Refinements in the Upper Permian to Lower Jurassic stratigraphy of Karakorum, Pakistan

New sampling on critical intervals of the uppermost Permian and Triassic successions of the Northern Karakorum Terrain in the Karakorum Range (Pakistan) has refined the stratigraphy. Two types of successions may be distinguished in the Karakorum Range: a carbonate platform succession, spanning the whole interval from Upper Permian to Upper Triassic, possibly with several gaps; and a basinal succession, deposited from the Middle Permian to Early Carnian (Late Triassic), when the carbonate platform prograded into the basin. With the approaching and later docking of the Karakorum Block against the Asian margin closing the Paleo-Tethys, a portion of Karakorum emerged while another part subsided as a fore-deep, receiving clastics from the emerging Cimmerian Range. Molassic sediments filled the basin, whilst shallow-water carbonates transgressed over the emerged carbonate platform sometime between the latest Triassic and the Pliensbachian (Early Jurassic), with Cimmerian deformation occurring to the north. The age control is provided by conodonts, with assemblages of late Wuchiapingian, Changhsingian, Induan (Griesbachian and Dienerian), late Olenekian, early Anisian, late Ladinian, and early Carnian ages, respectively. Some information on the section around the P/T boundary is provided by palynology and isotopic C¹³ values. The dating of the Norian/Rhaetian platform is provided by foraminifers.     

An early isocrinid sea lily from the middle to late Anisian boundary (Middle Triassic) of south‐west China – evidence for a far‐east Tethyan origin of the family Isocrinidae

Abstract: The isocrinid sea lily Tyrolecrinus wugangi sp. nov. from uppermost middle to lowermost upper Anisian (lower Middle Triassic) strata of Leidapo near Qingyan, Guizhou Province, south‐west China, provides new data on the early phylogeny of the order Isocrinida. The new species is the earliest unequivocal representative of the family Isocrinidae. It predates all other known species of the genus Tyrolecrinus that come from upper Ladinian to Rhaetian strata and also all other known taxa of the Isocrinidae. Nevertheless, its stem is of fully developed isocrinid type and is characteristic of the genus, with consistently synostosial, rarely cryptosymplectial, distal nodal articular facets. These findings indicate that the Isocrinidae evolved prior to the late middle Anisian in the far‐eastern part of the Palaeotethys ocean, perhaps in the region forming today’s south‐west China, and spread to various other regions later in the Middle and Late Triassic. The genus Tyrolecrinus is revised and the new genus Bakonycrinus gen. nov. is erected.     

Middle Triassic basin evolution and stratigraphy in the Carnic Alps (Austria)

Summary A local intraplatform basin developed in the Gartnerkofel-Zielkofel area of the Carnic Alps (southern Carinthia, Austria) during the Middle Triassic (Ladinian). This basin was filled with a transgressive basinal sequence composed of the Uggowitz Formation and overlying Buchenstein Formation. At the northwestern slope of the Gartnerkofel, the platform carbonates of the Schlern Dolomite interfinger with the Buchenstein Formation, causing the formation of two depositional sequences. The Uggowitz Formation consists of the Uggowitz Breccia and the Kühweg Member. Sediments of the Uggowitz Breccia were formed by different types of gravity induced processes. The Kühweg Member is a thin sequence of silt-and fine-grained sandstones which were deposited in a slope to basin margin environment by turbidity currents. The overlying Buchenstein Formation consists of hemipelagic to pelagic limestones of Fassanian age with intercalated pyroclastic rocks (Pietra verde). Nodular limestones were deposited under slow rates of accumulation during a relative sea-level highstand. The uppermost Buchenstein Formation is composed of hemipelagic limestone beds with intercalated graded calcarenites and breccias of platform-derived debris, showing characteristics features of a fore-reef slope of the prograding Schlern Dolomite. Uggowitz Formation and basal Buchenstein Formation are interpreted as a transgressive systems tract, nodular limestones from the middle part of the Buchenstein Formation mark an early highstand systems tract, forereef slope sediments of the upper Buchenstein Formation formed during the beginning regression of a late highstand systems tract, the basal part of the overlying Schlern Dolomite probably reflects a lowstand systems tract. The intercalated bedded limestone facies within the Schlern Dolomite is characterized by large, platform derived blocks, slump structures, breccia beds, graded calcarenites and hemipelagic limestones indicating a forereef slope environent. This intercalated facies belongs to the Buchenstein Formation and interfingers with the Schlern Dolomite. Conodonts from this intercalated slope facies point to Late Fassanian age. Therefore, the two Middle Triassic depositional sequences of the Gartnerkofel area can be correlated with the depositional sequences ‘Ladinian 1’ and ‘Ladinian 2’ of the Dolomites, proposed byDe Zanche et al. (1993). A brief comparison with the basinal sequences of similar age of the karawanken Mountains and the Carnia is presented.     

Late Triassic, Early and Middle Jurassic Radiolaria from ferromanganese-chert ‘nodules’ (Angelokastron, Argolis, Greece): evidence for prolonged radiolarite sedimentation in the Maliac-Vardar Ocean

In the Argolis, the Basal Sequence, constituting the eastern Pelagonian margin which bordered the Maliac-Vardar oceanic domain, includes shallow-water carbonates of Late Triassic-Early Jurassic, condensed pelagic limestones of Early-Middle Jurassic, radiolarian cherts of late Middle-Late Jurassic age and siliceous mudstones and sandstones rich in ophiolite fragments. Up-section, coarse breccias, including clasts of boninites derived from the ophiolite obducted onto the Pelagonian margin in Late Jurassic times crop out. Near Angelokastron a small quarry exposes pervasively sheared dark reddish-brown, radiolarian-bearing cherty shales with disrupted fragments of chert and chert nodules impregnated by ferro-manganese oxides. These shales occur in the footwall of a thrust bringing them into contact with the Pantokrator Limestone of the Basal Sequence. We collected more than 30 samples of the chert fragments and the shaly matrix. Thirteen nodules and one matrix sample yielded determinable radiolarians. Low to non-detectable concentrations of trace metals such as Co, Cr, Cu, Ni, Zn, and Pb indicate a hydrothermal origin of the ferro-manganese mineralization. The radiolarian taxa found indicate four age groups for the nodules that are embedded in the siliceous shale matrix that yielded a Middle Jurassic age (middle Bathonian). The first group includes a nodule of Late Triassic age (late Norian to Rhaetian); the second group nodules of Early Jurassic age (late early to late Pliensbachian and probably middle-late Toarcian); the third group nodules of early Middle Jurassic age (Aalenian–Bajocian); the last group finally includes nodules of late Middle Jurassic age (Bajocian–Bathonian). The presence of Upper Triassic to Middle Jurassic Mn-impregnated chert nodules in a Middle Jurassic matrix indicates a deep oceanic environment of deposition outside the Pelagonian realm (easternmost Adria Plate), which at that time was a shallow-water carbonate platform with a thin pelagic limestone cover. The chert nodules are with all certainty derived from the oceanic Maliac-Vardar domain and were, together with their host formation, tectonically emplaced onto the Pelagonian margin. We speculate that these nodules, more lithified than their matrix, were exhumed on the slope of an intra-oceanic accretionary wedge and were redeposited in the Middle Jurassic siliceous mudstones on the floor of the subducting Maliac-Vardar Ocean.     

RHAETIAN (LATE TRIASSIC) MONOTIS (BIVALVIA: PECTINOIDA) FROM THE EASTERN NORTHERN CALCAREOUS ALPS (AUSTRIA) AND THE END-NORIAN CRISIS IN PELAGIC FAUNAS

Abstract:  Species of marine bivalves of the pectinoid genus Monotis provide useful biochronologic indices for the Late Triassic (middle Norian–earliest Rhaetian). We report the succession of Monotis at Hernstein in Lower Austria where typical late Norian Monotis salinaria (Schlotheim) are overlain by strata with apparently the youngest Monotis known of demonstrable Rhaetian age: Monotis hoernesi Kittl and Monotis rhaetica sp. nov., a species closely related to M. rudis Gemmellaro. A Rhaetian (Sevatian 2) age is confirmed by the co-occurrence of Monotis with the platform conodont Misikella posthernsteini and close association with the ammonoid Paracochloceras . A review of late Norian monotid species indicates that a profound extinction event occurred in the pelagic realm at or close to the Norian/Rhaetian boundary where c. 15 monotids ( Monotis s.l . and Maorimonotis ) became extinct. The surviving Monotis are dwarfed when compared to their Norian predecessors and may represent an ecological/phylogenetic response to the crisis.     

Albian calcareous nannofossils from ODP Site 1258, Demerara Rise

Site 1258, drilled during Ocean Drilling Program (ODP) Leg 207 on Demerara Rise off the northern coast of South America, recovered organic-rich Albian sediments that yield abundant, moderately to well preserved calcareous nannofossils. Biostratigraphic analysis shows the section primarily spans Roth’s (Initial Reports DSDP 44. US Government Printing Office, Washington (1978) 731) middle to late Albian Zone NC9. A disconformity separates these sediments from overlying uppermost Albian laminated shales from Zone NC10. The presence of Seribiscutum primitivum within the Albian section represents the first known occurrences of this species at such low latitudes, as Demerara Rise was located within 15° of the equator during the mid-Cretaceous. This species exhibits a bipolar distribution and is considered a cool-water, high-latitude species. Its presence on Demerara Rise indicates cooler water incursions either through changes in surface circulation or upwelling conditions during the opening of the Equatorial Atlantic.     

Evidence for Prosauropod Dinosaur Gastroliths in the Bull Run Formation (Upper Triassic,Norian) of Virginia

Definitive criteria for distinguishing gastroliths from sedimentary clasts are lacking for many depositional settings, and many reported occurrences of gastroliths either cannot be verified or have been refuted. We discuss four occurrences of gastrolith-like stones (category 6 exoliths) not found within skeletal remains from the Upper Triassic Bull Run Formation of northern Virginia, USA. Despite their lack of obvious skeletal association, the most parsimonious explanation for several characteristics of these stones is their prolonged residence in the gastric mills of large animals. These characteristics include 1) typical gastrolith microscopic surface texture, 2) evidence of pervasive surface wear on many of these stones that has secondarily removed variable amounts of thick weathering rinds typically found on these stones, and 3) a width/length-ratio modal peak for these stones that is more strongly developed than in any population of fluvial or fanglomerate stones of any age found in this region. When taken together, these properties of the stones can be explained most parsimoniously by animal ingestion and gastric-mill abrasion. The size of these stones indicates the animals that swallowed them were large, and the best candidate is a prosauropod dinosaur, possibly an ancestor of the Early Jurassic gastrolith-producing prosauropod Massospondylus or Ammosaurus.

Skeletal evidence for Upper Triassic prosauropods is lacking in the Newark Supergroup basins; footprints (Agrestipus hottoni and Eubrontes isp.) from the Bull Run Formation in the Culpeper basin previously ascribed to prosauropods are now known to be underprints (Brachychirotherium parvum) of an aetosaur and underprints (Kayentapus minor) of a ceratosaur. The absence of prosauropod skeletal remains or footprints in all but the uppermost (upper Rhaetian) Triassic rocks of the Newark Supergroup is puzzling because prosauropod remains are abundant elsewhere in the world in Upper Triassic (Carnian, Norian, and lower Rhaetian) continental strata. The apparent scarcity of prosauropods in Upper Triassic strata of the Newark Supergroup is interpreted as an artifact of ecological partitioning, created by the habitat range and dietary preferences of phytosaurs and by the preservational biases at that time within the lithofacies of the Newark Supergroup basins.     

Middle Triassic Palynological assemblage in the Wangjiashan Basin of Jingyuan,Gansu and Its Stratigraphical Significance

The sediments, overlain by the Middle Jurassic strata in the Wangjia-shan Basin, are a sequence of grey-green sandstones and mudstones and sum more than700 m. It was originally refered to Late Triassic in age. The palynomorph-bearingsamples were collected from the upper part of the sequence in a bore hole. Three sam-ples yielded a, bundant plant microfossils. The sporepollen assemblage consists of 63 species (types) and 51 genera, including 3 new species, namely, Cyclogranisporites callosus, Verrucosisporites wangjiashanensis and Longdongspora jingyuanensis. The component and constituent percentages of palynological assemblage are listedin Tab. 1, in which Pteridophytic spores hold predominant place (67.1%), Gymnospermous pollen reach 32.9%. Among the Pteridophytic spares Punctatisporites (8 forms)plays an important role in assemblage up to 32.3%, followed by Verrucosisporites, Calamospora, Duplexisporites and Apiculatisporis. Besides, Lundbladispora, Kraeuselis-porites and Aratrisporites are represented only by individual specimens. In Gymnospe-rmous pollen Disaccites hold 11.3%, Disaccite striatiti 13.1%, by amplification of Tae-niaesporites reaching 7.2%,others are Piceites, Protoconiferus and Parataeniaesporites.Angustisulcites and Chordasporites are a few in the assemblage. The present assemblage is quite different from the early and upper Triassic paly-nological assemblages of Shan-Gan-Ning Basin, (Qu Li-fan, 1980, 1982) but rather similar to that of the middle Triassic Tongchuan Formation. (Qu Li-fan, 1980; Liu Zhao-sheng et al, 1981) There are 33 genera and 18 species in common between the present assemblage and that of the Tongchuan Formation. Therefore, the present assemblage maybe assigned to the younger age of Middle Triassic. The bearingbed of the spore-pollen assemblage is called the Nanyinger Group. Thisgroup was compared originally with the Yanchang Group of Shan-Gan-Ning Basin, referring to Late Triassic. Based on present data, the Nanyinger Group may include se-diments of younger age of Middle Triassic.     

广西凤山、西林等地中三叠统上部双壳类分带研究*

讨论广西西部中三叠统上部的双壳类生物地层序列,建立5个化石带;并附述黔南和滇东南晚三叠世1个双壳类带.这些带自下而上为:(1) Daonella (Longidaonella) producta-D. americana 带,属晚安尼早中期;(2) D. moussoni 高峰带,属晚安尼晚期;(3) D. rieberi-D. indica 带,属早拉丁期;(4) D. kotoi-D. jilongensis 带,属晚拉丁早期;(5) Halobia subcomata-D. varifurcata带,属晚拉丁晚期;(6) H. rugosoides-H. bifurcata 带,属早卡尼期.文内简要描述了各带的主要标志化石23种.     

Late Ordovician palaeoceanographic changes as reflected in the Hirnantian–early Llandovery succession of Jämtland, Sweden

A study of the Upper Ordovician–Lower Silurian strata in Jämtland, central Sweden, shows that large-scale changes in shelf deposition took place close to the systems boundary. These changes include unconformity development and the replacement of a siliciclastic shelf with a carbonate-dominated shelf, suggesting the interaction of allocyclic controls such as changing eustatic sea-level and climate. The 6-m-thick Ede Formation is a key lithosome for interpretation of this transition. Its sediments were deposited in the Caledonian foreland basin, situated east of the closing Iapetus Ocean on the western margin of the Baltic craton. A major part of the late Caradoc to late Ashgill (into the Hirnantian) was characterised by continuous and uniform deposition over wide areas (Kogsta Formation), whereas erosional surfaces and complex lateral facies relationships characterise the Ordovician–Silurian boundary strata (Ede Formation and lateral equivalents). The Ede Formation represents the end of terrigenous deposition, which in the middle Aeronian was followed by regional expansion of carbonate deposition (Berge Formation). A syn-sedimentary erosional surface, with at least 1 m of relief locally, forms the lower boundary of the Ede Formation. This surface is overlain by two types of conglomerate. Lower parts of the Ede Formation consist of medium to thick-bedded quartzites. A second erosional surface with only minor (few centimetres) relief occurs on top of these quartzites. The upper parts of the Ede Formation consist of a thin, basal favositid biostrome overlain by thin bedded, calcareous sandstones, limestones and intensely bioturbated shales. Analysis of stratigraphic boundaries and the facies succession suggests that the lower Ede Formation represents a major downward shift in coastal onlap and by-pass sedimentation that created the lower erosional surface. The erosional surface in the middle of the Ede Formation is inferred to have formed during the subsequent maximum lowstand or as a ravinement surface, and is interpreted as an unconformity. The succession is subdivided into four facies associations, each corresponding to a specific systems tract: (a) a Shale–Siltstone Association (uppermost Kogsta Formation), deposited during a highstand situation in mid-outer shelf areas; (b) a Quartzite Association (the lower Ede Formation), deposited during forced regression in a shoreface environment; (c) a Mixed Carbonate–Siliciclastic Association (the upper Ede Formation), deposited during transgression in a wave-dominated, proximal shelf environment when clastic supply was reduced; and (d) a Micritic Limestone Association (lowermost Berge Formation), deposited during a second highstand situation in a low-energy, offshore environment.

Conodont data, together with a previously reported Hirnantia fauna, constrain the position of the Ordovician–Silurian boundary to the lower 1.65 m of the Ede Formation, or less likely, to the uppermost metre of the underlying Kogsta Formation, i.e., within a 2.65-m-thick uncertainty interval. The base of the Berge Formation is about 4 m above the top of the uncertainty interval, and is dated as being mid-Aeronian in age, suggesting condensation and/or a hiatus close to, or at, the Ordovician–Silurian boundary. These data tie the unconformity and the regional facies change from a siliciclastic to a carbonate-dominated shelf to Late Ordovician–Early Silurian eustatic and climatic changes.     

Characteristics of peritidal facies and evidences for subaerial exposures in Dachstein-type cyclic platform carbonates in the Transdanubian Range,Hungary

In the Late Triassic, an extremely large carbonate platform system (Dachstein-type platforms) developed on the margin of the Neotethys. On the wide inner platform cyclic peritidal, lagoonal successions were deposited. In the Transdanubian Range (Hungary), the lower part of the 1.5–2-km-thick cyclic succession (Upper Tuvalian–mid-Norian) is pervasively dolomitised, the upper part (Upper Norian–Rhaetian) is non-dolomitised; there is a transitional interval between them made up of partially dolomitised cycles. The peritidal–lagoonal cycles are commonly bounded by well-developed disconformity surfaces reflecting subaerial erosion that punctuated the marine carbonate accumulation. Truncation of the cycles was preceded by pervasive cementation of the previously deposited cycle. In the early stage of the platform evolution, tidal flat dolomitisation under semi-arid conditions led to the consolidation of the previously deposited sediments. The truncation surfaces were commonly covered by dolocretes. During the more humid Late Norian–Rhaetian period, the early cementation was followed by karstification, accumulation of wind-blown dust and pedogenesis. Erosion during regularly recurring subaerial exposure that commonly reached the previously deposited subtidal beds suggests eustatic control of the cyclicity and supports the application of an allocyclic model, even if the Milankovitch signal is imperfect.     

Genesis of stromatactis in an upper jurassic carbonate buildup (Mlynka,Cracow region,Southern Poland): Internal reworking and erosion of organic growth cavities

Summary The Middle Oxfordian strata in the southern part of the Cracow-Wielun Upland consist of platy and bedded limestones (‘normal facies’), of massive limestones as well as locally of mass flow sediments. Massive limestones, prevailing in the Upper Oxfordian, form commonly carbonate buildups, which are made up predominantly of cyanobacterial allochems and to a minor amount of siliceous sponges. Stromatactis can be best observed in the Mlynka quarry. They occurs in the uppermost part of slope sediments close to a cyanobacterial-sponge buildup. The bedding-plane of the slope sediments is directly overlain by debris-flow and grain-flow sediments. Fragments of a primary laminar framework rich in growth-cavities occur in the uppermost part of the slope sediments as precondition for the formation of stromatactis. The stromatactis cavities were formed by internal reworking and erosion within these organic growth cavities, caused by strong bottom currents due to mass transport from higher parts of the buildup.     

Late Triassic sedimentary evolution of Slovenian Basin (eastern Southern Alps): description and correlation of the Slatnik Formation

Successions of the Slovenian Basin structurally belong to the easternmost Southern Alps. During the Late Triassic, they were part of the Adriatic continental margin. Norian–Rhaetian successions of the Slovenian Basin are characterized mainly by dolomite of the Bača Dolomite Formation. However, in the northern part of the basin, Late Triassic limestone is preserved above Bača Dolomite Formation and is formalized as the Slatnik Formation. It is composed of hemipelagic limestone alternating with resedimented limestones. The succession documents an upward progradation of the slope environment composed of three high-frequency cycles. Most prominent progradation is referred to the second, i.e., Early Rhaetian cycle. The Slatnik Formation ends with thin-bedded hemipelagic limestone that records the end-Triassic productivity crisis, or rapid sea-level fall. The overlying resedimented limestones of the Early Jurassic Krikov Formation, document the recovery of production and shedding from the adjacent carbonate platform.     

FIRST OCCURRENCE OF FOOTPRINTS OF LARGE THERAPSIDS FROM THE UPPER PERMIAN OF EUROPEAN RUSSIA

Abstract:  Large footprints of terrestrial tetrapods have been found in the Cis-Urals region of European Russia. The footprint horizon is in Late Permian (Changhsingian) deposits of the Vyatkian Gorizont (uppermost Tatarian) approximately 50 m below the local Permian/Triassic boundary. Seventeen randomly orientated footprints were excavated and are referred to the ichnospecies Brontopus giganteus . The footprints were emplaced in a reddish-brown mudstone that was deposited from suspension beneath shallow ponded water in a floodplain environment. They were subsequently cast by the base of the overlying fine-grained sandstone, which was deposited from a sheet-flood event. The footprints were produced by a large therapsid, possibly a dinocephalian, but more probably a dicynodont, and represent the first ichnological record of the Therapsida from the Upper Permian of Russia.     

Facies stacking and extinctions across the Triassic–Jurassic boundary in a peritidal succession from western Sicily

An uppermost Triassic–lowermost Jurassic carbonate platform succession, which is 430 m thick, in northwestern Sicily is described with the aim to provide new data on the sedimentological and biological variations across the Triassic–Jurassic boundary in peritidal environments. The studied succession belonged to the rimmed carbonate shelf that developed during the Late Triassic along the margins of the Ionian Tethys. The peritidal sediments consist of meter-scale shallowing-upward cycles formed by subtidal, intertidal, and supratidal facies. Three main informal units are differentiated along the section on the basis of the variations recorded by the subtidal facies. The lower and middle units are attributed to the Rhaetian, on the basis of the common presence of the foraminifer Triasina hantkeni, associated with several benthic foraminifers, such as Aulotortus sinuosus and Auloconus permodiscoides. Megalodontids are particularly abundant and large in the lower unit, whereas they become rare in the middle unit and disappear in the upper unit. The last occurrence of T. hantkeni, along with the disappearance of the benthic foraminifer fauna, and the bloom of the calcareous alga Thaumatoporella parvovesiculifera is assumed as a proxy of the Rhaetian–Hettangian boundary. Recovery biota during the early Jurassic occurs about 20 m upward of the boundary zone, marked by the appearance of benthic foraminifers, such as Siphovalvulina sp. The observed biostratigraphic signature in the studied section is easily comparable to similar Tethyan sections already described from Italy, Greece, and Turkey; thus, it is believed that the faunistic turnover does not reflect local facies variations, in response to changes in the accommodation space of the platform, but regional changes in a more wide area of ocean Tethys.     

AMMONOIDS AND THE TRIASSIC/JURASSIC BOUNDARY IN THE HIMALAYAS OF SOUTHERN TIBET

Abstract:  The sections of Germig in the Nyalam area, southern Tibet, provide a continuous exposure of ammonoid-bearing, uppermost Triassic and basal Jurassic strata. Eighteen species (three of them new) are described and illustrated: Choristoceras marshi Hauer , C. aff. marshi, C. cf. nobile Mojsisovics , C. nyalamense sp. nov., Eopsiloceras germigense sp. nov. , Pleuroacanthites aff. biformis (Sowerby) , Rhacophyllites sp., Nevadaphyllites cf. psilomorphus (Neumayr), Neophyllites sp. indet., Neophyllites cf. biptychus (Lange), Psiloceras tibeticum sp. nov., P. calliphyllum (Neumayr), Euphyllites cf. struckmanni (Neumayr), Discamphiceras pleuronotum (Canavari), Alsatites spp., Kammerkarites frigga , and K. sp. The ammonoid fauna shows a strong affinity to that of the Northern Calcareous Alps, although diversity in the Calliphyllum Zone is markedly lower. The ammonoid succession across the Triassic/Jurassic boundary is subdivided into four zones: the Rhaetian Marshi, the basal Hettangian Tibeticum, the lower Hettangian Calliphyllum, and the middle Hettangian Pleuronotum zones. It is the only known succession across the Triassic/Jurassic boundary in the Tethyan Realm that is not condensed. The Marshi and Calliphyllum zones are correlated with the same zones in the Northern Calcareous Alps. The Tibeticum Zone, a new local zone, is transitional between the Marshi and the Calliphyllum zones in that it yields both choristoceratids and psiloceratids. Its base is taken to mark the base of the Jurassic System in the eastern Tethys.     

Triassic temnospondyl biostratigraphy, biochronology and correlation of the German Buntsandstein and North American Moenkopi Formation

Based on temnospondyl distributions, we propose a correlation of the non-marine Lower and Middle Triassic strata of Germany and the western United States. This correlation relies principally on the presence of two genera, Nonesian (late Early Triassic) Parotosuchus and Perovkan (early Middle Triassic) Eocyclotosaurus . The distribution of Parotosuchus helgolandicus indicates that the Torrey Member of the Moenkopi Formation in Utah, USA is equivalent to the Volpriehausen Formation of the German Buntsandstein of Nonesian age. Magnetostratigraphy and the distribution of Eocyclotosaurus indicate that the Perovkan Holbrook Member of the Moenkopi Formation in Arizona corresponds to a short interval of Anisian time, as does the German Upper Buntsandstein. The Buntsandstein and Moenkopi biostratigraphic schemes agree with a recently proposed biochronology of Triassic terrestrial tetrapods.     

Biostratigraphy and geometric morphometrics of conchostracans (Crustacea,Branchiopoda) from the Late Triassic fissure deposits of Cromhall Quarry,UK

The enigmatic fissure deposits of south‐western England and southern Wales are famous for their unique assemblage of Late Triassic vertebrates, although their age is contentious. While recent studies of palynomorphs have dated some as Rhaetian, their conchostracan (Crustacea, Branchiopoda) assemblages have not been described in detail nor used in biostratigraphy. We find that species determination of British Late Triassic conchostracans requires detailed observations of size, shape and ornamentation. We provide evidence that although Euestheria brodieana is invariably smaller than E. minuta, with some slight differences in carapace ornamentation, the traditional view that they are very similar is upheld. The use of conchostracans as a biostratigraphical tool is here tested by application to the British Triassic fissures at Cromhall quarry where the usual stratigraphical evidence provided by superposition is absent. We find no distinction between conchostracans from bedded Rhaetian deposits of the UK and specimens collected from the fissure deposits of Cromhall Quarry, Gloucestershire, supporting a late Rhaetian age for these deposits.     

泥河湾盆地上新世小哺乳动物生物地层学及相关问题讨论

在泥河湾盆地稻地组与壶流河组、蔚县组命名剖面上分别筛洗小哺乳动物化石。其中花豹沟剖面选取 3个层位 ,发现小哺乳动物化石 1 0种 ;稻地剖面 6个层位 ,发现化石 2 5种。结合小哺乳动物化石组合 ,以及岩石地层追索对比 ,认为壶流河组与蔚县组从岩性上无法分开 ,应予合并 ,保留蔚县组 ;稻地组与蔚县组为同时期沉积 ,二者之间存在一定的相变关系 ,建议废止稻地组 ;仅保留蔚县组作为泥河湾组之下风成红黏土之上的岩石地层单位 ,其时代为中上新世晚期至晚上新世。     

End‐Triassic bivalve extinction: Lombardian Alps,Italy

A major biotic crisis affecting virtually all major marine invertebrate clades occurred at the close of the Triassic. Species‐level data on bivalves from the Lombardian Alps of Italy record the extinction and suggest a possible causal mechanism. A significant decline in species richness is observed during the lower Rhaetian, where 51% of bivalve species, equally distributed among infaunal and epifaunal filter‐feeders, went extinct. The taxonomic loss at the middle Rhaetian was more severe, where 71% of the bivalve species were eliminated, including all infaunal and 50% of the epifaunal species. The data indicate that the extinction selectively eliminated infaunal bivalves.

An initial loss of bivalve species richness during the middle and upper Rhaetian correlates with changes in sedimentary facies related to a fall in relative sea level. This sea level fall is marked by the onset of peritidal micrites and shifting ooid shoals which may have rendered substrates unsuitable for both epifaunal and infaunal bivalves. The possible influences of temperature and salinity fluctuations are difficult to assess, but they may also have had a deleterious effect on the local bivalve fauna. The loss due to peritidal conditions is not consistent with the selective survivorship of epifaunal taxa recurring in overlying Jurassic rocks.

We propose that physiologic differences and selective resistance to physical stress are consistent with the pattern of selective extinction. Facies shifts associated with the marine regression are not sufficient to account for the extremely high magnitude of infaunal extinction. This selection against infaunal bivalves is probably caused by their decreased capacity to filter feed relative to their metabolic demands. A decrease in primary productivity could have selectively eliminated the infauna. Oceanographic processes or atmospheric darkening, perhaps caused by an extraterrestrial impact, could drastically limit food resources (primary productivity) and is consistent with the selective extinction at the end of the Triassic.