Über zwei diploide,in Südwest-Anatolien endemische Arten aus dem Komplex derVeronica cymbalaria (Scrophulariaceae)

Veronica lycica Lehm. is a distinct, diploid member of theVeronica cymbalaria group, endemic in Lycia (S.W. Anatolia). Closely related is the newV. stamatiadae M. Fischer etW. Greuter which is also diploid and seems to be restricted to the small Greek island Ro close to the South coast of Lycia. The chromosome numbers for both species are reported for the first time (2n = 18).

Untersuchungen über den PolyploidkomplexVeronica cymbalaria agg., II. — Der erste Beitrag dieser Serie:Fischer (1975).     

Pollen morphology ofSonchus and related genera,and a general discussion

The pollen morphology of the taxa belonging to the generaAetheorhiza Cass.,Launaea Cass.,Reichardia Roth andSonchus L. in the Iberian Peninsula has been studied with light and electron microscopy. The pollen is 3(-4)-zonocolporate and echinolophate (without polar lacunae, but in general with prelacunae), with equatorial ridges and 15–20 lacunae: 3–4 poral, 6–8 abporal and 6–8 paraporal. Small to medium size, P × E = 19–36 × 23–42 µm; sometimes two different sizes have been found. Exine 3–9 µm thick and ornamentation microreticulate and echinate. The results clearly show the relationships between genera. Moreno-Socías, E., Mejías, J. A., Díez, M. J., 1994: Morfología polínica deLactuceae (Asteraceae) en la Península Ibérica, I.Lactuca y géneros relacionados. — Acta Bot. Malacitana.19: 103–113.     

Mobergellans (Problematica) from the Cambrian of Greenland, Siberia and Kazakhstan

Mobergellans are disc-shaped problematic fossils of phosphatic composition bearing prominent muscle attachment structures on the presumed internal surface. They are found exclusively in sediments of Early Cambrian age, mainly from Laurentia, Baltica and Siberia. Type material ofDiscinella brastadi Poulsen, 1932 from North-East Greenland is re-studied and synonomised withDiscinella micans (Billings, 1871). Specimens from Siberia described asD. brastadi are re-assigned toMobergella hexactina n. sp., and specimens from Siberia described asMobergella radiolata Bengtson, 1968 are re-assigned toM. sibirica n. sp.Aktugaia triangula Mis-Sarzhevsky, 1976 from Kazakhstan is redescribed, and its relationship to other mobergellans is discussed.      

<Emphasis Type="Italic">Enigmagraptus</Emphasis> n. gen., a new graptoloid (Přídolí, Silurian,New South Wales,Australia)

Enigmagraptus n. gen. comprises a group of the smallest known graptolites, with problematical evolutionary ancestors. The species described, all of Přídolí age are:E. yassensis (Rickards &Wright, 1999),E. cf.yassensis, E. mitchelli (Rickards &Wright, 1999) andE. pennyae n. sp.      

Sclerorhachis rechingeri (Asteraceae-Anthemideae), a new species from N. Khorasan

Sclerorhachis rechingeri Iranshahr, sp. n., from N. Khorasan is closely related to the Afghanistan speciesS. caulescens (Aitch. & Hemsl.)Rech. f. andS. polysphaera Rech. f. It differs from both in the thicker and woolly collar and the much larger capitulum.Vorarbeiten zuK. H. Rechinger, Flora Iranica, Nr. 21.     

Zur Cytotaxonomie der Verwandtschaftsgruppe umVeronica orientalis Mill., emend.Ait. in der Türkei

Zusammenfassung Die ArtenVeronica pectinata, V. cuneifolia, V. macrostachya, V. orientalis undV. multifida (SubsektionenAnatolicae-Lycicae Riek undOrientales [Römpp]Riek der SektionVeronica[=Chamaedrys Griseb.]) bilden zusammen mit einigen weiteren Arten in Kleinasien eine sehr formenreiche Verwandtschaftsgruppe, die darüber hinaus in den Gebirgen entlang der Ostküste des Mittelmeeres nach Süden reicht und nach Osten zu noch in Mesopotamien, den kurdischen Gebirgen sowie in Westund Nordpersien vorkommt. Im Nordosten findet die Gruppe Anschluß an eine im Kaukasus reich entfaltete Nachbargruppe (SubsektionGaucasicae Riek).Die jüngste zusammenfassende Bearbeitung dieses Verwandtschaftskreises gabRiek (1935). Der vorliegende erste Beitrag zu einer weiteren Erforschung bringt einige Angaben über Chromosomenzahlen (siehe Übersicht auf S. 159), die bisher völlig gefehlt haben, und stellt vier neue Sippen auf:V. elmaliensis M.Fischer sp. n.,V. macrostachya Vahl ssp.sorgerae M.Fischer ssp. n.,V. macrostachya Vahl ssp.iconiensis M.Fischer ssp. n. undV. macrostachya Vahl ssp.pisidica M.Fischer ssp. n. Die Analyse der Verwandtschaftsbeziehungen unter Berücksichtigung der chromosomalen Befunde ergibt das Bild eines sowohl auf dipoider Stufe stark differenzierten als auch vermutlich durch allopolyploide Sippenkombination äußerst polymorph gewordenen Formenkreises. Die einzelnen Arten des Polyploidkomplexes stehen einander zum Teil sehr nahe, charakteristisch sind retikulate Ähnlichkeits-(Verwandtschafts-?) beziehungen, große Variabilität und unscharfe Sippengrenzen.Die Nomenklatur vonV. dichrus Schott &Kotschy (=V. cilicica Stroh=V. pilosa (Benth.) Römpp) wird erörtert.Die Untersuchung soll auch eine Vorarbeit zu einer künftigen Bearbeitung der GattungVeronica für die Flora of Turkey (Herausgeber P. H.Davis, Edinburgh) darstellen und zugleich einen Beitrag zu der Frage nach den Gesetzmäßigkeiten der Sippendifferenzierung mediterraner Formenkreise liefern. Die GattungVeronica scheint für die zuletzt erwähnte Aufgabe gut geeignet wegen ihrer weiten Gesamtverbreitung und wegen ihrer Artenfülle und ihres kleinräumigen Endemismus in den anatolischen Gebirgen. Im Hinblick auf künftige Untersuchungen sei bemerkt, daß in der SektionVeronicastrum eine ähnliche kleinräumige Differenzierung in den jungen europäisch-asiatischen Gebirgszügen zu beobachten ist.Der Verfasser übernimmt gerne die Revision kritischen Herbarmaterials und wäre auch sehr dankbar für die Zusendung von lebenden Pflanzen und Samen oder von Fixierungen mit entsprechenden Herbarbelegen.The writer would be glad to revise herbarium material of the genusVeronica and gateful for reveiving living plants and seeds or fixations together with vouchers.     

Zwei neue Arten der GattungSaponaria (Caryophyllaceae) aus Afghanistan und Pakistan

Saponaria stenopetala sp.n. in Eastern Afghanistan is close toS. pachyphylla Rech. f. andS. subrosularis Rech. f.—The nearest allies ofS. makranica sp.n. from Western Pakistan and Southeastern Iran areS. kermanensis Bornm. andS. floribunda (Kar. & Kir.)Boiss.

Flora Iranicae praecursores 36–37. — Praecursores praecurrentes in Pl. Syst. Evol.139, 313–317 (1982).     

Interspecific hybridizations among species of theElymus semicostatus andElymus tibeticus groups (Poaceae)

Interspecific hybridizations were made between species of theE. semicostatus group, viz.,E. semicostatus (Nees exSteud.)Meld.,E. validus (Meld.)B. Salomon,E. abolinii (Drob.)Tzvel., andE. fedtschenkoi Tzvel., and species of theE. tibeticus group, viz.,E. pendulinus (Nevski)Tzvel.,E. tibeticus (Meld.)Singh,E. shandongensis B. Salomon, andE. gmelinii (Ledeb.)Tzvel., as well as among species within theE. tibeticus group. All species are tetraploid (2n = 4x = 28) and possess SY genomes. Meiotic pairing data from 24 hybrids involving 17 interspecific combinations are presented. The average number of chiasmata per cell ranged from 17.91 to 26.20 in hybrids within theE. tibeticus group, compared with 7.26 to 22.04 in hybrids between the two species groups. Despite the extensive collection of cytological data, there was no definite evidence for confirming or disproving the separate existence of the two groups.     

Veronica allahuekberensis (Scrophulariaceae), eine neue Art aus der Türkei

Veronica allahuekberensis A. Öztürk, spec. nova, probably endemic to the Allahuekber mountain in N.E. Anatolia, belongs toV. orientalis group of sectionVeronica and resemblesV. cuneifolia D. Don subsp.cuneifolia, endemic to S.W. Anatolia.

     

The genusAntennaria (Asteraceae: Inuleae) in western North America: Morphometric analysis ofAntennaria alborosea,A. corymbosa,A. marginata,A. microphylla,A. parvifolia,A. rosea,andA. umbrinella

Multivariate analysis of vegetative and reproductive characters was used to examine morphological relatedness amongAntennaria alborosea A. E. Porsild,A. corymbosa E. Nels.,A. marginata Greene,A. microphylla Rydb.,A. parvifolia Nutt.,A. rosea Greene, andA. umbrinella Rydb. Both pistillate and staminate plants were examined. Some of the characters examined were variable in one species, but stable in another (i.e., presence or absence of papillae on the achenes). Our analyses indicate that the seven species are morphologically distinct. It is hypothesized that theA. rosea agamic complex arose through hybridization amongA. corymbosa, A. microphylla, A. umbrinella, and possiblyA. dioica (L.)Gaertn. However, hybridization between the three former species and others, as well as their subsequent morphological responses to different environmental conditions causes confusion in recognizing the taxa.Antennaria angustifolia Rydb.,A. arida E. Nels.,A. confinis Greene,A. scariosa E. Nels.,A. foliacea humilis Rydb.,A. concinna E. Nels., andA. viscidula E. Nels. are considered to represent F 1 hybrids.     

Chromosome number and DNA ploidy level reports from Central Europe — 2

Second part of commented chromosome number and DNA ploidy level reports from Central Europe comprising the whole Carpatho-Pannonian region includes the data for following taxa: Tephroseris aurantiaca (2n = 96), T. capitata (2n = 64) and T. integrifolia (2n = 48) by J. Kochjarová from Poland and Slovakia (reports nos. 12–14); Urtica diocia and U. kioviensis (both 2n = 26) by M. Kolník & K. Goliašová from Slovakia (nos. 15–16); Viola hirta (2n = 20), V. odorata (2n = 20), V. reichenbachiana (2n = 20), V. riviniana (2n = 40, 2n ∼ 8x, based on x = 5), V. suavis (2n = 40) and V. × bavarica [V. reichenbachiana × v. riviniana] (2n ∼ 6x, based on x = 5) by P. Mereďa jun., I. Hodálová, P. Mártonfi & V. Kolarčík from Slovakia (nos. 17–22); Fallopia × bohemica [F. japonica × F. sachalinensis] (2n = 66), Thladiantha dubia (2n = 18) and Hieracium longifoliosum (2n = 36) by P. Mráz from Romania and Slovakia (nos. 23–25); Amsinckia calycina (2n = 34) by M. Perny & H. Šípošová from Slovakia (no. 26). For further details and arrangements of reports see the first part (Mráz, 2005).     

Distribution of low molecular weight carbohydrates in the subgenusCeratotropis of the genusVigna (Leguminosae)

The seeds of 9 members of the subgenusCeratotropis (Piper) Verdc., namelyVigna aconitifolia (Jacq.) Maréchal,V. angularis (Willd.) Ohwi et Ohashi,V. minima (Roxb.) Ohwi et Ohashi,V. nakashimae (Ohwi) Ohwi et Ohashi,V. reflexo-pilosa Hayata,V. umbellata (Thumb.) Ohwi et Ohashi,V. mungo (L.) Hepper,V. radiata (L.) Wilczek andV. sp., have been examined. On their low molecular weight carbohydrate compositions, this subgenus has been divided into 2 subgroups, mungo-radiata group and angularis group. Four other species referred to the subgeneraPlectotropis (Schumach.) Bak.,Lasiospron (Benth. emend Piper) Maréchal, Mascherpa et Stainier andVigna, V. vexillata (L.) A. Rich.,V. lasiocarpa (Benth.) Verdc.,V. marina (Burm.) Merr. andV. unguiculata (L.) Walp., were also analyzed and they had distinctive carbohydrate compositions. 1d-4-O-methyl-myo-inositol and 1d-5-O-(α-d-galactopyranosyl)-4-O-methyl-myo-inositol were detected in all species examined exceptV. vexillata, V. mungo andV. radiata.     

Experimental documentation of natural hybridization inCactaceae: Origin of Lloyd's Hedgehog Cactus,Echinocereus ×lloydii

The origin ofEchinocereus ×lloydii Britt. & Rose, pro sp. (Lloyd's Hedgehog Cactus) was investigated using comparative morphology, cytology, biochemistry, and particularly, artificial hybridization. Numerous artificial crosses between the putative parentsE. coccineus Engelm. (a species of claret-up cactus) andE. dasyacanthus Engelm. (Texas Rainbow Cactus) were successful, resulting in the production of hundreds of seeds with hybrid embryos. The F₁ hybrid progeny (i.e., syntheticE. ×lloydii) grew to sexual maturity in about four and one-half years, whereupon successful backcrosses and F₂ generation hybrids were also obtained. The known F₁ hybrids closely approximated naturalE. ×lloydii. The fertility of these syntheticE. ×lloydii was high, like their natural counterparts. The populations ofE. ×lloydii in Pecos County, Texas are inferred to have originated as the result of natural interspecific hybridization. It is assumed thatE. ×lloydii or similar plants may arise wherever the parental taxa grow sympatrically.     

Veronica quezelii undV. saturejoides Vis. subsp.munellensis,zwei neue Sippen der SektionVeronicastrum aus ostmediterranen Gebirgen

Zusammenfassung Veronica quezelii M.Fischer aus hochalpinen Felswänden in Lycien (Kleinasien) ist eine neue Art aus der Verwandtschaft vonV. kotschyana Benth.; sie ist durch die reichliche Bedrüsung der ganzen Pflanze ausgezeichnet.Veronica saturejoides Vis. subsp.munellensis M.Fischer ist eine neue Unterart aus der alpinen Region des Berges Munella in Nord-Albanien, die — ähnlich wie die gleichfalls als Subspecies einzustufendeV. kellereri Degen etUrum. — von der typischen Subspecies hauptsächlich durch Merkmale der Behaarung unterschieden und durch eine Areallücke getrennt ist.

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Summary Veronica quezelii M.Fischer from calcareous rocks of the alpine region of Lycia (Asia Minor) is a new species akin toV. kotschyana Benth. and characterised by the glandular pubescence of the whole plant.Veronica saturejoides Vis. subsp.munellensis M.Fischer is a new subspecies from the alpine region of Mount Munella in Northern Albania. SimilarlyV. kellereri Degen etUrum. has to be treated as subspecies ofV. saturejoides. Both differ from the typical subspecies by characters of the indumentum, and occur in separate areas.

     

A new species of Eumonhystera and three species of Monhystera (Monhysteridae: Nematoda) from Li River,China

A new Eumonhystera species, E. huruii n. sp., together with Monhystera nubiae Eyualem & Coomans, 1996, Monhystera somereni Allgén, 1952, and Monhystera wangi Wu & Hoeppli, 1932 are described from Li river at Guiling, China. Eumonhystera huruii n. sp. is closest to E. barbata Andrássy, 1981, but differs in having shorter anterior setae, much longer reproductive system, shorter and less dorsally bent tail and differently shaped spinneret. M. nubiae and M. wangi are reported here for the first time outside their type locality, whereas M. somereni is reported from China for the first time.     

A new scolenaspidid (Osteostraci, Vertebrata) from the Lower Devonian of Podolia, Ukraine

Anew osteostracan genus and species,Victoraspis longicornualis n. gen., n. sp., is described based on material from Rakovets’, present day Ukraine. This new taxon shares characters with the two generaStensiopelta Denison, 1951 andZychaspis Janvier, 1985. A phylogenetic analysis supports the position ofVictoraspis as the sister group to a monophyleticStensiopelta, while the interrelationships of the various species ofZychaspis are poorly resolved. A morphometric analysis is carried out in an attempt to further resolve the taxonomic affinity. This analysis groups all examinedZychaspis species closely together, and further supports the establishment ofVictoraspis as separate genus.      

A phylogenetic analysis of the lichen familySphaerophoraceae (Caliciales); a new generic classification and notes on character evolution

A phylogenetic analysis of the familySphaerophoraceae (Caliciales, lichenized ascomycetes) has resulted in a new generic classification. Notes on character evolution are given. The generaSphaerophorus s. str.,Bunodophoron andLeifidium, gen. nov., are accepted.Pleurocybe andPseudosphaerophorus are considered synonyms ofBunodophoron andThysanophoron is considered synonym toSphaerophorus. The following new combinations are proposed:Bunodophoron coomerense (Ohlsson)Wedin,B. diplotypum (Vain.)Wedin,B. dodgei (Ohlsson)Wedin,B. flaccidum (Kantvilas & Wedin)Wedin,B. formosanum (Zahlbr.)Wedin,B. imshaugii (Ohlsson)Wedin,B. insigne (Laurer)Wedin,B. kinabaluense (M. Satô)Wedin,B. macrocarpum (Ohlsson)Wedin,B. madagascareum (Nyl.)Wedin,B. microsporum (Ohlsson)Wedin,B. murrayi (Ohlsson)Wedin,B. notatum (Tibell)Wedin,B. ohlssonii (Wedin)Wedin,B. patagonicum (C. W. Dodge)Wedin,B. ramuliferum (I. M. Lamb)Wedin,B. scrobiculatum (C. Bab.)Wedin,B. tibellii (Wedin)Wedin,B. whakapapaense (Wedin)Wedin, andLeifidium tenerum (Laurer)Wedin.     

Cynoglossopsis somaliensis,sp. nov., und die GattungCynoglossopsis (Boraginaceae)

Cynoglossopsis somaliensis H. Riedl, sp. nov., is described as a new species of the hitherto monotypic genusCynoglossopsis Brand which is closest related toCynoglossum L. but has to be included inBoraginoideae-Eritrichieae from the way the nutlets are attached to the gynobasis.

     

Zur cytotaxonomie vonVeronica chamaedrys L. agg., II.: subsp.micans M.Fischer,subsp. nova,eine weitere diploide Sippe

Zusammenfassung Veronica chamaedrys L. subsp.micans M.Fischer, unterscheidet sich vonV. chamaedrys L. subsp.chamaedrys hauptsächlich durch die meist hellgrün gefärbten und im Leben oberseits stets schwach glänzenden Laubblätter, deren Blattrand durchschnittlich mehr (9–11) Zähne aufweist, durch die stets drüsenlose Infloreszenzbehaarung, die kleineren Blüten mit schmäleren Sepalen und hellblau (nicht leuchtend-sattblau) gefärbten Corollen sowie durch die diploide Chromosomenzahl (2 n=16). Sie tritt in der mittleren Bergstufe zusammen mit subsp.chamaedrys auf, ersetzt sie jedoch in der oberen Bergstufe und in der Krummholzstufe an±gleichen Standorten fast vollständig (mit Ausnahme der stark eutrophierten Standorte, die ausschließlich von subsp.chamaedrys besiedelt sind). Die neue Sippe ist bisher nur aus den nordöstlichen Alpen bekannt geworden (vgl. S. 77).

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Cytotaxonomy ofveronica chamaedrys L. agg., II.: subsp.micans M.Fischer, a new diploid race

Summary Veronica chamaedrys L. subsp.micans M.Fischer differs fromV. chamaedrys L. subsp.chamaedrys mainly by: leaves bright green, in the living state always slightly but distinctly shining on the upper surface; margins on the average with more (9–11) teeth at each side; indumentum of the inflorescence without any glands; flowers relatively small with bright blue corollas and narrow sepals; chromosome number diploid: 2 n=16. In the upper mountain and subalpine region, where it replaces the common tetraploid subsp.chamaedrys L.; it is known up to now only from the Northeastern Alps (see p. 77).

     

Chromosome numbers of the European seagrasses

The chromosome numbers of the five European seagrasses have been determined in material from several sites along the coasts of the Atlantic Ocean, the North Sea and the Mediterranean:Zostera marina L., 2n = 12;Z. noltii Hornem., 2n = 12;Posidonia oceanica (L.)Delile, 2n = 20;Cymodocea nodosa (Ucria)Aschers., 2n = 14, 2n = 28;Halophila stipulacea (Forsk.)Aschers., 2n = 18. The difference in chromosome morphology betweenZ. marina andZ. noltii supports the division of the genus into two subgenera.