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1.
The Cabo Mondego outcrops exposed along the cliffs, on the western margin of the Iberian Plate, show an expanded stratigraphic section of Lower Bathonian deposits containing abundant ammonoids. Upper Bajocian deposits correspond to similar facies, of muddy limestones alternating with marlstones, although ammonoids are scarce. A detailed succession of ammonites across the Bajocian/Bathonian boundary has been recognized at Cabo Mondego, which can form a useful bio‐ and chronostratigraphic standard for the Lusitanian Basin. The revision of previous collections from the classical section and new field samplings of two other separate sections allow the recognition through up to twenty metres of thickness, the highest zone of Bajocian (Parkinsoni Zone) and the lowest zone of Bathonian (Zigzag Zone). The Parkinsoni and the Zigzag zones established for NW European areas and belonging to the Northwest European Province, can be identified in the Lusitanian Basin, although the ammonite fossil assemblages are composed of Submediterranean taxa. However, a subdivision of the Parkinsoni Zone is not possible, due to the scarcity of well preserved ammonoids. The Zigzag Zone can be recognized and characterized as composed of two subunits (Parvum and Macrescens subzones) as represented in diverse European basins of the Submediterranean Province. Ammonite fossil assemblages of the Parvum Subzone may be grouped into two successive horizons, which are biochronostratigraphically equivalent to the subdivisions of the Convergens Subzone distinguished in the Digne‐Barrême area (SE France). New biochronostratigraphic data on the Bigotitinae, youngest members of Leptosphinctinae and oldest members of Zigzagiceratinae are relevant in understanding the evolution and faunal turnover of the West Tethyan Perisphinctidae during earliest Bathonian. The ammonite succession at the Bajocian/Bathonian boundary in the Cabo Mondego region (Portugal) represents one of the most complete biostratigraphic records so far recognized on the Iberian Plate.  相似文献   

2.
Jacques Thierry 《Geobios》1976,9(3):291-331
From Bajocian to lower Kimmeridgian, the subfamilyStephanocerataceae represents a large stock of the ammonite fauna. Though it is tributary of author's stratigraphical correlations and systematical interpretations, its geographical distribution with a drifting continents concept gives many teaching about the study of this superfamily, the continental drift and its consequences. Showing a world wide repartition at its apparition this stock then presents branches; the one (Stephanoceratidae) is not envisaged here; the second (Sphaeroceratidae and Cardioceratidae families) characterises rather the «boreal province; the third (Macrocephalitidae family) is chiefly specific of the «tethysian province. A «north-east pacific province with boreal affinities and a «south-east province with tethysian affinities are delimited. It is probably from its fauna that the tethysian macrocephalitid stock appears; first represented by ammonites specifically north american in middle and upper Bathonian stage, they extend during upper Bathonian and lower Callovian and reach South America and the Tethys. The presence of a narrow sea, on Patagonia, West Antarctica, East Africa, Madagascar and India, joining the Tethys and the south edge of Gondwania, is proved at various times, at Bajocian and Callovian stages.  相似文献   

3.
The new forma aegrafenestra n. f. sensuHölder (1956) is introduced to describe healed injuries to the body Chambers of Jurassic/Cretaceous ammonites, primarily behind the peristome but normally excluding the peristome itself. The different outlines and positions of these window-like sublethal injuries indicate two different Subtypes of injury inflicted by two different types of predator. Some of the large-scale ventral injuries occurring since the Late Devonian were presumably due to attacks by durophagous vertebrates, whereas the smaller, more or less rounded and mainly lateral puncture marks in Liassic to Cretaceous ammonite Shells can be traced back to unsuccessful attacks by stomatopod crustaceans. Identification of the specific predators responsible for characteristic injuries will give information about synecological relationships and life-modes of the attacked ammonites.  相似文献   

4.
Iterative evolution of Middle Jurassic ammonite faunas   总被引:2,自引:0,他引:2  
The phenomenon of iterative evolution of homeomorphs in identical temporal sequences is exhibited by Aalenian and Bajocian ammonite faunas in the south German depositional basin. Within each evolutionary cycle, inflated, evolute, and ornamented shells with complex suture lines are successively replaced with discoidal, involute, smooth shells with simpler sutures. Three distinct and identical cycles of morphological change occur, and involve ammonites belonging to four different familial/subfamilial groups: the Leioceratinae, Graphoceratinae, Hammatoceratidae, and Sonniniidae. Geometric and statistical analyses of morphological change within each of these higher taxa reveal remarkable iteration in both the pattern and proportion of change with time. The iterative morphological cycles were not driven by orthogenesis or anagenetic transformation within lineages, as has been previously suggested. A clear correlation exists between the successive ammonite faunas and repeated cyclical changes in the physical marine environment of the depositional basin. D Ammonoidea, Aalenian, Bajocian, south Germany, iterative evolution.  相似文献   

5.
Unusual phosphatic casts of the ammonites Mortoniceras (Subschloenbachia) sp. and Stoliczkaia sp. from the upper Albian condensed phosphorite bed at Annopol, Poland, are discussed in terms of their taphonomic history. These specimens are interpreted as ‘secondary’ external casts of ammonite replicas preserved originally as attachment scars on oyster shells. The following genetic history is suggested for this previously undocumented mode of ammonite preservation: (1) settling of shells of dead ammonites on the seafloor; (2) colonization of these shells by oysters and formation of ammonite replicas on left valves of oysters; (3) dissolution of ammonite shells; (4) reworking and fragmentation of oyster shells; (5) casting of ammonite replicas by phosphatic material; and (6) separation of ammonite casts from oyster shells, either through mechanical disintegration or dissolution of the latter. The specimens studied were formed after dissolution of the ammonite conchs, not prior to this event as in the case of typical ammonite steinkerns (internal moulds). Therefore, they are here referred to as ‘pseudo‐steinkerns’. The time interval between loss of the original ammonite shells and the formation of oyster‐mediated pseudo‐steinkerns may be very extensive. Therefore, the pseudo‐steinkerns may potentially mislead in biostratigraphic dating of condensed phosphorite deposits.  相似文献   

6.
Uppermost Bajocian (Parkinsoni Zone, Bomfordi Subzone) calcitic concretions from the epicratonic basin in South-Central Poland (Polish Jura) contain numerous, well-preserved ammonites unique for this area. Characteristic of the assemblages is the presence of small, juvenile specimens. Associated with dominant parkinsoniids (Parkinsonia (Parkinsonia) aff. dorni Arkell) are lytoceratids [Nannolytoceras tripartitum (Raspail)], lissoceratids [(Lissoceras (Lissoceras) oolithicum (d'Orbigny) and Lissoceras (Microlissoceras) solitarium Zatoń and Marynowski nov. sp.)], leptosphinctids (Vermisphinctes sp.) and strigoceratids [(Strigoceras (Strigoceras) sp. juv. gr. strigifer/pseudostrigifer)], as well as fragments of phylloceratids (Phylloceras sp.). Parkinsoniids (the most numerous) are considered here as host fauna. Other ammonites, especially phylloceratids and nannolytoceratids, are supposed migrants from the Mediterranean area into the shallow epicratonic Polish Basin during the Late Bajocian sea-level rise. N. tripartitum, which has a broad geographical range probably drifted as a plankton-feeder in the upper water column and, together with phylloceratids, are considered to be unsuccessful immigrants which did not adapt to the new environment. It seems that the whole fauna, inclusive of plant remains, was accumulated by bottom currents in randomly situated depressions on the sea-floor, where carcasses became ideal nucleation sites for concretion genesis. Early diagenesis, which played an important role, resulted in well-preserved ammonite assemblages close to the original biological assemblage. Most of described ammonites (nannolytoceratids, lissoceratids, strigoceratids and leptosphinctids) are reported from the Upper Bajocian epicratonic deposits of Poland for the first time. This makes the concretions a true “taphonomic window” into latest Bajocian census biocenosis. The organic geochemistry results suggest predominance of terrestrial organic matter in both concretions and surrounding clays, with well oxygenated conditions in the sedimentary basin. The large concentrations of Δ13(17)diasterenes in the analysed carbonate concretions indicates that these compounds must have been formed by contact with clay minerals before formation of the carbonate concretions. The processes described may have followed early stage of diagenesis, but certainly under tens of cm to even a few meters of burial. The high negative δC13 values in the inner zone of the concretions suggest that concretion growth began in the sulphate reduction zone and may have continued in the methanogenesis zone.  相似文献   

7.
In the Jajarm area (eastern Alborz Mountains, NE Iran) the ?Upper Triassic — Lower Bajocian siliciclastic Shemshak Formation is up to 2000 m thick. Whereas the lower third of the formation is nearly exclusively non-marine, the upper two-thirds are fully marine. The middle part is characterized by several levels containing a relatively diverse and well preserved assemblage of ammonites of the Toarcian to Aalenian stages. Two sections of the ammonite-bearing strata, spaced 20 km, are presented and correlated by means of ammonite biostratigraphy. The ammonite fauna consists of 27 taxa, some of which are recorded for the first time from the Alborz Mountains. The ammonites are briefly described and their palaeobiogeographic context is reviewed. The ammonite fauna of the Shemshak Formation at Jajarm, as elsewhere in North and Central Iran, is exclusively Northwestern Tethyan in character and is closely related to the faunas of Northwestern and Central Europe.  相似文献   

8.
《Geobios》1986,19(6):689-704
Brachiopod species have been collected in lithologicalunits dated with ammonites in the Bajocian and the Bathonian of the Vienne valley (South-West of the Parisian Basin, France) where detailed surveys were carried out. These observations brought out informations about stratigraphical range and palaeoecology of the species.Some of these brachiopods had alrewdy been observedin the Creuse valley (Fischer 1962, 1964). Comparison between the two lithological and faunal sequences in these valleys, allows to give a new interpretation about the stratigraphical sequence in the Creuse valley. The occurrence of Eudesia infrabathonica Fischer in the Upper Bajocian (Parkinsoni Zone) is noticeable.  相似文献   

9.
Late Bajocian to Early Bathonian ammonites, now attributed to Benatinites and Lugariceras, and Late Bathonian Hemigarantia, previously considered as cryptogenic taxa, share very similar sutural and ornamentation characters, suggesting their derivation from the stephanoceratid stock, most probably from the Cadomites-Polyplectites line. The palaeogeographical distribution of these taxa is limited to the North-Western African and European margins of Western Tethys. Palaeontologically, they represent the result of the iterative evolution rather than a poorly known lineage.  相似文献   

10.
Maeda, H. 1991 01 15: Sheltered preservation: a peculiar mode of ammonitc occurrence in the Cretaceous Yeto Group, Hokkaido. north Japan. Lethaia , Vol. 24, pp. 69–82. Oslo. ISSN 0024–1164.
'Sheltered preservation', in which many small ammonites ('refugees') arc preserved in the inside and/or the lower umbilical void of a large ammonite shell ('shelter'), is a common preseivational style in the Cretaceous Yezo Group in Hokkaido, north Japan. The best example is large Calycoceras of Middle Cenomanian age, attaining 300 to 500 mm diameter, whose shell usually hours more than 100 individuals of Desmoceras comprising mainly juveniles less than 12 mm in diameter. In contrdst with other preservational styles of Desmoceras from the same horizon. the shells of refugee ammonites have not suffered phragmoeone collapse, and the inner whorls and a protoconch are intact. Many refugee ammonites are allochthonous, as indicated by the incomplete preservation of their living chamber and the absence of a jaw apparatus. The sheltered preservation is, therefore, a kind of 'Konzcntrat Lagerstatten' completed by both pre- and post-burial processes. Acting as a 'concentration trap' on the sea floor, a large empty ammonite shell probably triggered off such accumulation of the small ammonite shells in post-mortem transport across the sea floor. This biostratinomical process biased the original thanatococnosis of ammonites, so that its immature specimens were selectively accumulated and preserved in the shelter. Ammonoidea, taphonomy, Konzentrat Lagerstätten'. differenrial preservation, Cretuceous. Hokkaido .  相似文献   

11.
Taphonomic analysis of Lower and Middle Oxfordian ammonites from the Cracow Upland, southern Poland (localities at Pod???e, Zalas, M?ynka) revealed differences in ammonite preservation. The studied ammonites, usually termed as external and internal moulds, show a more complex state of preservation. In the Middle Oxfordian glauconitic marls, ammonites are preserved as internal moulds with neomorphic calcite shells showing relics of the original internal structure. In the Middle Oxfordian platy peloidal limestones, ammonites are preserved mostly as external moulds, without septal suture, however under microscope might show relics of internal whorls and septa and/or subtle differences in sediment filling phragmocone chambers. In sponge–microbial bioherms and biostromes, ammonite internal moulds have shells, which in contrast to ammonites from glauconitic marls are not strictly neomorphic ones, but originated by shell dissolution and subsequent filling of moldic porosity by calcite cement. In sponge–microbial nodular limestones, the ammonites are strongly deformed and the outer wall is usually removed by dissolution under pressure. Other important taphonomic differences include the rate of compaction (highest in platy limestones), sedimentary infillings, microborings, encrustations and preservation of siphuncular tubes. The majority of the ammonites appear to be phragmocones; aptychi in all facies are rare. Siphuncular tubes are fossilized exclusively in oppeliids, only in specimens from glauconitic marls and platy limestones, although their other taphonomic attributes are different. Tubes seem to have fossilized due to microbially mediated phosphatization that could be favoured by a set of parameters which operated rather at the scale of ammonoid carcasses: closed, poorly oxygenated conditions, and reduced pH. Taphonomic processes were controlled by the sedimentary environment (fragmentation, sedimentary filling, phosphatization of siphuncular tubes), as well as by early and late diagenesis (neomorphic transformation, dissolution, cementation, compaction) influenced by lithology.  相似文献   

12.
《Comptes Rendus Palevol》2013,12(3):137-148
A study of the ammonite assemblages from the “Oolithe ferrugineuse de Bayeux” Formation of Bajocian age is presented herein. The section at the locality of Maizet shows a high level of stratigraphic condensation, and taphonomic reworking is common within the sequence. All the ammonites being studied are classified herein as having been reworked, transported or displaced on the sea-floor prior to burial, and as such, are determined taphonomically as being resedimented or reelaborated fossil elements. Seven evidences of reelaboration within the sequence under investigation are detailed here. The palaeontological units, so-called taphorecords, characterized by distinctive taphonomic features, are used to directly or indirectly assign beds to biochronostratigraphic units. In addition, identification of taphorecord relationships regarding successive or contemporaneous deposition allows their registratic succession and order of depositional events to be inferred. The latter may be deduced on the basis of bed succession or by reference to stratigraphical intervals that now are only represented in the stratigraphic column by fossils. Deposition by tractional currents and winnowing is indicated by sharp bed-base and by reworked fossil elements. Biochronostratigraphic correlation with other sections of the inland Bajocian successions in the Calvados area (Bretteville, Feuguerolles) highlights a common depositional evolution that may be related to an eastward-deepening carbonate ramp.  相似文献   

13.
Over 50 species of liparoceratid ammonites have previously been recognized from the Ibex and Davoei Zones (Lower Pliensbachian, Lower Jurassic) of England alone. Such a high specific diversity is an artifact caused by their stratigraphical importance and the complex nature of their shells. Simple bivariate analysis indicates that Aegoceras species are arbitrary subdivisions of an allometric growth trend. Principal component multivariate analysis shows that Liparoceras and Androgynoceras (sensu Callomon) are quite distinct from Aegoceras , and they display a more random pattern of variation. The existence of stratigraphically useful size changes in liparoceratids mirrors precisely size increases in benthic molluscan taxa, such as the bivalve 'Astarte' platymorpha and the archaeogastropod Eucyclus subimbricata. The ammonite size changes, at least, probably result from phenotypic variation analogous to that observed in the living cephalopod Sepia officinalis. This evidence taken in conjunction with the fact that kosmoceratid ammonite size changes correspond somewhat with benthic faunal changes suggests that ammonites maybe less useful for stratigraphy than is believed.  相似文献   

14.
Eight clusters of small spherical and subspherical objects, some isolated and some associated with shells of perisphinctid ammonites, have been recovered from the Lower and Upper Kimmeridge Clay (Upper Jurassic) of the Dorset coast, England. They have been interpreted as ammonite egg sacs and represent the freshest and best-preserved examples known so far. Their structures and the ecological framework in which they occur are discussed. The parents are thought to be members of the two eudemic genera Aulacostephanus and Pectinatites that dominate the biostratigraphy of the ammonites in the range of the Kimmeridge Clay in which they occur. Isolated nuclei of ammonitellae have also been recovered.  相似文献   

15.
Distinct faunal aggregates are described from the Middle Jurassic (uppermost Bajocian/lowermost Bathonian and Middle Bathonian) clay deposits of Częstochowa area, south-central Poland. These aggregates are composed of molluscs (scaphopods, gastropods, bivalves, ammonites and belemnites), articulate brachiopods and echinoderms (asteroids, crinoids and echinoids). A large percentage of the fossils, especially bivalves, are fragmented, but some fossils are complete. Although most of the fossils are crushed and fragmented, they are still identifiable to at least the genus level. Thorough statistical analysis of taphonomic features indicates that the preservation of asteroid marginal plates is distinct from the ossicles derived from the host clays. The high frequency of bite marks and the good state of preservation suggest that the accumulations are the products of predation activities and most probably are the effect of regurgitation. Taking into account the rich and diverse fauna, the predator was a bottom-feeding generalist. Possible predators include palaeospinacid sharks, a tooth of which was collected from the same bedding surface, but not associated with regurgitated remains. Although the bite marks on the asteroid ossicles point to sharks as potential producers of regurgitates, other vertebrates, like durophagous pycnodontiform fish, cannot be excluded.  相似文献   

16.
The presumed link between bilateral asymmetry and lateralized habitual activity in extinct hominins is the basis upon which inferences of ‘hand preference’ often derive. While this presumption is reasonable, in-vivo comparisons of skeletal asymmetries and self-reported handedness are rare, and as a result the accuracy of these inferences is questionable. To assess this relationship in living humans, reported ‘handedness’ was compared against peripheral quantitative computed tomography (pQCT) derived bilateral measurements of humeral, ulnar, and tibial midshaft cortical area (CA) and torsional rigidity (J). Significant bilateral differences were found in the humerus for all groups, and in the ulna for the cricketer and field hockey sub-samples. Additionally, cricketers’ non-dominant tibiae were more robust than their dominant tibiae. An assessment of ‘Dominance Asymmetry’ revealed that measures of CA and J were higher in the dominant humeri in ∼90% of participants; in the ulna this was true in ∼75% of cases, and in the tibia CA and J were higher in the dominant limb less than 50% of the time. Comparisons of (self)‘Reported’ hand preference against ‘Predicted’ hand preference (based on the calculation of % Directional Asymmetry) revealed a low level of error for predictions based on both humeral (∼4-5% error) and ulnar (6-11% error) asymmetry. Error was decreased with the exclusion of individuals displaying less than 2.5-5% asymmetry. Contrarily, predictions based on tibial analyses had a much higher level of ‘error’ (∼45%). Overall, the results support currently accepted approaches for inferring ‘hand preference’ from measures of upper limb geometric asymmetry in the hominin skeleton.  相似文献   

17.
Analyses of ammonite shell forms of two Barremian stratigraphic sections from Southwest Mexico consist of two well-defined morphotypes: (1) Small uncoiled, mostly leptoceratoid ancyloconic shells of the families Ancyloceratidae and Hamulinidae, and (2) middle-sized involute to moderately evolute oxycone to discocone shells of the family Pulchelliidae. Index taxa allow the recognition of standard ammonite biozones for the Barremian, which permit the relative dating of different processes that occurred through the water column in the environment of deposition. The vertical distribution of ammonite morphotypes and facies suggests changes of the palaeoceanographic and sedimentological conditions that prevailed in the area during Barremian time. Petrologic data, analyses of the organic carbon and carbonate contents of the rocks support the idea that oxygen-deficient bottom waters existed within a shallow marine, tectonically active area with little carbonate deposition during the early early Barremian (upper part of the Taveraidiscus hugii Zone through the base of the Nicklesia pulchella Zone). These conditions in the basin caused a proliferation of middle-water depth ammonites of Morphotype 1 but prevented the abundance of nektobenthic forms of Morphotype 2. Oxic conditions on a more calcareous and open normal marine environment seem to have been reestablished progressively during a transgressive episode from late early-early late Barremian (upper part of the Nicklesia pulchella Zone through the Gerhardtia sartousiana Zone). This environmental setting supported more facies dependent nektobenthic ammonites of Morphotype 2 to flourish within the basin.  相似文献   

18.
A new ammonite genus of the subfamily Garantianinae, family Stephanoceratidae, from the Upper Bajocian Strenoceras niortense Zone in the Bolshoi Zelenchuk River basin (Karachay-Cherkessia), with two new species from two different localities, is established. The type species Keppleritiana rostovtsevi gen. et sp. nov. is homeomorphic to some species of the Upper Bathonian–Lower Callovian genus Kepplerites (Fam. Kosmoceratidae) but is distinguished by the presence of a ventral furrow in adults. An isolated valve of an aptychus possibly belonging to this species is illustrated. Keppleritiana graebensteini sp. nov. has a more archaic morphology and is apparently ancestral to the type species. The macroconchs and microconchs of both species are described.  相似文献   

19.
西藏北部安多地区中侏罗统(巴通阶-卡洛夫阶)菊石   总被引:1,自引:0,他引:1  
最近从藏北安多地区中侏罗统发现较丰富的菊石类化石,巴通期的菊石有OxyceritesoppeliElmi,Ho moeoplanulitescf. homoeomorphus(Buckman),H. cf. acuticosta(Roemer), Choffatiacf. vicentiMangold, Siemiradzkiacf. matisconensis(Lissajous), Proceritessp., Neuquenicerascf. yokoyamaiKobayashiandFukada和Indosphinctessp.。卡洛夫期的菊石有Oxyceritescf. subcotarius(Oppel), Macrocephalites? sp., Homoeoplanulitescf. furculus(Neumayr),Indospinctes(Elatmites) cf. reveliMangold和Reineckeitessp.。其中Indosphinctes,Indospinctes(Elatmites),Neu queniceras,Siemiradzkia和Procerites等属(或亚属)在西藏北部地区为首次报道。安多地区中—晚巴通期和早卡洛夫期菊石动物群凸现较强的地方性色彩; 晚巴通期ORBIS菊石带和早卡洛夫期KOENIGI菊石带是藏北地区菊石动物群扩散的重要时期。  相似文献   

20.
The life histories of ammonites and the life strategies they employed are difficult to assess without robust modern analogues but placing constraints on ammonite growth rates provides a fundamental first step to understanding this abundant, but poorly understood, fossil group. Here we interpret periodic variations in carbon and oxygen stable isotope profiles from Campanian and Maastrichtian ammonites (Baculites) as seasonally driven and use these records to determine their rate of shell precipitation. Several of these samples are housed in museums and were originally prepared using sealants for display and preservation but testing of these sealants indicated no alteration of the isotopic values of treated carbonate. Diagenetic alteration, as determined by shell microstructure, affected the preservation of isotopic signals, resulting in the loss of seasonal variation in less well‐preserved specimens, and the δ13C signal is more robust than δ18O. The periodicity of isotopic profiles from Baculites shells presented here suggest that these organisms grew at rapid rates (c. 340 mm per year), which may imply an r‐type life strategy in which the animals reach maturity quickly, spawn large quantities of progeny, and die at a young age. Because of the potential mobility of Baculites, reconstructing palaeoenvironmental conditions from these isotopic records is challenging and should be conducted cautiously. Unfortunately, well‐preserved Baculites shells much longer than 350 mm are rarely recovered, which complicates the statistical treatment of potential periodicity in isotopic profiles.  相似文献   

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