Espéces affines microsympatriques chez Puellina (Bryozoa, Cheilostsmata) et description d'espèces nouvelles

The existence of microsympatry (close coexistence in microhabitats) allows morphologically very similar forms belonging to intraspecifically highly variable taxa to be distinguished by consistent microanatomical characteristics at the specific level. Four pairs of closely related species of Puellina Jullien (Bryozoa, Cheilostomata, Cribrimorpha), microsympatric in the Mediterranean Sea to different degrees according to the magnitude of their coexistence, are considered at the systematic, ecological and biogeographic levels. P . ( Cribrilaria ) innominata (Couch) coexists with: (1) P . ( C .) picardi sp. n., in a wide range of littoral and deep habitats; (2) P . ( C .) hincksi (Friedl), in ecological transition zones within sciaphilic littoral communities. P . ( Glabrilaria ) pedunculata Gautier, an apparently Mediterranean endemic with a wide depth range, coexists with: (1) P . ( G .) orientalis orientalis sp. et ssp.n., in deep hard bottoms of the eastern Mediterranean, two other subspecies, P . ( G .) orientalis azorensis ssp.n. and P . ( G .) orientalis lusitanica ssp.n., being distributed in the near Atlantic; (2) P . ( G .) corbula Bishop & Househam, recently described from the English Channel, in cryptic littoral microhabitats of the northwestern Mediterranean.     

Structural and Systematic Studies on Turbellaria Schizorhynchia (Platyhelminthes)

Supplementary structural and systematic records are given on Thylacorhynchus caudatus, Trapichorhynchus tapes (Schizorhynchidae), Karkinorhynchus primitivus, Cheliplana marcusi, C. setosa and Cheliplanilla caudata (Karkinorhynchidae). Cheliplana vaginalis sp.n. and C. evdonini sp.n. are described. The systematics of the Karkinorhynchidae are reviewed and in part revised.     

Description of two new species of Haemaphysalis Koch, 1844 (Acari: Ixodidae) from the H. (Rhipistoma) spinulosa subgroup,parasites of carnivores and other mammals in Africa

Systematic Parasitology - Haemaphysalis (Rhipistoma) latitudinis n. sp. (Acari: Ixodidae) and H. (R.) setosa n. sp. are described based on adults. Adults of H. (R.) latitudinis n. sp. were mostly...     

中国斑织蛾属四新种记述(鳞翅目,织蛾科)

记述中国斑织蛾属Ripeacma Moriuti,Saito et Lewvanich 4新种.梯斑织蛾Ripeacma trapezialis sp.nov.采自贵州梵净山,与疣斑织蛾Ripeacma verruculosa Wang et Li 外表相似,可通过外生殖器特征区别:新种雄性颚形突的侧臂短于腹板,抱器腹背缘无突起,囊形突略呈三角形;雌性交配孔两侧呈齿轮状突出,囊突具小齿.杯斑织蛾Ripeacma cotyliformis sp.nov.采自湖北五峰,与秦岭斑织蛾Ripeacma qinlingensis Wang et Zheng外部特征近似,可通过雄性外生殖器特征加以区别:新种爪形突小三角形,颚形突前缘生出一杯状突起,抱器腹明显具端突,阳茎基环略呈矩形.毛斑织蛾Ripeacma setosa sp.nov.采自贵州梵净山,与Ripeacma yaiensis Moriuti,Saito et Lewvanich相似,但其雄性爪形突短三角形,颚形突不呈带状,抱器腹端突具稠密的刚毛,可与后者区别.茎裂斑织蛾Ripeacma bilobata sp.nov.采自贵州梵净山和湖南桑植,可通过以下特征与该属所有已知种区别:前翅前缘基部及中部偏外各有一个大三角形黑斑延伸至翅中部,雄性颚形突呈"X"状;雌性生殖板强烈骨化.该文同时首次报道了角斑织蛾Ripeacma trigonia Wang et Li的雌性个体.文中提供了新种的成虫及外生殖器特征图.模式标本保存在南开大学生物系.     

Phylogenetic revision of the Charis cleonus complex (Lepidoptera: Riodinidae)

Abstract.  A revision of the Charis cleonus group of Neotropical riodinid butterflies is presented, including discussions on the taxonomy, biogeography and biology of its species, and illustrations of the adults and male and female genitalia (where known) of all taxa. We recognize twenty-two species, including nineteen new species: C. ariquemes sp.n., C. brasilia sp.n., C. breves sp.n., C. cacaulandia sp.n., C. cuiaba sp.n., C. humaita sp.n., C. ipiranga sp.n., C. iquitos sp.n., C. ma sp.n., C. manicore sp.n., C. manu sp.n., C. matic sp.n., C. maues sp.n., C. negro sp.n., C. palcazu sp.n., C. rocana sp.n., C. santarem sp.n., C. tapajos sp.n and C. tefe sp.n. A cladistic analysis using thirty-six characters of male and female genitalia and external facies generated a single most parsimonious cladogram highlighting the existence of two clades, the major and cleonus subgroups. As each species largely occupies a parapatric range, this cladogram provides the opportunity to reassess historical relationships among areas of endemism in the Amazon basin.     

Palaearctic species of the ichneumon-fly genus Campodorus Förster (Hymenoptera, Ichneumonidae): III. Species with long-haired ovipositor sheath, species with entirely rufous hind tibia, and species with white-banded tibiae

A complex of about 40 Palaearctic species of the genus Campodorus with long haired ovipositor sheath and with entirely rufous or white-banded hind tibia is considered and keyed. Twenty-one new species are described. The species with the pale rufous hind tibia have black hind coxa and trochanters, most of them also have rather long genae and scabrous mesopleura. The majority of these species are distributed in subarctic Siberia: C. yakutator sp. n., C. minutor sp. n., C. arctor sp. n., C. atripes sp. n., C. pequenitor sp. n., C. mordax sp. n., C. lucidator sp. n., C. subarctor sp. n., C. nikandrovskii sp. n., and C. genator sp. n. C. suomi sp. n. is described from northern Finland, C. labytnangi sp. n., from Kola Peninsula, the Polar Ural, and Transbaikalia, and only C. gallicator sp. n., from France (? Alps). The species with a white band on the hind tibia are mostly boreal; most of them have red coxae and trochanters (except for C. sakhalinator sp. n. and C. taigator sp. n.), rather short genae, and granulate mesopleura. In this group, eight new species are described: C. atrofemorator sp. n., C. barbator sp. n., C. sakhalinator sp. n. (all from the southern part of the Russian Far East), C. taigator sp. n. (southern Siberian), C. ciliator sp. n., C. melanopygus sp. n., C. marginator sp. n., and C. boreator sp. n. (European-Siberian).     

Four new genera of the PIesiobracon Cameron group (Insecta, Hymenoptera, Braconinae)

Crinibracon gen.n. (type species: Crinibracon malayensis sp.n. from Malaysia), C. indicus sp.n. from India, C. striatus sp.n. from New Guinea, Lyricibracon gen.n. (type species: Lyricibracon bicolorus sp.n. from Brunei), Rugosibracon gen.n. (type species: Rugosibracon maculithorax sp.n. from S. Africa), R. nigerianus sp.n. from Nigeria and Simplicibracon gen.n. (type species: Simplicibracon maculigaster sp.n. from Taiwan) are described and illustrated. A key to the Plesibracon group of genera is provided, together with specific keys for the new genera Crinibracon and Rugosibracon .     

Parental origin and genome evolution in the allopolyploid Iris versicolor

BACKGROUND AIMS: One of the classic examples of an allopolyploid is Iris versicolor, 'Blue Flag' (2n = 108), first studied by Edgar Anderson and later popularized by George Ledyard Stebbins in cytogenetics and evolutionary text-books. It is revisited here using modern molecular and cytogenetic tools to investigate its putative allopolyploid origin involving progenitors of I. virginica (2n = 70) and I. setosa (2n = 38). METHODS: Genomic in situ hybridization (GISH), fluorescent in situ hybridization (FISH) and Southern hybridization with 5S and 18-26S ribosomal DNA (rDNA) probes were used to identify the parental origin of chromosomes, and to study the unit structure, relative abundance and chromosomal location of rDNA sequences. KEY RESULTS: GISH shows that I. versicolor has inherited the sum of the chromosome complement from the two progenitor species. In I. versicolor all the 18-26S rDNA units and loci are inherited from the progenitor of I. virginica, those loci from the I. setosa progenitor are absent. In contrast 5S rDNA loci and units from both progenitors are found, although one of the two 5S loci expected from the I. setosa progenitor is absent. CONCLUSIONS: These data confirm Anderson's hypothesis that I. versicolor is an allopolyploid involving progenitors of I. virginica and I. setosa. The number of 18-26S rDNA loci in I. versicolor is similar to that of progenitor I. virginica, suggestive of a first stage in genome diploidization. The locus loss is targeted at the I. setosa-origin subgenome, and this is discussed in relation to other polyploidy systems.     

Taxonomic revision and phylogenetic analysis of the cestode genus <Emphasis Type="Italic">Paraprogynotaenia</Emphasis> Rysavy, 1966 (Cyclophyllidea: Progynotaeniidae)

The type-series of Paraprogynotaenia jimenezi Rysavy, 1966 [type-species of Paraprogynotaenia Rysavy, 1966] from Charadrius wilsonia Ord in Cuba and P. charadrii (Yamaguti, 1956) (= Proterogynotaenia charadrii Yamaguti, 1956) from C. alexandrinus L. in Japan are redescribed. Two new species are described: Paraprogynotaenia minuta n. sp. from C. alexandrinus on the Black Sea coast of Bulgaria [type-host and type-locality, based on specimens previously reported as P. charadrii], from the same host species in Tunisia and from "Tringa totanus" (the host identification is uncertain) in France [the latter two samples were previously reported as Progynotaenia odhneri Nybelin, 1916]; and Paraprogynotaenia canarisi n. sp. from C. marginatus Vieillot in the Republic of South Africa [type-host and type-locality] and from C. alexandrinus and Pluvialis dominica (Müller) in Taiwan [specimens from both localities were previously reported as Paraprogynotaenia charadrii]. P. jimenezi can be distinguished from its congeners by having 12 hooks of equal length. Among the remaining species with 18-22 hooks of varying length, P. minuta n. sp. can be distinguished by the smaller number of proglottides: up to 8 vs 15-20 for P. charadrii and P. canarisi n. sp. The latter two species can be differentiated from one another by the blade length/total length ratio of the smallest (lateral) rostellar hooks, which is less than 0.5 for P. canarisi n. sp. and 0.5 or more for P. charadrii. An identification key to the species of Paraprogynotaenia is provided and the generic diagnosis is amended. A phylogenetic analysis based on 17 morphological characters resulted in the following hypothesis for the relationships within the genus: (P. jimenezi (P. minuta (P. charadrii, P. canarisi))).     

Pipunculidae (Diptera) from Papua New Guinea: the genera Cephalosphaera, Cephalops and Beckerias

Seven new pipunculid species are described from Papua New Guinea: Cephalosphaera gymne sp.n., C. immodica sp.n. C. wauensis sp.n., Cephalops bifidus sp.n., C. multidenticulatus sp.n., C. papuaensis sp.n., and C. parmatus sp.n. Cephalops argutus (Hardy) is considered as a true Beckerias sp. and thus placed in a new combination: Beckerias argutus. The phylogenetic relationship of the new species is shortly discussed.     

Trois mitrata (echinodermata: stylophora) nouveaux de ľordovicien de tchécoslovaquie

Three new mitrate stylophoran carpoids from the Letná Formation (Lower Caradocian) of Háj, near Zaho?any, Tchecoslovaquia, are described. They are: a mitrocystitid,Aspidocarpus bohemicus n. g., n. sp., a kirkocystid,Anatiferocystis spinosa n. sp., and a species tentatively referred to the anomalocystitids,Barrandeocarpus jaekeli n. g., n. sp.     

Revisión del género Diacodexis (Artiodactyla, Mammalia) en el Eoceno inferior del Noreste de España

Most of the material review in this paper was included in the work of Sudre et al. (1983), some other provides from new sites. Two new species of primitive artiodactyls are described here: Diacodexis corsaensis n. sp. and Eygalayodon isavenaensis n. sp., which is considered the most plesiomorphic form of the subfamily Eurodexeinae and probable derived from Diacodexis corsaensis n. sp. It is for the first time that a representative of the Eurodexeinae is found in the Iberian Peninsula, where they might originate.     

Four new species of Cichlidogyrus Paperna, 1960 (Monogenea,Ancyrocephalidae), all gill parasites from African mouthbreeder tilapias of the genera Sarotherodon and Oreochromis (Pisces,Cichlidae), with a redescription of C. thurstonae Ergens, 1981

A study of Oreochromis niloticus (Linnaeus), O. aureus (Steindachner), Sarotherodon caudomarginatus (Boulenger), S. galilaeus (Linnaeus) and S. galilaeus sanagaensis (Thys van den Audenaerde) (Teleostei, Cichlidae) from different locations in Africa (Burkina Faso, Cameroon, Guinea, Niger and Senegal) revealed the presence of 11 species of monogenean gill parasites. Four, belonging to Cichlidogyrus Paperna, 1960 and considered as new species, are described: C. rognoni n. sp., C. douellouae n. sp., C. giostrai n. sp. and C. njinei n. sp. They are distinguished by the shape and/or size of the sclerotised parts of the haptoral and copulatory complexes. C. thurstonae Ergens, 1981 from O. niloticus is redescribed.     

Phylogeography of the planktonic chaetognath Sagitta setosa reveals isolation in European seas

Numerous planktonic species have disjunct distribution patterns in the world's oceans. However, it is unclear whether these are truly unconnected by gene flow, or whether they are composed of morphologically cryptic species. The marine planktonic chaetognath Sagitta setosa Müller has a discontinuous geographic distribution over the continental shelf in the northeastern Atlantic, Mediterranean Sea, and Black Sea. Morphological variation between these populations has been described, but overlaps and is therefore unsuitable to determine the degree of isolation between populations. To test whether disjunct populations are also genetically disjunct, we sequenced a 504-bp fragment of mitochondrial DNA comprising the cytochrome oxidase II region of 86 individuals. Sequences were highly variable; each represented a different haplotype. Within S. setosa, sequence divergence ranged from 0.2 to 8.1% and strong phylogeographic structure was found, with four main groups corresponding to the northeastern Atlantic, Mediterranean Sea (including Ligurian Sea, Tyrrhenian Sea and Gulf of Gabes), Adriatic Sea, and Black Sea. Two of these (Atlantic and Black Sea) were resolved as monophyletic clades, thus gene flow between disjunct populations of S. setosa has been extremely limited and lineage sorting has taken place. The deepest divergence was between Atlantic and Mediterranean/Black Sea populations followed by a split between Mediterranean and Black Sea populations. The Mediterranean/Black Sea clade comprised three groups, with the Adriatic Sea as the most likely sister clade of the Black Sea. These data are consistent with a colonization of the Black Sea from the Mediterranean. Furthermore, a possible cryptic species was found in the Black Sea with 23.1% sequence divergence from S. setosa. Two possibilities for the evolutionary origin of this species are proposed, namely, that it represents a relict species from the ancient Paratethys, or that it represents another chaetognath species that colonized the Black Sea more recently. Even though the exact timing of disjunction of S. setosa populations remains unclear, on the basis of the geological and paleoclimatic history of the European basins and our estimates of net nucleotide divergence, we suggest that disjunct populations arose through vicariance resulting from the cyclical changes in temperature and sea levels during the Pleistocene. We conclude that these populations have remained disjunct, not because of limited dispersal ability, but because of the inability to maintain viable populations in suboptimal, geographically intermediate areas.     

Taxonomy of European species of Cassagnaudiella and Bourletiella (Collembola, Symphypleona, Bourletiellidae)

The taxonomy of the European members of Cassagnaudiella and Bourletiella (Collembola, Symphypleona, Bourletiellidae) is reviewed. The subgenus Cassagnaudiella Ellis, 197.5 is raised to generic rank with a new definition. Specimens earlier considered to represent a transformed form of Bourletiella radula Gisin, 1946 are referred to Cassagnaudiella , and the application of 'phenotypic neutralization' within the group is Considered erroneous. Four Bourletiella species and six Cassagnaudiella species are described, five of the latter which are new: C. cœnigouensis sp. n., C. luteovernalis sp. n., C. cadiensis sp. n., C. canigouensis sp. n., and C. trichroma sp. n., A key to species is provided.     

Description de deux especes nouvellesd'Atopochara Peck (Clavatoraceae,Charophyta)

The species Atopochara restricta n. sp. which represents an evolutive stage in the lineage established by L. Grambast (1974) is described here, as well as another one: A. brevicellis n. sp.; they are both of presumed Albian age.     

Revision of the subgenus Ceratitis (Pardalaspis) Bezzi, 1918 (Diptera, Tephritidae, Ceratitini)

Abstract. The fruit fly genus Ceratitis (Diptera: Tephritidae) comprises several important pest species attacking a wide range of unrelated fruits. In this paper the subgenus Ceratitis ( Pardalaspis ) Bezzi is revised. Ten species are recognized of which five are described as new: C.(P.)hamata sp.n., C.(P.)munroi sp.n., C.(P.)serrata sp.n., C.(P.)semipunctata sp.n. and C.(P.)zairensis sp.n. A key to both sexes is provided. All species are restricted to the Afrotropical Region and distributional and known host plant data are given.     

New species of Choleoeimeria (Apicomplexa: Eimeriidae), coccidia of bile-bladders of reptiles, illustrating a multiplicity of host cell-parasite interrelations

Oocyst characteristics and histological features of the endogenous development of bile-bladder coccidia of the genus Choleoeimeria Paperna and Landsberg, 1989 are described and the main features for species differentiation are discussed for the following new species: C. allogamae n. sp. from Agama sp., Cameroon, West Africa; C. allogehyrae n. sp. from Gehyra australis, Magnetic Island (type) and mainland N Queensland, Australia; C. boulii n. sp. from Gehyra variegata, SW Queensland, Australia; C. calotesi n. sp. from Calotes mystaceus, Xiang-Mai, Thailand; C. heteronotis n. sp. from Heteronotia binoei, N Queensland, Australia; C. lygosomis n. sp. from Lygosoma buringi, Kon-Kaen, Thailand; C. sylvatica n. sp. from Carlia rhomboidalis, N Queensland, Australia, and C. xiangmaii n. sp. from Hemidactylus frenatus, Xiang-Mai, Thailand. Oocyst characteristic of Choleoeimeria are also reported from Oedura castelnaui, N Queensland. The described species demonstrate a diversity of associations with the bile-bladder epithelial lining, from a single parasite in a single hypertrophic host cell to multiple infections inducing the hypertrophied cells to form stratified layers, or merge into branched clumps.