Holocene vegetation and water level history in two bogs of the Cordillera de Talamanca, Costa Rica

Pollen records of Holocene sediment cores from the Costa Rican Cordillera de Talamanca (La Chonta bog, 2310 m and La Trinidad bog, 2700 m) show the postglacial development of the montane oak forest zone from ca. 9500 to 1500 yr BP. During the early Holocene (ca. 9500–700 yr BP), alder vegetation covered the La Chonta and La Trinidad bogs and their adjacent hills. The upper forest line is inferred to be at 2800–3000 m elevation. A Podocarpus-Quercus forest characterised the middle Holocene (ca. 7000–4500 yr BP). The upper forest line is located at >3000 m reaching the present-day altitudinal distribution. A Quercus forest characterised the late Holocene (ca. 4500–1500 yr BP). Compared to modern conditions, the early Holocene has similar average temperatures, but the moisture level was probably higher. Pollen evidence for the late Holocene indicates drier environmental conditions than today. In order to improve the paleoecological interpretation, we described the local vegetation and used moss samples as pollen traps at both montane bogs along strong soil moisture gradients.The Netherlands Centre for Geo-ecological Research, ICG     

Regional and local controls on postglacial vegetation and fire in the Siskiyou Mountains, northern California, USA

The Siskiyou Mountains of northwestern California and southwestern Oregon are a floristic hotspot, and the high diversity of conifers there likely results from a combination of geological, ecological, climatological and historical factors. To evaluate how past climate variability has influenced the composition, structure and fire regime of the Siskiyou forests, pollen, charcoal, and lithological evidence was examined from two lakes along a moisture gradient to reconstruct the vegetation, fire and climate history. The late-glacial period was characterized by subalpine parkland and infrequent fire at both sites. During the late-glacial/Early Holocene transition period, subalpine parkland was replaced by a closed forest of Pinus, Cupressaceae, Abies and Pseudotsuga and more frequent fires a 1000 years earlier at the wetter site, and it is likely that reduced Pacific Ocean upwelling created warmer drier conditions at the coast. In the Early Holocene, Pinus, Cupressaceae were less abundant and fire less frequent at the coastal site during a period of increased coastal upwelling and fog production. In the Late Holocene, Abies, Pseudotsuga, Pinus, and Quercus vaccinifolia increased in the forest at both sites suggesting a widespread response to cooling. Fewer fires at the wetter site may account for the abundance of Picea breweriana within the last 1000 years. The comparison of the two records implies that large-scale controls in climate during the last 14,000 cal yr BP have resulted in major changes in vegetation and fire regime. Asynchrony in the ecosystem response of wetter and drier sites arises from small-scale spatial variations in effective moisture and temperature resulting from topographically-influenced microclimates and coastal-to-inland climate gradients.     

Long-term human impact as registered in an upland pollen profile from the southern Black Forest, south-western Germany

A detailed,¹⁴C-dated, pollen profile from Steerenmoos, a raised bog in the uplands of the southern Black Forest (Schwarzwald) is presented. The Late-glacial and early Holocene vegetation history conforms to the known pattern of forest dynamics for that region. At ca. 6100 cal. B.P.,Abies replaced the mixed oak forest, which is in contrast to adjacent regions whereFagus spread beforeAbies. From the Subboreal onwards,Fagus expanded and slowly largely replacedAbies. The mire developed from a fen to a raised bog. The mountain pine (Pinus mugo ssp.rotundata) on the present-day bog surface is a result of medieval burning. Cereal pollen are first recorded in the Neolithic (7600 cal. B.P.) and there is a closed curve forPlantago Lanceolata — a good indicator of human impact — since the Bronze Age (4000 cal. B.P.). On the basis of the cereal pollen record nine human impact phases (HIP) are described. HIP 1 and 2, which are short, date to ca. 7600 and 6700 cal. B.P., respectively, in a mixed oak forest context and are characterized by declines inCorylus, Tilia, Ulmus and bySalix (but no major deforestation) and peaks in charcoal and loss-on-ignition curves. HIP 3 and 4, which are short and weak, date to ca. 6000 and 5300 cal. B.P., respectively, and occur in the context of anAbies alba forest. The Bronze Age and Iron Age HIPs 5-7 are more intense and of longer duration than the Neolithic phases and result in a decline inAbies and an increase inFagus. The early medieval HIP 8, although rather weak, probably finds expression also in an archaeological artefact, namely a dug-out boat from the near-by Schluchsee. Finally, the late Medieval HIP 9 resulted in a major transformation in the landscape. It is argued that the earlier HIPs are not a reflection of distant events in the lowland valleys of the Rhine, Danube or Neckar but reflect more or less local developments.     

7500 yr of mire-pool development and the history of Pinus sylvestris (L.) in Sub-Arctic coastal Norway

In coastal North Norway, mire areas and mire pools frequently exist, but their development and time of origin are poorly known. In order to investigate the development of a coastal mire pool and relate its changes to known climatic changes, a sediment sequence from the mire-pool Lillevardhaugvatnet (c. 0.05 ha large), was ¹⁴C-dated, investigated for loss on ignition and analysed for pollen and botanical macrofossils.The bottom of the sequence dates c. 7500 cal. BP. The site gradually developed from a swamp forest c. 6200 cal. BP via a more open dwarf-shrub phase to a pool c. 5000 cal. BP. The pool grew in depth as the result of continuous peat growth damming the water body. The water level of the pool was probably lowered by erosional drain c. 2700 cal. BP. Redeposited peat in the sequence occurred c. 2100 cal. BP and c. 1100 cal. BP. ¹⁴C-dates and pollen indicate that erosion and redeposition of the peat surrounding the mire pool is a normal process, connected with the expansion of the water body.The combination of macrofossils and pollen accumulation rates (PAR) of P. sylvestris indicates that in small lakes in coastal areas of North Norway, a PAR of 200–400 cm^− 2 yr^− 1 is sufficient for indicating local presence of P. sylvestris. P. sylvestris is represented with abundant macrofossils between 4800 and 2100 cal. BP. It is suggested that a marked P. sylvestris decrease about 2100 cal. BP may be a combined effect of human impact and climatic deterioration. A possible final termination of the P. sylvestris population about 1600 cal. BP may be considered human-made.     

Late Pleistocene and Holocene environments of the Southeastern United States from the stratigraphy and pollen content of a peat deposit on the Georgia Coastal Plain

A 5.4 m peat core from the Sandy Run Creek (SRC) valley in the upper Coastal Plain of Georgia shows that before 30 ka the valley had an aggradational sandy floor with shallow swales and low ridges of 1–2 m amplitude suggesting a braided stream pattern and a low water table. The climate from 30–25 ka was cool and dry and the vegetation open grassland with stands of pine and spruce that produced few fires. At ca. 16 ka a warmer, wetter climate caused SRC to meander and incise the valley fill removing some previously deposited sediment at the site of our peat core so that sediments of 13 ka rest directly on sediments deposited at 25 ka. After ca. 13 ka higher groundwater levels initiated vertical accretion of floodplains allowing peat accumulation in shallow depressions across the valley. Pollen from the Younger Dryas period (ca. 13–11 ka) indicates a cool, moist environment of open oak woodland, mesic trees, riparian populations of alder, and reduced levels of pine. By the early to mid Holocene, tupelo and oak dominated over pine indicating wetter conditions, while sand stringers in peat record periodic heavy rains, an elevated water table, and intervals of substantial runoff. Cooling after ca. 4.5 ka brought drier and more variable conditions. Fires increased and southern pine replaced tupelo and oak. The absence of sand stringers in peat younger than 4.5 ka indicates fewer intense rainfall events.     

Environmental changes off North Iceland during the deglaciation and the Holocene: foraminifera, diatoms and stable isotopes

A combined study of foraminifera, diatoms and stable isotopes in marine sediments off North Iceland records major changes in sea surface conditions since about 15 800 cal years (yr) BP. Results are presented from two gravity cores obtained at about 400 m water depth from two separate sedimentary basins on each side of the submarine Kolbeinsey Ridge. The chronology of the sedimentary record is based partly on AMS ¹⁴C dates, partly on the Vedde and the Saksunarvatn tephra markers, as well as the historical Hekla AD 1104 tephra. During the regional deglaciation, the planktonic foraminiferal assemblages are characterised by consistently high percentages of sinistrally coiled Neogloboquadrina pachyderma. However, major environmental variability is reflected by changes in stable isotope values and diatom assemblages. Low δ¹⁸O values indicate a strong freshwater peak as well as possible brine formation by sea-ice freezing during a pre-Bølling interval (Greenland Stadial 2), corresponding to the Heinrich 1 event. The foraminifera suggest a strong concurrent influence of relatively warm and saline Atlantic water, and both the foraminifera and the diatoms suggest mixing of cold and warm water masses. Similar but weaker environmental signals are observed during the Younger Dryas (Greenland Stadial 1) around the level of the Vedde Ash. Each freshwater peak is succeeded by an interval of severe cooling both at the beginning of the Bølling–Allerød Interstadial Complex (Greenland Interstadial 1) and during the Preboreal, presumably associated with the onset of intense deep water formatiom in the Nordic Seas. The Holocene thermal optimum, between 10 200 and about 7000 cal years (yr) BP, is interrupted by a marked cooling of the surface waters around 8200 cal yr BP. This cold event is clearly expressed by a pronounced increase in the percentages of sinistrally coiled N. pachyderma, corresponding to a temperature decrease of about 3°C. A general cooling in the area is indicated after 7000–6000 cal yr BP, both by the diatom data and by the planktonic foraminiferal data. After a severe cooling around 6000 cal yr BP, the planktonic foraminiferal assemblages suggest a warmer interval between 5500 and 4500 cal yr BP. Minor temperature fluctuations are reflected both in the foraminiferal and in the diatom data in the upper part of the record, but the time resolution of the present data is not high enough to pick up details in environmental changes through the late Holocene.     

History of Late Holocene vegetation at Quintana Roo,Caribbean coast of Mexico

A pollen record of a Late Holocene sediment core from the Mexican Caribbean coast (Quintana Roo) shows the development and changes of a mangrove system. Humid conditions seem to have persisted for the period approximately 2500–1500 ¹⁴C yr BP (pollen zone I), and mangrove Rhizophora mangle dominated with a good representation of elements from the nearby semi-evergreen tropical forest. During the period approximately 1500–1200 ¹⁴C yr BP (pollen zone II) the mangrove Conocarpus erecta dominated. R. mangle almost disappeared and other taxa appeared, suggesting drier climatic conditions and generally more open vegetation. This dry period coincided with the period of the Maya cultural decline. The following period (pollen zone III, approximately 1200–1000 ¹⁴C yr BP) was characterized by the recovery of R. mangle, indicating more humid conditions than in the preceding pollen zone. Pollen zone IV (approximately 1000 ¹⁴C yr BP till present) suggests a drier period reoccuring with C. erecta; this marks the transition to present day conditions.     

Holocene vegetation and fire dynamics in the supra-Mediterranean belt of the Gredos Range (central Iberian Peninsula)

Abstract

High-resolution pollen, macrofossil and charcoal data, combined with multivariate analysis, were used to reconstruct Holocene vegetation and fire dynamics at Manaderos. The studied mire is located in a highly valued location in the supra-Mediterranean belt of the Gredos Range (central Iberian Peninsula). The record started at 1260?calendar years (cal yrs) BP, according to eight radiocarbon dates, with the occurrence of an open Quercus pyrenaica forest and Pinus nigra stands. The palynological data show a dramatic change ～380?cal yrs BP, when the oak forest was progressively replaced by a maritime pine one. The interpretation of this record is related to an increase in agricultural and livestock activities and changes in the role of fire, thus Pyrenean oak canopy was consumed by fires providing openings for Pinus pinaster, better adapted to fire. The results are compared with other sequences from the Spanish Central System in order to better understand the past dynamics of the main forest constituents and to provide a critical view of the role of fire, anthropic dynamics and climatic events.     

Palaeoecological evidence for Holocene vegetation, climate and land-use change in the low Don basin and Kalmuk area, southern Russia

Two¹⁴C-dated pollen profiles from mires in the steppe belt of southern Russia are presented. On the basis of these and data from earlier investigations, the Holocene forest history of the southern part of Russia and Ukraine is reconstructed. The steppe belt is very sensitive to climatic oscillations and, in particular, to changes in evapotranspiration. The most favourable climate occurred between 6000 and 4500 B.P. (6800–5200 cal. B.P.), when forest attained its maximum extent in the steppe belt. The period 4500–3500 B.P. (5200–3800 cal. B.P.) was characterised by drier climate with the most arid phase occurring between 4200–3700 B.P. (4700–4000 cal. B.P.). During arid phases, the area under forest and also peat accumulation rates declined. Subsequently, a number of less pronounced climatic oscillations occurred such as in the period 3400/3300–2800 B.P. (3600/3500–2900 cal. B.P.) when there was a return to more humid conditions. During the last 2500 years, the vegetation cover of the steppe belt in southern Russia and Ukraine took on its present-day aspect. On the one hand, there is close correlation between the Holocene vegetation history of southern Russia and Ukraine and, on the other hand, the steppe belt of Kazakhstan and transgressions in the Caspian sea. Human impact on the natural vegetation became important from the Bronze Age onwards (after 4500 B.P.; 5200 cal. B.P.). Particular attention is given to the history of Scots pine (Pinus sylvestris), which had a much wider distribution in the southern part of eastern Europe in the early Holocene. The reduction in range during recent millennia has come about as a result of the combined effects of both climatic deterioration and increased human impact.     

Mangrove ecosystem changes during the Holocene at Spanish Lookout Cay, Belize

Mangroves are halophytic plants living at the land–sea interface and are therefore natural trackers of sea-level. Multiple proxies of a continuous (8 m) mangrove peat core (BT-79) from Spanish Lookout Cay, Belize illustrate mangrove ecosystem changes during the Holocene. Radiocarbon measurements show this site was colonized by mangroves ~ 8000 cal. yrs BP, with a significant decrease in the peat accumulation rate from ~ 6000 to 1000 cal. yrs BP. Stratigraphic characteristics of this peat core such as bulk density, magnetic susceptibility, and loss on ignition show relative uniformity, inferring an uninterrupted mangrove ecosystem during a majority of the Holocene. This is supported by pollen data from BT-79 that show that the site has been consistently dominated by Rhizophora mangle (red mangrove), with Avicennia germinans (black mangrove) and Laguncularia racemosa (white mangrove) present as well. Subfossil R. mangle leaves are used for stable nitrogen, carbon, and oxygen isotope (δ¹⁵N, δ¹³C, and δ¹⁸O) analyses. δ¹⁵N and δ¹³C values provide a proxy of this plant's past physiology and stand structure showing that dwarf (δ¹⁵N < − 3‰) and tall (δ¹³C < − 27‰) R. mangle stands were previously present at the site, which are a result of nutrient limitations that we equate with seawater inundation. δ¹⁸O values show differences in source water of R. mangle, with higher values attributed to the source water being composed of a greater proportion of seawater relative to precipitation. A decrease in inundation at the site is shown by lower δ¹⁸O values (< 19‰) from ~ 7000 to ~ 1000 cal. yrs BP that covary with the decreased sedimentation rate. Existing Caribbean sea-level data do not show evidence of a decrease in the rate of relative sea-level rise or fluctuations that we take to be the major causes of environmental changes at site BT-79.     

Lateglacial climate development in NW Romania — Comparative results from three quantitative pollen-based methods

This paper provides quantitative reconstructions of the Lateglacial changes in four climate parameters from two fine-resolution pollen profiles in the Gutaiului Mountains, NW Romania. Climate estimates are based on two modern analogue techniques (with and without considering vegetation types) and weighted averaging partial least squares regression (WA-PLS), giving evidence for several climatic fluctuations during the period from > 14,700 to 11,500 cal. yr BP. The comparative results of the two modern analogue techniques show consistent trends of climate changes that are also coherent at both sites, but these results appear to largely disagree compared with climate reconstruction provided by WA-PLS.The modern analogue techniques revealed four intervals with low temperatures: prior to 14,700 cal. yr BP, between 13,950 and 13,800; 13,400 and 13,200; and 12,700 and 11,700 cal. yr BP. The temperature declines were more pronounced for winter than for summer, suggesting an intensification of seasonality, which together with a drop in precipitation indicates an increase in continentality. The Younger Dryas is the most pronounced cooling phase with winter temperatures ~ 14–16 °C colder than modern conditions, annual and summer temperatures ~ 2–5 °C and ~ 2 °C, respectively below present ones. Precipitation was ~ 400–500 mm, half that of present. During the Bølling and Allerød, summer temperatures were close to modern values (13 to 17 °C), whereas winter (− 6 to − 12 °C) and annual temperatures (0.5 to 6 °C) as well as precipitation were (550 to 700 mm) lower, indicating more continental conditions compared to the present-day climate.     

Sea ice seasonality during the Holocene, Adélie Land, East Antarctica

Thin sections of laminated cores from different Antarctic coastal areas have demonstrated the potential of diatom species to document climate change at the seasonal scale. Here we present the relative abundances of four diatom species and species groups (Fragilariopsis curta group as a proxy for yearly sea ice cover, F. kerguelensis as a proxy for summer sea-surface temperature, Chaetoceros Hyalochaete resting spores as a proxy for spring sea ice melting and the Thalassiosira antarctica group as a proxy for autumn sea ice formation) in core MD03-2601 retrieved off Adélie Land on the Antarctic continental shelf. These abundances were compared to surface temperatures and sea ice cover modelled over the last 9000 years. Both the marine records and the simulated climate demonstrated a cooler Early Holocene (9000–7700 years BP), a warmer Mid-Holocene (7700–4000 years BP) and a colder Late Holocene (4000–1000 years BP). Yearly sea ice cover followed an inverse pattern to temperatures with less sea ice during the Mid-Holocene Hypsithermal than during the Late Holocene Neoglacial. However, diatom census counts and model output indicate that sea ice spring melting happened earlier in the season, as expected, but that autumn sea ice formation also occurred earlier in the season during the Hypsithermal than during the colder Neoglacial, thereby following seasonal changes in local insolation.     

Ectomycorrhizal responses to organic and inorganic nitrogen sources when associating with two host species

While it is established that increasing atmospheric inorganic nitrogen (N) deposition reduces ectomycorrhizal fungal biomass and shifts the relative abundances of fungal species, little is known about effects of organic N deposition. The effects of organic and inorganic N deposition on ectomycorrhizal fungi may differ because responses to inorganic N deposition may reflect C-limitation. To compare the effects of organic and inorganic N additions on ectomycorrhizal fungi, and to assess whether host species may influence the response of ectomycorrhizal fungi to N additions, we conducted an N addition experiment at a field site in the New Jersey pine barrens. Seedlings of two host species, Quercus velutina (black oak) and Pinus rigida (pitch pine), were planted at the base of randomly-selected mature pitch pine trees. Nitrogen was added as glutamic acid, ammonium, or nitrate at a rate equivalent to 227.5 kg ha⁻¹ y⁻¹ for eight weeks, to achieve a total application of 35 kg ha⁻¹ during the 10-week study period. Organic and inorganic N additions differed in their effects on total ectomycorrhizal root tip abundance across hosts, and these effects differed for individual morphotypes between oak and pine seedlings. Mycorrhizal root tip abundance across hosts was 90 % higher on seedlings receiving organic N compared to seedlings in the control treatment, while abundances were similar among seedlings receiving the inorganic N treatments and seedlings in the control. On oak, 33–83 % of the most-common morphotypes exhibited increased root tip abundances in response to the three forms of N, relative to the control. On pine, 33–66 % of the most-common morphotypes exhibited decreased root tip abundance in response to inorganic N, while responses to organic N were mixed. Plant chemistry and regression analyses suggested that, on oak seedlings, mycorrhizal colonization increased in response to N limitation. In contrast, pine root and shoot N and C contents did not vary in response to any form of N added, and mycorrhizal root tip abundance was not associated with seedling N or C status, indicating that pine received sufficient N. These results suggest that in situ organic and inorganic N additions differentially affect ectomycorrhizal root tip abundance and that ectomycorrhizal fungal responses to N addition may be mediated by host tree species.     

Autogenic succession, land-use change, and climatic influences on the Holocene development of a kettle-hole mire in Northern Poland

We reconstructed the Holocene developmental history of a kettle-hole peatland in the Tuchola Forest of Northern Poland, using pollen, testat amoebae and plant macrofossil indicators. Our aims were to determine the timing and pattern of autogenic succession and natural and anthropogenic influences on the peatland. Northern Poland is under mixed oceanic and continental climatic influences but has so far been less studied in a palaeoecological context than more oceanic regions of Europe. In the first terrestrial developmental phase of the mire, the testate amoebae-inferred depth to water table revealed two major dry shifts at ca. 9400 (end of lake phase) and ca. 7100 cal BP (a period of global cooling and dry shift in Western Europe). Conditions became wetter again in two steps at ca. 6700 and ca. 5800 BP after a dry event at ca. 6100 BP. The timing of the wet shift at 5800 BP corresponds to wet periods in Western Europe. Peat accumulation rates were low (0.1 mm yr^− 1) between ca. 5600 and ca. 3000 BP when sedges dominated the peatland. In the last 2500 yrs surface moisture fluctuated with wet events at ca. 2750–2400, and 2000 BP, and dry events at ca. 2250–2100 and 1450 BP. After 1450 BP a trend towards wetter conditions culminated at ca. 500 cal BP, possibly caused by local deforestation. Over the mire history, pH (inferred from testate amoebae) was mostly low (around 5) with two short-lived shifts to alkaline conditions (7.5) at ca. 6100 and 1450 BP indicating a minerotrophic influence from surface run-off into the mire. Up to about 1000 BP the ecological shifts inferred from the three proxies agree with palaeoclimatic records from Poland and Western Europe. After this date, however correlation is less clear suggesting an increasing local anthropogenic impact on the mire. This study confirms that kettle-hole peatlands can yield useful palaeoenvironmental data as well as recording land-use change and calls for more comparable studies in regions are the interface between major climate influences.     

Palynological evidence for Holocene environmental change in the Changjiang (Yangtze River) Delta, China

Holocene vegetation changes in response to climate fluctuations and human impacts are reviewed on the basis of pollen analyses from borehole cores taken from the Changjiang (Yangtze River) delta, China, and other previously reported data. During the earliest Holocene (10,930−9000 cal yr BP), the climate was warm and wet, allowing thermophilous hardwoods to occupy mid- and low-elevations surrounding a palaeo-Changjiang estuary. The climate became gradually cooler, and cool-tolerant conifers, grasses and ferns became dominant until 7600 cal yr BP, when the estuary became a delta. A mid-Holocene climatic optimum occurred between 7600 and 4800 cal yr BP, when evergreen and broadleaved deciduous trees flourished at mid- and low-elevations surrounding the delta front-prodelta. After this time, climate became cooler again until 1340 cal yr BP. During this period, evergreen and broadleaved deciduous trees were replaced by conifers and grasses inhabiting the inter/subtidal flat-delta front. This development of conifer-grassland vegetation was shortly interrupted between 3860 and 3200 cal yr BP when thermophilous tree cover increased and open vegetation with scattered conifers was reduced. Since 1340 cal yr BP, the vegetation has been similar to that at present under warm, wet conditions. Human impacts are recognized by the first appearance of Fagopyrum and a sudden increase in herb pollen at 4500 and 1340 cal yr BP, respectively.     

Late‐glacial and Holocene vegetation,climate and fire dynamics in the Serra dos Órgãos,Rio de Janeiro State,southeastern Brazil

We present a high‐resolution pollen and charcoal record of a 218 cm long sediment core from the Serra dos Órgãos, a subrange of the coastal Serra do Mar, located at 2130 m altitude in campos de altitude (high elevation grass‐ and shrubland) vegetation near Rio de Janeiro in southeastern Brazil to reconstruct past vegetation, climate and fire dynamics. Based on seven AMS ¹⁴C ages, the record represents at least the last 10 450 ¹⁴C yr bp (12 380 cal years bp ), The uppermost region was naturally covered by campos de altitude throughout the recorded period. Diverse montane Atlantic rain forest (ARF) occurred close to the studied peat bog at the end of the Late‐glacial period. There is evidence of small Araucaria angustifolia populations in the study area as late as the early Holocene, after which point the species apparently became locally extinct. Between 10 380 and 10 170 ¹⁴C yr bp (12 310–11 810 cal yr bp ), the extent of campos de altitude was markedly reduced as montane ARF shifted rapidly upward to higher elevations, reflecting a very wet and warm period (temperatures similar to or warmer than present day) at the end of the Younger Dryas (YD) chronozone. This is in opposition to the broadly documented YD cooling in the northern Hemisphere. Reduced cross‐equatorial heat transport and movement of the Intertropical Convergence Zone over northeastern Brazil may explain the YD warming. Markedly extended campos de altitude vegetation indicates dry climatic conditions until about 4910 ¹⁴C yr bp (5640 cal yr bp ). Later, wetter conditions are indicated by reduced high elevation grassland and the extension of ARF into higher elevation. Fire frequency was high during the early Holocene but decreased markedly after about 7020 ¹⁴C yr bp (7850 cal yr bp ).     

A new Late-glacial and Holocene record of vegetation and fire history from Lago del Greppo, northern Apennines, Italy

Detailed Late-glacial and Holocene palaeoenvironmental records from the northern Apennines with a robust chronology are still rare, though the region has been regarded as a main area of potential refugia of important trees such as Picea abies and Abies alba. We present a new high-resolution pollen and stomata record from Lago del Greppo (1,442 m a.s.l., Pistoia, northern Apennines) that has been dated relying on 12 terrestrial plant macrofossils. Late-glacial woodlands became established before 13000 cal b.p. and were dominated by Pinus and Betula, although more thermophilous taxa such as Quercus, Tilia and Ulmus were already present in the Greppo area, probably at lower altitudes. Abies and Picea expanded locally at the onset of the Holocene at ca. 11500 cal b.p. Fagus sylvatica was the last important tree to expand at ca. 6500 cal b.p., following the decline of Abies. Human impact was generally low throughout the Holocene, and the local woods remained rather closed until the most recent time, ca. a.d. 1700–1800. The vegetational history of Lago del Greppo appears consistent with that of previous investigations in the study region. Late-glacial and Holocene vegetation dynamics in the northern Apennines are very similar to those in the Insubrian southern Alps bordering Switzerland and Italy, across the Po Plain. Similarities between the two areas include the Late-glacial presence of Abies alba, its strong dominance during the Holocene across different vegetation belts from the lowlands to high elevations, as well as its final fire and human-triggered reduction during the mid Holocene. Our new data suggest that isolated and minor Picea abies populations survived the Late-glacial in the foothills of the northern Apennines and that at the onset of the Holocene they moved upwards, reaching the site of Lago del Greppo. Today stands of Picea abies occur only in two small areas in the highest part of the northern Apennines, and they have become extinct elsewhere. Given the forecast global warming, these relict Picea abies stands of the northern Apennines, which have a history of at least 13,000 years, appear severely endangered.     

The mid-Holocene extinction of silver fir (Abies alba) in the Southern Alps: a consequence of forest fires? Palaeobotanical records and forest simulations

Pollen records suggest that Abies alba played a dominating role in both the montane and lowland forests at the border of the Southern Alps between ca. 8500 and 5700 years ago. Two major declines in fir, at about 7300–7000 cal b.p. and at ca. 6000 cal b.p., followed by the local extinction of the species are characteristic of the area below ca. 1000 m a.s.l. In order to test the impact of fire on the population dynamics of silver fir, a dynamic model (DisCForm) with a fire module was applied to simulate the early- and mid-Holocene forest development. Simulation outputs based on different fire scenarios were compared with the pollen record from Lago di Annone (226 m a.s.l.). The marked Abies decreases shown in the pollen record can be simulated with very intensive fire scenarios, but they do not result in an extinction of silver fir in the model. Low charcoal influx values related to the Abies declines in the palaeobotanical record suggest that fire was not the only reason for the extinction of silver fir. Human impact, as well as Holocene climatic changes leading to temporary moisture deficits and reduced adaptability due to low genetic variation may have had a significant impact on the Abies forests.     

Pollen-based quantitative reconstructions of Holocene climate variability in NW Romania

A modern analogue technique is applied to two high-resolution pollen sequences from NW Romania to provide the first quantitative evidence for winter, summer and annual temperatures and for precipitation across the Holocene in this region.

The pollen-based climate reconstructions allow the identification of four main intervals: i) an early, less stable period between 11,700 and 11,200 cal. yr BP; (ii) generally stable conditions between 11,200 and 8300 cal. yr BP with winter and annual temperatures and precipitation higher than at present, and summer temperatures about the same; (iii) lower winter and annual temperatures, and higher summer temperatures and precipitation between 8000 and 2400 cal. yr BP; (iv) warmer winter and annual temperatures and lower precipitation for the last 2400 years, whereas summer temperatures became cooler at Steregoiu and remained stable at Preluca Tiganului.

The pollen-based climate reconstructions at the two sites show similar patterns in annual and winter temperatures and precipitation changes during the Holocene, but the trends appear to be less consistent for summer temperatures.

Our pollen-based reconstructions revealed several short-term climatic oscillations during the Holocene, the strongest of which occur between 10,300–10,100, 8300–8000, 6800–6400, 5100–4900, 4000–3600 and 3200–3000 cal. yr BP.