 (2005).     

New records of early Middle Jurassic belemnites in the French Subalpine Basin and their paleobiogeographic significance

A biostratigraphic and systematic study based on belemnites collected along with ammonites was performed on four sections in the Subalpine Basin (SE France): Lac du Castillon and La Baume (Castellane area), Galabrun and Grand Lara (Gap area). The specimens, originating from hemi-pelagic marl-limestone alternations in the lower part of the “Calcaires à Zoophycos” Formation, are dated from Middle Aalenian (Murchisonae Zone) to Lower Bajocian (Humphriesianum Zone). Five belemnite taxa (Megateuthis elliptica, Holcobelus munieri, H. trauthi, Pachybelemnopsis roettingensis, Hibolithes sp.) have been identified, and two more taxa are reported in an open nomenclature (Belemnitida incertae sedis sp. 1 and sp. 2). The biostratigraphic range of the belemnite fauna is established. The new findings contribute to a more detailed understanding of the paleobiogeography of holcobelid belemnites that flourished at the northern margin of the Tethys Ocean and formed a distinct sub-Mediterranean fauna. The association herein described is similar to the fauna of the Calabro-Peloritani Arc (Calabria, Italy), a further hint for the supposed paleogeographic position of the latter during the Middle Jurassic.     

Early Carboniferous spiriferoid brachiopods from the Qaidam Basin,Northwest China: Taxonomy,biostratigraphy and biogeography

Six Early Carboniferous brachiopod species in four genera of the Superfamily Spiriferoidea are described from the Qaidam Basin, northwestern China, including a new genus, Qaidamospirifer, and two new species: Grandispirifer qaidamensis and Qaidamospirifer elongatus. Additionally, a new genus, Triangulospirifer, is also proposed to replace Triangularia Poletaev, 2001 that was preoccupied by a Devonian molluscan genus.On the basis of the new material as well as published information, we have reviewed the taxonomic composition and the stratigraphic and palaeobiogeographic distributions of the three previously established genera from the viewpoint of palaeobiogeography. The study reveals that Grandispirifer has a relatively long stratigraphic range from the late Tournaisian to Serpukhovian. During this interval, the genus attained a wide geographical distribution, reaching Northwest China, western Yunnan of West China, Japan, as well as Iran and North Africa. Angiospirifer first occurred in western Europe in the Viséan, and later migrated to North Africa during the late Viséan. In the Serpukhovian, it migrated eastward, reaching the Donets Basin of Ukraine and the Qaidam Basin in Northwest China. Anthracothyrina evolved from Brachythyrina in North Africa in late Viséan, then dispersed north-westward to western and eastern Europe and, further eastward to the Qaidam Basin during the Serpukhovian.     

新疆北部侏罗、白垩纪非海相腹足类

记述新疆准噶尔盆地和吐鲁番盆地侏罗、白垩纪非海相腹足类１０属１５种，其中６新种、１未定种。根据这些腹足类的地层分布特点将它们划分为３个组合，自下而上为１）Ｖｉｖｉｐａｒｕｓｓａｎｇｏｎｇｈｅｅｎｓｉｓ组合，时代为早侏罗世；２）Ｖｉｖｉｐａｒｕｓｑｉｋｅｔａｉｅｎｓｉｓ－Ａｍｐｌｏｖａｌｖａｔａｓｕｔｕｒａｌｉｓ组合，时代为中侏罗世；３）Ｃｉｎｃｉｎｎａｔｕｒｇｅｎｓｉｓ组合，时代为早白垩世。对这些腹足类生活习性的分析，这两个盆地在早侏罗世气候温暖湿润，草木繁茂，大部分地区为平静、滞流的湖泊、沼泽；中侏罗世盆地相对下陷，水域扩大加深；中侏罗世末期至晚侏罗世早期，盆地抬升，水域范围明显缩小；至早白垩世，盆地又扩大加深。     

An integrated reconstruction of the early Pleistocene palaeoenvironment of Homo erectus in the Denizli Basin (SW Turkey)

The early Pleistocene travertines from Kocabaş in the Denizli Basin (SW Turkey), from which the only known Homo erectus from Anatolia derives, are covered by a succession of lake deposits. So far, the taphonomic history of the site has precluded a detailed palaeoecological analysis. This paper details the sedimentary succession and palaeoenvironmental conditions by analysing the micro- and macropalaeontological species compositions. These data provide direct evidence of the opportunities and limiting factors of the environment inhabited by hominins during the early Pleistocene. Four distinctive lithostratigraphic units are recognized in the Quaternary succession, consisting of: Lower Travertine, Lower Conglomerates, Upper Travertine, and Upper Conglomerates. These units correspond to an alternation of lacustrine limestone, fluvial-lacustrine siliciclastic deposits and subaerially precipitated travertine accumulations. The age of the succession is constrained by cosmogenic nuclide concentration, palaeomagnetic measurements and large mammal biostratigraphy which suggest deposition occurred between ∼1.6 and 1.2 Ma. The travertine succession contains a moderately diverse macromammal fauna, including Homo erectus, decapod crustaceans and leaf imprints. The herbivore association likely dwelled in a mixed landscape during temperate and humid climatic conditions, supported by negative δ¹⁸O values from analysed gastropod and bivalve shells. The presence of an anomalohaline lake is indicated by an abundant and well-preserved ostracod community, consisting of alkaline tolerant taxa and freshwater genera further supported by the presence of cardiid bivalves.     

New evidence for the Early Devonian age of the Jauf Formation in northern Saudi Arabia

Well-preserved Early Devonian continental and sparse marine palynomorph assemblages recovered from an exploration borehole located in northern Saudi Arabia are reported here and add to the understanding of Arabian Devonian stratigraphy. The units examined are the conventionally cored lower Murayr, upper Hammamiyat and Sha’iba Members of the Jauf Formation. A comparison with data sets from elsewhere on the Arabian Plate suggests that the units examined are Emsian in age. The age of the Murayr and Hammamiyat boundary interval is revised, based on the occurrence of Dibolisporites eifeliensis in northern Saudi Arabia, and placed in the Rhabdosporites minutus (Min) Interval Zone of the Emphanisporites foveolatus-Verrucosisporites dubia (FD) Oppel Zone. A new miospore, Apiculiretusispora densa with distinctive interradial thickenings, is described.     

New data on mollusks from the Miocene (Tarkhanian-Chokrakian) of northern Sinop Province,Turkey

An insufficiently known bivalve and gastropod assemblage from the Early-Middle Miocene (Tarkhanian-Chokrakian) of northern Sinop Province (Turkey), is analyzed. Environments of the assemblage are reconstructed for the Chokrakian as subtidal, with prevailing lime and sandy bottom and good aeration, and partially well vegetated. Impoverishment of the mollusk biocoenose in this part of the marine basin (only 18 bivalve and 22 gastropod species recorded) compared to other areas, including the closest regions, Bulgaria on the west and Georgia on the east, is emphasized. The relatively low diversity of the fauna is probably connected not only with insufficient collecting, but with special hydrological conditions. A special aspect of the fauna is highlighted by the presence of the bivalve Circomphalus foliaceolamellosus subplicatus (Orb.), which is rare in the Chokrakian.     

Aratrisporites saharaensis sp. nov.: A characteristic lower carboniferous miospore species of North Africa

A new miospore species Aratrisporites saharaensis is described. Attributed to a genus previously known mainly from the Trias, it constitutes the first record of monolete camerate miospores from the Carboniferous.This taxon occurs abundantly in the Lower Carboniferousof northern region of the Sahara, North Africa. It has, in the past, been incorrectly attributed by several authors to Spelaeotriletes balteatus (Playford) Higgs 1975. Its stratigraphic range is late Tournaisian to early Namurian.     

Discovery of the Misellina Zone (latest Kungurian) in the Lentas unit of the Pindos Series (Crete)

The origin and significance of the Lentas unit in southern Crete (Greece) is still subject to controversies. New stratigraphical and palaeontological results allow correlations with the classical Pindos-type series in Greece. In particular, new datings from reworked fossiliferous limestone pebbles within the Lentas unit are presented. These data concern especially the carbonate microfacies and carbonate microfossils from the latest Early Permian (i.e., from the late Kungurian Substage or Misellina Zone). For the first time in Greece, this period is recognized as being rich in foraminifers and calcareous algae. These fossils are described; especially, two dasycladales Likanella (?) cretae nov. sp. and Uragiellopsis bonneauii nov. sp. are established. The presence of these reworked Palaeotethyan sediments at the base of the Lentas unit suggests a derivation from the Palaeotethyan active margin (arc/fore-arc series).     

Discovery of the oldest known Pterygota in the Lower Carboniferous of the Upper Silesian Basin in the Czech Republic (Insecta: Archaeorthoptera)

The earliest pterygote (winged insect), dated from the Lower Carboniferous (Namurian A/E1, circa 324 millions years ago) is described from the Upper Silesian Basin in the Czech Republic. On the basis of its wing venation, it is attributed to the Archaeorthoptera Béthoux and Nel, 2002, crown group of the “Orthoptera”. Besides Apterygota (Collembola and Archaeognatha) known from the Lower Devonian, extremely rare pterygote insects are known from Lower Carboniferous deposits when they first appeared. The present discovery supports the hypothesis of the presence of the ancestor lineage of the orthopteroid in the Lower Carboniferous ecosystems.     

Palaeoginkgoxylon zhoui, a new ginkgophyte wood from the Guadalupian (Permian) of China and its evolutionary implications

A new taxon of ginkgophyte affinity Palaeoginkgoxylon zhoui gen. nov. et sp. nov. is described from the Guadalupian Lower Shihhotse Formation of the Hulstai coalfield, Inner Mongolia Autonomous Region (Nei Mongol), northern China, on the basis of the anatomical structures of the broad eustele and pycnoxylic secondary xylem. The anatomical structure of the new woody tree trunk resembles both the early gymnosperms of Eristophyton-Pitus types and the modern Ginkgo. Therefore, the new tree trunk is interpreted as representing a transitional stage in the evolution of Ginkgo from early arborescent lignophytes since the Early Carboniferous.     

Microfossil assemblage from the Late of Middle Carboniferous in Western Shaan-Gan-Ning Basin,Northwest China

In this paper, some results of the study on the sporepollen of the Yanghugou Formation in the western Shaan-Gan-Ning (Ordos) Basin of Northwest China axe reported. Plenty of well preserved microfossil, which 136 types belong to 62 microspore genera and one megaspore genus, including 2 new genera , 13 new species, I Acritarch (Chamosphaera pseudozonatus gen. et sp. nov.), from 8 bore-holes and outcrop have been discribed. In generally, the microfiora of the Yanghugou Formation is similar to the Taiyuan Formation (Upper Carboniferous). According to the spore-pollen assemblage of Yanghugou Formation may correlate with those in the Middle Carboniferous of Great Britain, Belgium the north of France, German and North America. Besides, the assemblage are similar to Middle Carboniferous of Nortth Shandong and Hengshanbu of Ningxia in the microfossil assemblage. The geological age of the beds is considered to Late of Middle Carboniferous.     

Detrital zircon and micropalaeontological ages as new constraints for the lowermost tectonic unit (Talea Ori unit) of Crete,Greece

The Talea Ori unit is the lowermost known tectonic unit of Crete and the most external part of the Hellenides. Its stratigraphy ranges from Late Carboniferous to Oligocene and outcrops of the lower part are only known in the Talea Ori mountains (central Crete).In this area, a black sandstone at the base of the Galinos Beds, thought to be the oldest formation, contains zircons which were dated using the single grain evaporation method. The majority of these grains yielded Late Carboniferous ages (Variscan), while a small group yielded Early Proterozoic ages. The age distribution of these zircons suggests that, at the Carboniferous–Permian boundary, not much of the older North Gondwanan basement was exposed and that a river system carried detrital material from the Variscan belt towards the forming Neotethyan rift.Additionally, higher up in the stratigraphy benthic foraminifers (miliolids) were found in clasts from a conglomerate which was so far thought to be of Early Triassic age [Epting, M., Kudrass, H.-R., Leppig, U., Schäffer, A., 1972. Geologie der Talea Ori/Kreta. N. Jb. Geol. Paläont. Abh. 141, 259–285.]. These miliolids belong to the species Hoyenella inconstans [Michalik, J., Jendrejakova, O., Borza, K., 1979. Some new foraminifera species of the Fatra-Formation (Uppermost Triassic) in the West Carpathians. Geol. Carpath. 30 (1), 61–91.], thus attributing a Late Triassic (Carnian–Norian?) maximal age to this conglomerate. The carbonate platform from which the miliolids-bearing clasts come is not known.The presence to the north of a continuous hemipelagic record from the Carboniferous to the Triassic (Phyllite–Quartzite and Tripali units), attributed to the Palaeotethys realm, allows the Talea Ori unit and its lateral equivalents (the Ionian zone) to be assigned to the westward continuation of the Cimmerian block and therefore to the northern margin of the East Mediterranean Neotethys ocean.     

Le passage Lias–Dogger de la Dorsale de Tunisie septentrionale: Nouveaux apports biostratigraphiques. Âge Toarcien supérieur de la distension téthysienne

In the Jebel Bou Kornine of Hammam Lif (northern Tunisia), the transition between Lias and Dogger is located in a carbonate formation that is approximately 150 m thick (Kef El Orma Formation). Based on ammonite faunas, its age can be determined as Upper Toarcian (Aalensis Zone) up to Lower Bajocian (Propinquans Zone). Four successive conglomeratic units are recognized in this formation. The two earlier ones are situated in the Uppermost Toarcian (Aalensis Zone, Lugdunensis Subzone), while the later two are in the Opalinum Zone of the Lower Aalenian (Opalinum and Comptum Subzones). These deposits, which associate carbonate-bearing conglomerates with laminated/slumped calcarenites, result from a gravitational flow on a palaeoslope induced by extensive palaeofaults. They testify to a tectonic instability during the Upper Toarcian, causing the formation of two distinct palaeostructural units: the ‘Tunisian trough’ and the ‘Tunisian ridge’ carbonated platform, both belonging to the Maghrebian passive margin of the Tethys. We must insist on the part played, as soon as the Upper Toarcian, by the polyphased tilted-blocks tectonics on the deposition of the different conglomeratic units of the Bou Kornine Jebel.