Cyst formation,sedimentation, and preservation: Factors affecting dinoflagellate assemblages in recent sediments from trondheimsfjord,Norway

Plankton records and 25 samples of Recent sediment from Trondheimsfjord and the adjoining shelf were studied to investigate production, sedimentation, and preservation of cysts, as factors which influence the eventual composition of dinoflagellate cyst assembleges. All sediment samples were examined for dinoflagellate cysts using routine semiquantitative palynological procedures. In addition, fjord sediments were subjected to a limited sediment analysis, and, for three samples, results from preparations both with and without acid treatments were compared. For the first time, cyst assembleges from Recent sediments were directly compared with extensive plankton records from overlying waters. Results indicate that approximately 20% of the 55 locally recorded dinoflagellate species contribute cysts to bottom sediments. Once formed, cysts behave as fine silt particles in the sedimentary regime, increasing in abundance as the percentage abundance of finer sediment increases, usually with increased water depth. Cyst-forming species are almost entirely restricted to a few genera, particularly Gonyaulax and Peridinium, within the order Peridiniales. For some groups, reasonably good correspondence was found between percentage abundances of dinoflagellates in plankton and their cysts in sediment, though plankton records covering at least five years were required to establish this. Gonyaulax grindleyi Reinecke (Von Stosch 1969) appeared to be consistently overrepresented by cysts in sediment relative to available plankton evidence; possible explanations are suggested. At least 30% of the cyst species present, including most Peridinium species, were eliminated, or rendered unreliable for semiquantitative palynology, by application of routine palynological preparation treatments. Such cysts may provide useful, non-quantitative, palynological information from Recent and possibly Quaternary sediments, but their persistence would seem unlikely. Thus, factors of preservation probably further restrict the dinoflagellate fossil record. Cyst assemblages from Trondheimsfjord are comparable with those previously recorded from the northeastern coast of U.S.A., and from Scotland and northeastern England. Fjord assemblages are dominated by small, simple, spinose cysts which would be regarded as acritarchs if culture experiments had not proved that they are dinoflagellate cysts. Much potential biogeographic and palaeoenvironmental information was included within the less abundant species.Attention is drawn to the role which future culture experiments may be expected to play in helping to resolve taxonomic difficulties currently affecting dinoflagellate studies. Palynological significance of results from the present study is discussed especially with reference to recent work by Von Stosch which strongly suggests that cysts may be hypnozygotes formed routinely in sexual cycles of dinoflagellates.     

Non-pollen palynomorphs in the Black Sea corridor

There have been few studies of non-pollen palynomorphs (NPP) in Holocene brackish water environments. The Black Sea is one of the world’s largest and deepest bodies of stable brackish water and a natural laboratory for study of marine carbon cycling to anoxic sediments. The main NPP in the modern sediments of this brackish water sea are dinoflagellate cysts (dinocysts), acritarchs (mainly the prasinophytes Cymatiosphaera, Micrhystridium, Sigmopollis and Pseudoschizaea) and diverse fungal remains. Other NPP include colonial algae, tintinnids, copepod and cladoceran egg covers, testate amoebae and microforaminiferal linings. These NPP assemblages are similar to those in the marginal marine environment of the Pliocene St. Erth Beds (England), but have more abundant NPP, and virtually lack scolecodonts. In the Black Sea corridor, modern assemblages from areas with salinity >22‰ have higher percentages of microforaminiferal linings and fewer prasinophytes, colonial algae and fungal spores. Prasinophytes dominate only in mid-Holocene sediments, during a 2000 years interval of sea level transgression and sapropel deposition. Early Holocene sediments have lower dinocyst diversity, increased fresh–brackish water colonial algae (Pediastrum spp. and Botryococcus braunii), zygnemataceous spores and desmids (including Zygnema, Cosmarium), ostracod linings and fewer foraminiferal linings. These assemblages are similar to those in the Baltic Sea where the annual salinity is about 6–8‰.     

Le passage Cénomanien–Turonien dans le Nord-Ouest algérien (coupe de Douar Menkouchi,Ouarsenis) : biostratigraphie des radiolaires

The Ouarsenis Massif belongs to the Algerian Tell domain which is considered as the eastern part of the Maghrebian Tethys former margin. The Cenomanian-Turonian Douar Menkouchi section outcropping in the eastern side of the Ouarsenis massif is composed of marls and laminated limestone alternations, including massive limestone and a biosiliceous bed toward the upper part of the succession. It reveals the presence of well-preserved and diversified radiolarians. In this study, we have identified for the first time in the Algerian domain, age-diagnostic radiolarian assemblages characterizing the Cenomanian-Turonian transition (C/T). The organic-rich laminated beds (black shales) confined to Silvae Zone, Cenomanian in age and underlying a biosiliceous interval, have provided radiolarian assemblages dominated by Pseudodictyomitrae. The successions overlying the biosiliceous interval have provided diversified radiolarian assemblages including Hagiastridae (i,e. Crucella cachensis, Alievium cf. superbum) and abundant Dictymitrae taxa (i,e. Dictyomitra cf. napaensis and Dictyomitra undata) indicating early Turonian age. These marl-limestone alternations are assigned to the Alievium superbum Zone. Considering taphonomic and diagenetic implications on assemblages content, we conclude that organic-rich beds and biosiliceous interval outline the latest Cenomanian. The radiolarian assemblage composition exhibits similarities to microfauna recovered from Bonarelli Horizon in Central Italy. Morphological similarities are also deduced from assemblages containing Pseudodictyomitrae tiara morphotypes (synonym of P. nakasekoi) which resemble those recovered from ophiolitic intervals in Cyprus and Oman areas.     

Dinoflagellate cysts,Acritarchs and Tasmanitids from the uppermost Callovian of England and Scotland: With a reconsideration of the “xanthidium pilosum” problem

Four assemblages of fossil organic-walled microphytoplankton from the Upper Callovian (Lamberti Zone), three from England and one from Scotland, are described. They comprise at least 50 species of dinoflagellate cysts, 18 species of acritarchs and 2 species of tasmanitids. After a review of the «Xanthidium pilosum problem, a revised diagnosis is proposed for Sentusidinium pilosum (Ehrenberg) and three new species, Sentusidinium creberbarbatum, S. sparsibarbatum and S. erythrocomum are proposed. The new family name Scriniodiniaceae is substituted for the now invalid family name Endoscriniaceae. The new genus Escharisphaeridia is proposed to contain two existing species, E. pococki (Sarjeant) and E. dictydia (Sarjeant). The present content of the genus Compositosphaeridium is examined; the new combination C. polonicum (Górka) is proposed and a new species, C. bulgaricum, is erected. One further new species of dinoflagellate cyst, Prolixosphaeridium anasillum, is proposed. The new term «peniplate is proposed for paraplates whose boundaries are penitabular in situation.One new species of acritarch, Solisphaeridium?rhachos, is described and the new combination Polygonium aster (Sarjeant) is proposed. Evidence for reworking of some of the species of Micrhystridium present in the Hackness Rock assemblage is noted, though it is considered that the majority of species in that assemblage are indigenous. It is considered that all four assemblages are derived from a single phytoplankton province, the differences between them resulting from environmental factors such as water depth or distance from a shoreline.     

Callovian (Middle Jurassic) marine microplankton from southwestern Germany: Biostratigraphy and paleoenvironmental interpretations

Late Middle Jurassic dinoflagellate cyst assemblages are documented from a section in Kandern and in samples from additional localities in southern Germany. The changes in the composition of the marine microfloras through the Callovian sections are related to fluctuations in sea-levels and changes in depositional environments. The Lower Callovian claystones, which represent inner shelf, soft bottom deposits, contain rich, diverse and well preserved dinoflagellate cyst assemblages. In the Kandern section a marked transition from the Lower Callovian claystone facies to the Anceps-oolite occur within the Calloviense zone. This transition was caused by a regression, and coincides with the LZA−3.1/LAZ−3.2 boundary on the cycle chart of Haq et al. (1987). Diverse dinoflagellate cyst assemblages also occur in the Anceps-oolite, but they are not so abundant as the acritarchs which dominate the overall palynofloras. The Anceps-oolite is interpreted as representing an offshore, but shallow marine environment. The transition to the overlying Upper Callovian “Renggeri Clay” is more gradual, but accompanied by marked changes in the overall composition of the marine microfloras. The “Renggeri Clay”, which represent an outer shelf, soft bottom depositional environment, shows a higher species diversity and abundance of dinoflagellate cysts than the underlying sequences. The distribution pattern of selected species through the Kandern section indicates that some species were related to specific environments, i.e. open marine forms, and that others could have been opportunistic forms, tolerating more shallow marine environments.

The distribution of the dinoflagellate cysts from southern Germany is related to the standard Northwest European ammonite zonation. The ranges of selected biostratigraphic key species are compared with ranges previously reported from the British and northwest European Jurassic. There are several discrepancies with respect to the earliest appearance and extinction datums, the only relevant Callovian palynostratigraphic “events” coinciding in southern Germany and Britain being: The extinction of Aldorfia aldorfensis at the top of the Calloviense zone, and the earliest incoming of Belodinium spp. and Liesbergia scarburghensis at the base of the Lamberti zone.     

Sequence biostratigraphy and paleoenvironmental reconstruction in the Early Eocene Figols Group of the Tremp-Graus Basin (south-central Pyrenees, Spain)

The dinoflagellate cyst, palynofacies and foraminiferal records from the Early Eocene Figols Group of the Spanish Pyrenees exhibit major changes associated with relative sea-level fluctuations and variations in efficiency of the fluvial systems. The mixed siliciclastic-carbonate marine succession exposed in the Merli-Esdolomada area was deposited on the shelf in a tectonically active basin, under oxic conditions, in shallow to moderately deep marine waters (close to the lower boundary of the photic zone) influenced by river discharge from adjacent emerged lands. Sedimentary cycles are mainly recorded by fine-grained deposits. The paleoenvironmental interpretation of palynological and calcareous microfaunal records allows the definition of depositional sequences and enables to trace basinward surfaces identified in more proximal settings by means of sedimentological facies analysis. At the same time, the solid regional stratigraphic framework already available for the Figols Group allows new insights on the paleoecology of extinct taxa.The Merli-Esdolomada section (ME) spans two 3rd order sequences. The upper sequence reflects more marginal marine conditions than the lower one, thus pointing to an overall shallowing trend with time. In the lower sequence the maximum flooding interval is characterised by a rich and diverse microfauna, highest relative abundances of marine phytoplankton and typically neritic dinoflagellate cyst (dinocyst) assemblages dominated by Spiniferites and Cordosphaeridium. The final phase of the highstand systems tract, in proximity of the main sequence boundary, is characterised by a decrease in abundance and diversity of dinocysts, with the dominance of the lagoonal genus Polysphaeridium, and by decreased microfaunal diversity (discorbids, miliolids, larger foraminifers).Palynological records permit the reconstruction of activation and deactivation phases of the fluvial systems, variations in runoff from the hinterland, nutrient delivery to the sea and productivity in surface waters of the Tremp-Graus Basin. The overall abundance of Spiniferites and the scarcity of peridinioid cysts, Lingulodinium, Pediastrum and Botryococcus algae are interpreted to indicate oligotrophic conditions during the deposition of the investigated succession. Recurrent intervals where the temporary disappearance of Spiniferites corresponds to highest relative abundances of Operculodinium and herbaceous debris are the main evidence for river plumes and denote periods of enhanced fluvial discharge reflecting a Milankovitch-type cyclicity. The Rotalia group, which shows abundance peaks of Cuvillierina spp. in the shallower intervals, is often associated with these episodes, thus suggesting for this taxon more tolerance to turbidity than larger foraminifera. Within the maximum flooding zone, a Thalassiphora patula acme corresponding to the local disappearance of Homotryblium is deemed to record the distal expression of a fluvial activation which triggered water salinity stratification. Despite broad morphological similarities, Homotryblium exhibits more cosmopolitan preferences than Polysphaeridium, which is instead confirmed as a lagoonal euryhaline taxon. The increase of Spinizonocolpites pollen toward the top of the Figols Group records the northwestward migration of Nypa mangrove-palms from southern Europe during the mid Early Eocene.     

Microbial carbonates and corals on the marginal French Jura platform (Late Oxfordian,Molinges section)

Molinges was located on an Upper Jurassic ramp system of low-energy regime that developed at the southern margin of the French Jura platform. The sedimentary succession is characterized by the transition from a mixed siliciclastic-carbonate to a carbonate depositional setting that occurred during a long-term shallowing-upward trend. The disappearance of siliciclastics is explained by a climatic change, from humid and cold to drier and warmer conditions, previously identified in Late Oxfordian adjacent basins. The base of the section shows marl-limestone alternations of outer ramp. In its middle part, the section displays oncolitic marls, coral-microbialite beds and oncolitic limestones that deposited in a mid ramp position. Finally, the upper section part is made of oolitic limestones of inner ramp. In outer- to mid-ramp settings submitted to terrigenous inputs, the stacking pattern of deposits and facies evolution allow the identification of elementary, small-, medium-, and large-scale sequences. Small amplitudes of sea-level variations probably controlled rapid shifts of facies belts and reef window occurrences. In small-scale sequences, the coral beds developed during periods of sea-level rise. The decreasing rate of sea-level rise is marked by the downramp shift of the oncolitic limestone belt that led to the demise of coral-microbialite beds. These bioconstructions are mainly represented by thin biostromes in which corals never reach great sizes. The coral assemblages mainly include the genera Enallhelia, Dimorpharaea, Thamnasteria, and some solitary forms (Montlivaltia and Epistreptophyllum). They suggest relatively low-mesotrophic conditions in marine waters during the edification of the primary framework. Relatively cold water temperatures and periods of more elevated nutrient contents are probably responsible of the reduced coral development and the formation of a large amount of microbialites.     

Paleoenvironmental and paleoclimatic changes during the Late Cretaceous and Cretaceous–Paleogene (K/Pg) boundary transition in Tattofte,External Rif,northwestern Morocco: implications from dinoflagellate cysts and palynofacies

This paper presents a quantitative study of dinoflagellate cysts (dinocysts) and palynofacies of the Campanian–Danian marly succession at the village of Tattofte (western External Rif, northwestern Morocco). The paleoenvironmental and paleoclimatic interpretations, inferred from this palynologic analysis, are compared to coeval sequences of other areas in the Northern Hemisphere. Changes in the relative abundances of dinocyst taxa, which are paleoenvironmental indicators, throughout the section suggest a deposition in a marine inner to outer neritic setting. The upper Campanian dinocyst assemblage is characterized by the presence of outer neritic-oceanic and low productivity indicator taxa (e.g., Spiniferites spp., Odontochitina spp.) and cold-water taxa (e.g., Manumiella spp., Chantangiella spp., Laciniadinium spp.), whereas, the lower Maastrichtian assemblage is characterized by inner neritic, high productivity and warm-water indicator taxa (e.g., Palaeocystodinium spp., Andalusiella spp.). The upper Maastrichtian dinocyst assemblage displays a return to an outer neritic environment under a transgressive regime, but with a cooling pulse and a moderate productivity. However, the Cretaceous–Paleogene (K/Pg) boundary interval records remarkable changes in the relative abundances of dinocyst taxa, indicating an inner neritic (coastal) setting, which is the most proximal in the study section; such changes reflect instable paleoenvironmental conditions which may be related to global cooling periods, likely caused by the Deccan volcanism in India and/or the Chicxulub asteroid impact in Mexico at the K/Pg boundary. In the Danian, the dinocyst relative abundances indicate a gradual return to stable environmental conditions.A quantitative analysis of the kerogen plots (palynomorphs, phytoclasts and amorphous organic matter (AOM)) reveals five types of palynofacies, generally indicating oxic to suboxic marine environments. The upper Campanian and upper Maastrichtian (lowermost part) strata are characterized by a playnofacies (V), indicating a distal shelf, while the lower Maastrichtian and lower Danian (uppermost part) strata record a playnofacies (III), reflecting an intermediate inner-outer neritic environment. However, the K/Pg boundary transition is characterized by playnofacies types (I) and (II), indicating a proximal (coastal) environment.     

Last interglacial surface water conditions in the eastern Nordic Seas inferred from dinocyst and foraminiferal assemblages

Marine sediments from the Vøring Plateau (Norwegian Sea) have been studied for their dinoflagellate cyst (dinocyst) and foraminiferal content in order to reconstruct sea-surface conditions in the eastern Norwegian Sea during Marine Isotope Stage (MIS) 5e. In combination with stable oxygen isotope and ice rafted detritus (IRD) data, the variations in foraminiferal and dinocyst assemblage composition reflect a stepwise transition from the final phase of deglaciation (Termination II) into typical interglacial conditions. This stepwise change is repeated subsequently during the cooling conditions of glacial inception towards MIS 5d. The interval studied is characterized by relatively high abundances of Bitectatodinium tepikiense, in comparison to present-day values in the area, indicating a larger seasonal temperature amplitude with enhanced surface water stratification during MIS 5e. The important occurrence of the warm-temperate dinocyst Spiniferites mirabilis s.l. concurrent with subpolar foraminifers Turborotalita quinqueloba, Globigerina bulloides, and Globigerinita glutinata reveals that most pronounced interglacial marine conditions prevailed in the area just prior to the transition towards MIS 5d. The late stratigraphic position of this phase in the interglacial is verified by comparison with dinocyst data from south of Iceland, manifesting its over-regional implication. Besides the good agreement in dinocyst and foraminiferal assemblage changes, the variations in and between both fossil assemblages also point to the existence of some significant surface water variability in the eastern Norwegian Sea during MIS 5e.     

Palynology of the Toarcian-Aalenian Global Boundary Stratotype Section and Point (GSSP) at Fuentelsaz (Lower-Middle Jurassic, Iberian Range, Spain)

The Fuentelsaz Section is located in the Castilian Branch of the Iberian Range (Guadalajara, Spain). Its exceptional Lower-Middle Jurassic transition outcrops led to its designation as the Global Boundary Stratotype Section and Point for the base of the Aalenian. The sediments of the Toarcian-Aalenian transition at Fuentelsaz are composed of marls with interbedded limestones in rhythmic alternation. Marls are dominant in the Toarcian and the Opalinum Zone up to the middle part of the Comptum Subzone. In the remaining Comptum Subzone, marly materials are scarce and limestones predominate. Sedimentation at Fuentelsaz took place over an extensive marine epeiric carbonate platform that was well connected with the open sea. In the Fuentelsaz Section, a total of 43 palynomorph taxa were recorded: 23 spore taxa, 13 pollen taxa, 4 acritarchs, 2 prasinophytes and 1 dinoflagellate cyst. The studied sediment samples were always dominated by terrestrial allochthonous miospores. In general, miospore assemblages are biased due to the transport of pollen from land or islands to the continental platform. Spheripollenites, Classopollis or indeed both pollens numerically dominate the assemblages. Other miospores appear in low numbers. Aquatic palynomorphs are also scarce; Micrhystridium lymensis is the most common. Five palynological assemblages (PA) were distinguished: PA1, PA2 and PA3 are Toarcian in age, PA4 is located at the boundary between the Toarcian and the Aalenian and PA5. This latter palynological assemblage which is lower Aalenian in age shows a strong reduction in palynomorph diversity and preservation in the Comptum Subzone. Palaeoecological analysis of the palynomorphs indicated the presence of calm, oligotrophic sea water. The palaeofloral communities of gymnosperms and vascular cryptogams—which grew in subtropical arid conditions—were poorly diversified.     

Dinoflagellate cyst distribution in marine surface sediments off West Africa (17–6°N) in relation to sea-surface conditions, freshwater input and seasonal coastal upwelling

An organic-walled dinoflagellate cyst analysis was carried out on 53 surface sediment samples from West Africa (17–6°N) to obtain insight in the relationship between their spatial distribution and hydrological conditions in the upper water column as well as marine productivity in the study area.Multivariate analysis of the dinoflagellate cyst relative abundances and environmental parameters of the water column shows that sea-surface temperature, salinity, marine productivity and bottom water oxygen are the factors that relate significantly to the distribution patterns of individual species in the region.The composition of cyst assemblages and dinoflagellate cyst concentrations allows the identification of four hydrographic regimes; 1) the northern regime between 17 and 14°N characterized by high productivity associated with seasonal coastal upwelling, 2) the southern regime between 12 and 6°N associated with high-nutrient waters influenced by river discharge 3) the intermediate regime between 14 and 12°N influenced mainly by seasonal coastal upwelling additionally associated with fluvial input of terrestrial nutrients and 4) the offshore regime characterized by low chlorophyll-a concentrations in upper waters and high bottom water oxygen concentrations.Our data show that cysts of Polykrikos kofoidii, Selenopemphix quanta, Dubridinium spp., Echinidinium species, cysts of Protoperidinium monospinum and Spiniferites pachydermus are the best proxies to reconstruct the boundary between the NE trade winds and the monsoon winds in the subtropical eastern Atlantic Ocean. The association of Bitectatodinium spongium, Lejeunecysta oliva, Quinquecuspis concreta, Selenopemphix nephroides, Trinovantedinium applanatum can be used to reconstruct past river outflow variations within this region.     

Ediacaran acanthomorphic acritarchs from the Outer Krol Belt,Lesser Himalaya,India: Their significance for global correlation

Large acanthomorphic acritarchs have been previously reported from the Ediacaran successions of the Pachmunda and Krol Hill synclines in the Outer Krol Belt of Lesser Himalaya, India. Thin sections of chert from Krol ‘A’ Formation in Khanog and Rajgarh synclines, Outer Krol Belt, record an equally well developed and diversified assemblage of Ediacaran large acanthomorphic acritarchs. This assemblage contains specimens belonging to seven genera and ten species, identified as: Appendisphaera fragilis, A. grandis, Asterocapsoides sp. A, Asterocapsoides sp. B, Cavaspina acuminata, C. basiconica, Eotylotopalla dactylos, Knollisphaeridium sp., Papillomembrana sp., and Weissiella cf. grandistella. It also contains five unnamed forms, viz. A, B, C, D, and E. The Krol acritarch assemblage shows a close resemblance with the Upper Doushantuo or Tanarium anozos–Tanarium conoideum assemblage of China. However, the absence of biostratigraphically important markers such as Tanarium anozos and T. conoideum from the Krol assemblage, so far, makes it difficult to establish a definite biostratigraphic correlation between the two assemblages. The current observations from this new locality provide additional data for regional and global biostratigraphic correlation, and significantly increase the purview of Ediacaran sequences for global biostratigraphic zonation.     

Control of modern dinoflagellate cyst distribution in the Irish and Celtic seas by seasonal stratification dynamics

Surface sediments from seven stations located in the seasonally stratified, frontal and mixed water regions in the Celtic and Irish seas have been analysed for their dinoflagellate cyst assemblages and dinosterol content. A total of 45 dinoflagellate cyst taxa have been identified and the assemblages related to surface and sediment conditions. Sediments from the mixed water region, at 30 m water depth, are characterised by a relatively low cyst concentration (∼2300 cysts/g dry weight) and high relative abundances of Lingulodinium machaerophorum accompanied by Spiniferites membranaceus, Brigantedinium spp. and Dubridinium caperatum. Assemblages from stratified and frontal water stations are dominated by Spiniferites ramosus associated with Operculodinium centrocarpum, Brigantedinium spp., cysts of Polykrikos schwartzii and Selenopemphix quanta. Ordination techniques performed on a restricted number of 35 taxa from the assemblages differentiated the stratified and frontal assemblages based on the abundance of the less abundant species Bitectatodinium tepikiense and Spiniferites elongatus. Among the environmental parameters (sea-surface temperature and salinity, stratification index, chlorophyll concentration and sediment grain-size classes), the seasonal stratification and sedimentological context, itself a function of tidal dynamics, explain most of the variance in the environmental conditions. These results indicate that dinoflagellate cyst analyses of shelf sediment records can be used to document the planktonic signal of seasonal stratification dynamics.     

Modem organic-walled dinoflagellate cysts in arctic marine environments and their (paleo-) environmental significance

The Arctic Ocean is one of the least known marine regions of the world. Because of its major influence on global climate and its hostile environmental conditions it is a fascinating area for paleoecological, paleoclimatic and paleoceanographic research. The composition of planktic microfossil assemblages, and both the trace-element and stable isotope compositions of hard parts, provide us with valuable information about the physical and biochemical parameters of surface waters in the high northern latitudes. Calcareous and biosiliceous microfossils that are traditionally used in Quaternary paleoenvironmental studies are of limited value in the Arctic Ocean because of their low abundances, low diversity and/or low preservation potential. The past several decades have seen considerable progress in our knowledge of the ecology and biogeography of dinoflagellates and their organic-walled cysts in the high northern latitudes, and these dinoflagellate cysts are now important proxies for reconstructing surface water conditions in the Quaternary. This arcticle gives an overview of the ecology of dinoflagellates and their cysts, the processes that transform the living communities into sediment communities, and the environmental gradients that may be reconstructed from fossil dinoflagellate cysts assemblages in the high northern latitudes.      

First record of organic-walled microfossils from the Tonian Shiwangzhuang Formation of the Tumen Group in western Shandong,North China

Organic-walled microfossils offer important information on the biospheric evolution in pre-Cryogenian and provide biostratigraphic implications for many Proterozoic fossiliferous sequences that are poorly age constrained for the lack of reliable radiometric date. Recently, macroscopic carbonaceous compression fossils have been reported for the first time from the Tonian Shiwangzhuang Formation of the Tumen Group in western Shandong, North China. However, organic-walled microfossils have never been discovered from this formation up till now. To improve our knowledge about Proterozoic biodiversity in North China, we conducted a micropaleontological survey on the argillaceous limestone samples of the Shiwangzhuang Formation, which also contain macroscopic carbonaceous compression fossils, from the Baishicun section in Anqiu, western Shandong, North China. Our investigation shows that the Shiwangzhuang microfossil assemblage is dominated by smooth-walled sphaeromorphic acritarchs and cyanobacterium-like filaments and relatively low abundance of other acritarchs, including 16 taxa, such as Polysphaeroides filliformis, Ostiana microcystis, Simia annulare, ?Jacutianema sp., Arctacellularia tetragonala, Pellicularia tenera, Polythrichoides lineatus, and Navifusa actinomorpha. The Shiwangzhuang organic-walled microfossil assemblage, although consisting of long-ranging and not age diagnostic taxa, is consistent with a Tonian age suggested by macroscopic carbonaceous compression fossils, including the Chuaria-Tawuia and Sinosabellidites-Protoarenicola-Pararenicola assemblages, revealed from the same fossiliferous horizon of the Shiwangzhuang Formation and by organic-walled microfossil assemblage, including the late Mesoproterozoic to Tonian index fossil Trachyhystrichosphaera aimika, from the underlying Tongjiazhuang Formation. However, it is also worth noting that a Cryogenian or Ediacaran age cannot be completely excluded based just on the Shiwangzhuang microfossils because of their limited biostratigraphic utility.     

Impact of grazing and bioturbation of marine benthic deposit feeders on dinoflagellate cysts

The impact of benthic deposit feeders on marine dinoflagellate cysts was studied by adding a concentrated natural Swedish cyst assemblage to sediment with different deposit feeders in replicate 4-l aquaria. The deposit feeders used were the bivalve Abra nitida, the echinoderm Amphiura filiformis, and the polychaetes Melinna cristata and Nereis diversicolor. These species occur naturally near the Swedish west coast and were selected to represent different ways of feeding. The results showed a significant relative decrease of unfossilizable cyst species; whereas, the common fossilizable species Lingulodinium polyedrum significantly increased in the cyst assemblage after grazing. This work suggests that differences in dinoflagellate cyst compositions can in part be caused by different animal grazing behaviors.     

Late Devonian Acritarchs from the Gneudna formation in the Western Carnarvon basin,Western Australia

The Gneudna Formation is a Late Devonian(Frasnian) sequence of marine calcareous sediments that occurs in the Carnarvon Basin, Western Australia. The present palynological study is based upon subsurface silty strata from a borehole (Pelican Hill or Bibbawarra Bore) that was drilled early this century near the western coastal limit of the Carnarvon Basin.The subject strata have previously been attributed to the Gneudna Formation on lithostratigraphic grounds. They contain a rich and varied assemblage of marine microphytoplankton (acritarchs), associated with trilete miospores of which Geminospora lemurataBalme, 1962 is the dominant form. Forty-seven species of acritarchs are recognizable in the palynoflora, which corresponds very closely with that described recently (Playford & Dring, 1981) from the Gneudna Formation in the vicinity of its type section on the opposite (eastern) side of the Carnarvon Basin. The apparently parochial complexion of the Gneudna acritarch suite is probably illusory, insofar as early Late Devonian acritarchs have not been studied extensively or intensively from either the northern or southern hemispheres.The following new species of acritarchs areformally instituted herein: Elektoriskos villosa, Lophosphaeridium pelicanensis, and Pterospermella tenellula.     

南海北部大陆架琼东南盆地西南部新近纪沟鞭藻类

首次报道的低纬度晚第三纪(新近纪)丰富多样的海相沟鞭藻类,产自南海北部大陆架琼东南盆地西南部的两钻孔(主要是Ya2112井,少量为Ya2111井),经鉴定达40余种。文中仅对其中的31种4亚种进行了照相和对15种2亚种和1新种(Impletosphaeridiumcapillaresp.nov.)在我国进行了首次描述;在新近系建立2个沟鞭藻组合:即Hystrichosphaeropsisobscura组合(下部)和ReticulatosphaeraactinocoronataPolysphaeridiumzoharyi组合(上部),并讨论了它们的组合特征及其地质时代,认为其下部的组合时代为中新世,上部的组合时代为上新世。     

Palynological evidence of Early Carboniferous sedimentation in the Llanos Orientales Basin, Colombia

Early Carboniferous palynological assemblages from the SM-4 well located in the Llanos Orientales Basin constitute the only definitively geological evidence of Carboniferous strata in this Basin. Stratigraphically significant species include: Anapiculatisporites concinnus, Apiculiretusispora mutiseta, Grandispora spiculifera, Indotriradites dolianitii morphon, Spelaeotriletes pretiosus and Prolycospora rugulosa. The presence of scarce acritarchs indicates a shallow marine environment. A Tournaisian age, probably reaching the Viséan, is proposed for the interval 2010-2340 ft based on the presence of distinctive spore species with previous records in the Viséan and Tournaisian of Western Europe and Western Gondwana. This report increases the knowledge about the distribution and composition of the Early Carboniferous palynological assemblages in Northern Gondwana.