Répartition stratigraphique et biozones des kystes de dinoflagellés au passage Jurassique moyen–Jurassique supérieur (Bathonien supérieur–Oxfordien inférieur) dans le Bassin de Guercif,Maroc nord-oriental

Despite the stratigraphical significance of dinoflagellate cysts as reliable markers for correlating and dating Jurassic–Cretaceous strata, investigations into this palynomorph group in the southern Tethyan Realm, specifically northwest Africa, are sparse and somewhat parochial. Most research on Jurassic dinoflagellate cysts is focussed on European depocentres in the Boreal and Sub-Boreal realms. This study is on biostratigraphical data from two petroleum boreholes (MSD1 and KDH1) drilled during 1985 in the Guercif Basin, northeast Morocco by ONAREP (Office National de Recherche et d’Exploitation Pétrolier), now ONHYM (Office National des Hydrocarbures et des Mines). These boreholes penetrated a thick siliciclastic succession, attributed to the Middle and Upper Jurassic, below Miocene marls. Over sixty dinoflagellate cyst taxa were identified. Four dinoflagellate cyst biozones, named GI to GIV, are established for the late Bathonian to early Oxfordian interval. These biozones are defined on the basis of the first appearance datum (FAD) and/or the last appearance datum (LAD) of some biomarker taxa which have wide geographical distributions. These are: Ctenidodinium combazii and Ctenidodinium sellwoodii for the GI Biozone (late Bathonian–early Callovian); Ctenidodinium continuum and Meiourogonyaulax caytonensis for the GII Biozone (middle Callovian); Gonyaulacysta centriconnata and Wanaea thysanota for the GIII Biozone (late Callovian–earliest Oxfordian); and Liesbergia liesbergensis and Systematophora penicillata for the GIV Biozone (early Oxfordian). These biozones are correlated to those already established for the respective intervals in other palaeogeographic regions, such as the Boreal, Sub-boreal and Tethyan realms.     

Le genre RichterellaAvram (Ammonitina,Périsphinctidés) dans le Tithonique inférieur de la bordure ardéchoise (Sud-Est de la France): Dimorphisme et variabilité

Dimorphism in the genus RichterellaAvram is confirmedon the basis of material collected in Ardeche (South east France) principally in the beds of the Fallauxi zone, from the localities of Le Pouzin and Broyon. The microconch, R. richteri (Oppel), lacks lateral lappets but a ventral horn, formed by the strongly projected forward secondary ribs, is present. The macroconch is «Lemencia praerichteriDonze & Enay. These two forms are united in the same species, R. richteri, and then in the same genus, Richterella. The microconch, which is generally better preserved than macroconch in the studied material, shows a greater variability of ribbing than previously thought, due to the poor illustrations of the lectotype of richteri (reproduced here) which showed only dichotomous ribs. The genus Richterella loses its original dimorphic sense and since its origins and its possible derivations are unknown, it became a taxon defined on morphological bases. The forms from Ardeche bear a greater resemblance to the Spanish ones than they do to the Roumanian forms. The stratigraphical and faunal contexts of Richterella in Ardeche are discussed in this work.     

Hommes et climats au Paléolithique supérieur dans le bassin de la Somme (France)

Nearly 40 years of intensive research conducted in the Somme basin has demonstrated the discontinuity in the human occupation of the area during the Upper Palaeolithic. The northern position of the Somme basin within the loessic region appears to be an explanatory factor in this discontinuity, the nature of which increasingly appears related to climate and environmental factors. Archaeological occupations have been systematically attributed to phases of warming or climate improvement. On the other hand, an absence of human occupation has been consistently observed during the coldest periods. The extreme poverty of the biomass contemporary to the loessic deposits of the Upper Pleniglacial could partly explain this discontinuity specific to the far northwest of Europe (southern England, northern Belgium, Netherlands, and northern Germany). Although only 150 or 200 km away, the southern Paris Basin, which is located on the fringes of the loessic region of northern France and Normandy, presents a more continuous human occupation, which appears to have benefitted from more favourable conditions.     

Découverte d'une faune de vertébrés dans l'Oligocène inférieur de Vayres-sur-Essonne (bassin de Paris,France) : biodiversité et paléoenvironnement

The new rich vertebrates deposit from the Lower Oligocene, at Vayres-sur-Essonne (Essonne, France), contains 34 marine and seven continental taxa, 17 taxa being recorded for the first time from the Paris Basin. The richness of the ichtyofauna is higher than in anyone of the Parisian Oligocene deposits and equals the one of the ‘Argiles de Boom’ (Belgian Rupelian). Its affinities with the septentrional ichtyofauna is confirmed, meridional elements being rare. The terrestrial fauna, the oldest known from stratotypical Stampian (base of the Upper Stampian) shares affinities with the one of Montalbán and attests the occurrence of nearly freshwater environments. To cite this article: D. Merle et al., C. R. Palevol 1 (2002) 111–116.     

Suoidea du Miocène inférieur de Laugnac (Lot-et-Garonne, France)

The fossil vertebrate locality of Laugnac is the type locality of the Neogene mammalian zone MN2b. It has yielded many remains of Suoidea belonging to two different genera.Xenohyus venitor is characterized by its large size and especially its large central upper incisors, I1/ and I2/. It is quite difficult to know its phyletic relationships because the material is not abundant.Hyotherium cf.meisneri is more abundant with a quite good skull, pieces of skull, jaws and isolated teeth. It belongs to a peculiar lineage different from that ofH. major from Saint Gérand-le-Puy, France. It has some similarities withAureliachoerus aurelianensis from later geological levels (MN3-MN5).     

Modalités d'installation et d'évolutiondes Harpocératinés (ammonitina A) au Domérien inférieur dans le sud-Ouest de l'Europe (France,Portugal)

In Mesogea, the Harpoceratinae were known asearly as the end of the Lower Carixian. They later reached the sub-boreal province through the Alps or the South of the Iberic Peninsula. The analysis of the populations met in four crosssections of the Lower Domerian (Causses, Mâconnais, Franche-Comté, Portugal) showed the settling down of the Protogrammoceras isseli group that set up and evoluated in a similar way in the different areas in spite of a certain originality.On the one hand, it gave the cladogenetic groupP. monestieri-nitescens by diminution of the ribing density, strengthening of the ribs and simplification of the suture.On the other hand, a slower evolution give, near the end of the Stokesi zone, P. normanianum in the Paris Basin and in Burgundy, and P. lusitanicum in Portugal. At the same time, a new invader, P. celebratum, arrived in the Causses (and in Portugal).     

Nouvelles données sur le Magdalénien inférieur de la Région Cantabrique : le Niveau F de la grotte de El Cierro (Ribadesella,Asturies, Espagne)

El Cierro Cave (Ribadesella, Asturias, Spain), located near the mouth of the River Sella, has yielded one of the most important Upper Palaeolithic sequences in northern Spain. To date, three major occupation periods at the cave have been identified and dated. The first was at the beginning of the Holocene (ca. 8500 BP; ca. 9000 cal BP); the second at the end of the Upper Palaeolithic, in the Younger Dryas (YD) or Greenland Stadial 1 (GS1) (ca. 11.200 BP; 12.700 cal BP) and the third during Greenland Stadial 2 (GS2) (ca. 16.300–15.500 BP; ca. 19.200–18.700 cal BP). This paper describes the stratigraphy documented in the excavations performed by F. Jordá Cerdá and A. Gómez-Fuentes between 1977 and 1979 and presents the first radiocarbon determinations for the first two occupation periods, together with the study of the archaeological materials found in Level F. This level, dated to 15.500 BP (ca. 18.700 cal BP) is characterised by specialised red deer hunting and the gathering of marine resources (winkles). Various artefacts made from animal raw materials have been documented; both finished products and items in the process of being manufactured, as well as portable art objects. The lithic assemblage, consisting mainly of local raw materials with a small proportion of allochthonous flint, is characterised by an abundance of small bladelet cores and backed bladelets. These archaeological remains and the radiocarbon date mean Level F can be attributed to the so-called “Cantabrian Lower Magdalenian”. This period has been documented archaeologically and dated to a similar time at other sites in the River Sella valley and in the rest of northern Spain.     

Le Paléolithique supérieur de la région centre de l’île de Honshû

Tokai area is situated in the middle of Japan and has various configurations of ground. The eastern area has thick natural layers with volcanic ash which has been supplied from Mt. Fuji. About five chronological stages of Paleolithic culture from trapeze and edge-ground axe culture to microlith culture through backed blade culture have been found in the layers. These stages of Paleolithic culture are indicators of chronological studies in Tokai area. On the other hand, it is difficult to advance chronological studies in the middle east, middle, and west areas, because they have little thickness of layers with volcanic ash. However, different Paleolithic artifacts are found in top and bottom layers which put the AT layer between at Tsubakibora Site in Gifu Prefecture, and they have helped very much chronological studies of Paleolithic culture in the west area. Here is the first result of Paleolithic studies in Tokai area with artifacts which have been found on the forth layer of scoriae at Idemaruyama Site are dated about 35,000 years, one of the oldest Paleolithic culture in Japan. In addition, some lines of pits which were dated 30,000 years were found at Hatsunegahara Site, and they provided valuable data to study Paleolithic hunting. Finally, a holed pendant which had 10 notches on one side was unearthed from Fujiishi Site, and it was dated 19,000 years ago. At Terada and Hino Site, a flat big stone which had carvings was found, though the shape and use were seemed to be different from the pendant at Fujiishi Site.     

Nouvelles espèces de Madréporaires dans le Kimméridgien supérieur du Jura (France)

The study of an Upper Kimmeridgian madreporarianfauna collected in the southern french Jura (between Valfin and Belley), allowed: 1) to describe five Scleractinian new species: Mitrodendron biennensis nov. sp., Hexapetalum nemorosum nov. sp., Thalamocoenia gironensis nov. sp., Cryptocoenia stelliserrata nov. sp., Columnocoenia miniflora nov. sp. and a new variety of Heliocoenia (Kobycoenia) claudiopolisensisBeauv., 1964, characterized by its small calices; 2) to modify the diagnosis and the systematic position of the species Baryhelia crassa (de From.) which was created as Heterocoenia crassade From.; 3) to increase of a new species the genus Hexapetalum only known, till now, by its Stramberg Tithon type-species; and 4) to give some palaeoenvironmental conclusions = the Upper Jurassic reef formations in the Southern Jura grew in shallow water characterized by an abundant carbonate sedimentation. Genera associations and distribution confirm the presence, between Oyonnax and Belley, of a low energy area, protected by a barrier reef and, northern of this barrier, of a higher hydrodynamic environment.     

Sur les transports de réserves et les transports de couvain dans la ruche

Résumé Dans cet exposé, nous avons montré que les Abeilles sont capables de changer de place les réserves sucrées et le couvain. Ces comportements ne correspondent pas à des conditions de vie normales de la colonie.Cependant, le transport des réserves indique la possibilité, pour les Abeilles, de réagir à une perturbation importante selon un ordre que nous avions déjà établi dans l'élevage des larves (Montagner, 1962). C'est ainsi qu'en cas de nourrissement brutal et pendant une période de disette, elles tendent à grouper la majeure partie des réserves près de la reine, tout en laissant une place sur un cadre pour la ponte. La reine représenterait donc bien, dans la ruche, le potentiel survie à partir duquel les ouvrières auraient tendance à répartir leurs activités de façon décroissante.Par contre, les transports de couvain ne semblent répondre à aucun plan d'organisation comme chez les Fourmis. Nous avons montré qu'ils étaient associés à de profondes perturbations, telles que le manque de nourriture ou la perte de la reine.

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Summary The results of these studies indicate that bees are able to remove sugary stores and brood.We have established that bees remove their stores according to a certain order that we also made evident when studying brood rearing (Montagner, 1962). So, the bees hoarded the most part of their stores in the side of the hive where the queen stood, when we fed them suddenly during a dearth time.Then, the queen would be in beehive the attractive center from which the workers would share theirs activities in a decreasing way.On the contrary, it never seemed to us that the removals of brood were connected to any right order as for ants. We have established that they were associated to great troubles in beehive such as a want of food or the death of the queen.

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Zusammenfassung In diesem Bericht haben wir bewiesen, dass die Bienen die Zuckervorräte und die Brut an andere Plätze zu bringen vermögen. Dieses Verhalten entspricht nich den normalen Lebenszuständen der Kolonie.Soch bezeugt der Transport der Vorräte die Möglichkeit für die Bienen, gegen eine erhebliche Störung, nach einer bestimmten Regel zu reagieren, die wir schon bei der Aufzucht der Larven festgestellt haben (Montagner, 1962).Also, im Falle einer plötzlich verstärkten Zufuhr und während einer Mangelperiode haben sie die Neigung, den grössten Teil der Vorräte um die Königin zu sammeln, indem sie aber einen Platz für das Eierlegen auf einem Rahmen frei lassen. Die Königin würde also im Bienenstock die Möglichkeit der Fortdauer darstellen. Von dieser Tatsache ausgehend würden die Arbeiterinnen also ihre Tätigkeit in abnehmender Weise ausüben.Im Gegenteil scheinen die Brutversetzungen keinem Organisationsplan zu entsprechen, wie es bei den Ameisen der Fall ist. Wir haben gezeigt, dass sie mit schwerwiegenden Störungen, wie z. B. dem Nahrungsmangel oder dem Tod der Königin, in Verbindung stehen.

     

Les Pachyerymnoceras (Pachyceratidés,Périsphinctacés,Ammonites) du Callovien moyen et supérieur de la région de Saida (Algérie occidentale). Origine phylétique et biogéographie des Pachyceratidés

The genus Pachyerymnoceras, with four new species,is described for the first time in Western Algeria. The genus first appears in the Middle Callovia (Coronatum Zone) as in Saudi Arabia. It is represented in the Saida region by uncommon P. praecox nov. sp. followed during the Upper Callovian (Athleta Zone) by specimens belonging to P. flamandi sp. nov., P. kmerense nov. sp. and P. saidense nov. sp. Every species is represented by a dimorphic pair.Pachyceratidae are put into Perisphinctaceae. If Pachyerymnoceras arises from Erymnoceras, a genus suddenly appearing during the Middle Callovian on northern and southern Tethyan borders, the origin of Erymnoceras is still unknown. Three phylogenetic hypothesis are considered here. Erymnoceras may have evolved during Lower Callovian: 1) from serpenticonic Tethyan Pseudoperisphinctinae like Choffatia (Subgrossouvria); 2) from Tulitids via ellipticoncic Bullatimorphites (Kheraiceras); 3) through a double lineage arisen in Subboreal and Tethyan platyconic Proplanulitids.Pachyceratids have a wide paleobiogeographic distribution in NW Europe and on the Tethyan margins. The genus Pachyerymnoceras evolves in Ethiopian Province of the Tethyan Realm. Some species migrate into Indo-Malagasian Province and others, through the south Tethyan border came to Europe were they are found restricted to Upper Callovian (Athleta and Lamberti Zones). Therefore this migration has a duration of two or even three zones.     

Découverte de charophytes et ostracodes de l’Yprésien inférieur dans les Monts des Ksour (Algérie) : biostratigraphie et paléoécologie

A stratigraphic, palaeontological and sedimentological study was carried out in the red beds cropping out on the left side of Oued Tafarahit, south-east of the Ksour Mountains (Algeria). The studied succession consists of a non-marine fining-upward detrital formation, including microconglomerate, sandstone and clay beds, which were previously attributed to the Cenozoic sensu lato, since no palaeontological evidence was available. Sedimentology suggests deposition in a fluvial environment. The clayey levels yielded the assemblage formed by the charophytes Peckichara atlasensis, Maedleriella cristellata, Nitellopsis (Tectochara) thaleri, Grovesichara sp., and Lamprothamnium papulosum and the ostracods Neocyprideis meguerchiensis, Herpetocypris? sp. and Cyprinotus? sp. This assemblage allows constraining the age of the detrital series from Oued Tafarahit to the Ypresian. Ostracods are typical of freshwater to euryhaline environments. The freshwater taxa (Herpetocypris? sp. and Cyprinotus? sp.) would indicate phases of desalinisation during periods of flood. However, the brackish water species (Neocyprideis meguerchiensis) is characteristic of saline phases related to low water table periods. Moreover the charophyte and ostracod assemblage confirms a close palaeobiogeographic relationship between North Africa and Southern Europe during the Lower Eocene.     

Eodmitria,genre nouveau de Brachiopode Cyrtospiriferidaedu Frasnien inférieur et moyen

A new genus Eodmitria (Brachiopod Spiriferida)with type-species: Spirifer supradisjunctus boloninensis n. subsp., typical of the lower and middle Frasnian, is defined. It is akin to DmitriaSidiachenko, 1961.     

Une nouvelle faune de vertébrés continentaux,associée à des artefacts dans le Pléistocène inférieur de l’Hérault (Sud de la France), vers 1,57 Ma

A new vertebrate fauna associated with lithic artefacts from the Early Pleistocene of the Hérault Valley (southern France) dated around 1.57 Ma. Some lithic artefacts associated with an Early Pleistocene (Upper Villafranchian) vertebrate fossil assemblage have been found from a quarry exploited for basalt in the lower Hérault Valley (Languedoc, southern France) at the Lézignan-le-Cèbe locality. A preliminary patrimony expertise led us to identify about 20 vertebrate taxa, and the autumnal rainfalls revealed the presence of roughly 30 lithic artefacts of “pebble culture” type. A basalt layer dated at 1.57 My directly overlies the fossiliferous level, extends along the little hill (locus 2) yielding artefacts. These new promising data offer new perspectives to improve our understanding of Early Pleistocene ecosystems (and possibly ancient hominin occupation) of southern Europe.     

Nouvelles données paléontologiques,paléoécologiques et sédimentologiques sur le Berriasien de la région de Lamoricière (ouled Mimoun et E1 Rhoraf,Monts de Tlemcen,Algérie)

In the Western Algerian area, the Ouled Mimoun and El Rhoraf sections are particularly appropriate for the study concerning the berriasian formations.From tha lithostratigraphical point of view, the upper section of the carbonated platform corresponds to the lower Berriasian and a part of the middle Berriasian. The «Argiles de Lamoricière which dominate it belong to the middle and upper Berriasian. The subjacent detritic formation known as «Grès de Berthelot probably begins with the lower Valanginian. The stratigraphical results are based on an abundant fauna (ammonites, ostracods) collected in situ, the main species of which have been the subject of a paleontological review and abondantly illustrated.The studied series offer a good example of the evolution of sedimentary conditions which accompany the temporary opening of a carbonated platform towards the open sea.The variations in fauna composition are closely related to changes in environment. In particular, the presence of ammonites in the Boissieri zone can be connected to the highest point of marine advance, although the depth remained relatively shallow (an upper infratidal zone being likely). It is to note that during this period the low level of sedimentary activity favored the settling of some remarkable epilithic fauna.Above the «Argiles de Lamoricière, the massive detritic sedimentation (grès de Berthelot) could indicate an important paleogeographical change in the interior. In the area studied, the ostracods encountered in the basal section of the last formation still mean a shallow marine environment.     

Trace nouvelle de locomotion de Chélonienet figures d'émersion associées dans les calcaires lithographiques de Cerin (Kimméridgien supérieur,Ain, France)

The first tetrapod Vertebrate track discovered in the late Kimmeridgian lithographic limestones of Cerin (Ain-France), is descrbed as a new ichnospecies, named Chelonichnium cerinense. The characteristics of the track, which is about 7 m long, suggest that it was made by a large Chelonian, but none of the fossil turtles known so far from Cerin is likely to be the author of these footprints. The ground on which the animal walked was apparently very wet, superficially soft, and sloping. Moreover, this track is associated to soft sediment slides, confirming emersion and slope. The depositional environment is interpreted as a platform located behind a «ribbon of islands. This irregular surface was at times emerged and terrestrial tetrapods could then leave tracks on the superficial layer.     

Mise en évidence de la zone à Platynota (Kimméridgien inférieur) dans le Massif du Montejunto (Portugal); conséquences stratigraphiques et paléontologiques

The section has been logged on the southern slope of the Montejunto Mountain, near Cabanas de Torres (North of Tage River, Portugal). Typical ammonites of the Platynota zone with species-index Sutneria platynota (Reinecke) have been found in the lower part of the «Marnes d'Abadia formation. Orthosphinctes and Desmoides subzones of early Kimmeridgian are paleontologically well characterized. Unlike what is usually thought, the lower Kimmeridgian does occur. More fossiliferous beds of the «Marnocalcaires de Tojeira formation are coincident with Upper Oxfordian (Planula zone and Upper part of the Bimammatum zone). They yield Ardescia gr. enayi Atrops which appear earlier in Portugal than in South-eastern France.     

Découverte de faunes d'Ammonites de la zone à Platynota(Kimméridgien inférieur) dans les Monts de Chellala (avant-pays Tellien,Algérie); Conséquences stratigraphiques et paléogéographiques

In Chellala mountains (Seba el Abiod), in Western Algeria, new biostratigraphic data, based on Ammonites, allow characterize, for the first time in North Africa, the Platynota zone with the species-index Sutneria platynota. This species was known, to the present time, only in the South European border. These new biostratigraphic data show the diachronism of flyschoid facies. They do not exceed middle Oxfordian in the Tlemcenian zone; on the contrary they go up to lower Kimmeridgian in the preatlasic zone.