Technological variation in the earliest Oldowan from Gona,Afar, Ethiopia

Inter-site technological variation in the archaeological record is one of the richest potential sources of information about Plio-Pleistocene hominid behavior and evolution. However, appropriate methods for describing and comparing Oldowan assemblages have yet to be agreed upon, and interpretation of the early record remains highly controversial. Particularly salient is disagreement over whether the Oldowan is a single technological phenomenon or is more accurately divided into multiple regional and/or chronological traditions, perhaps including a less developed Pre-Oldowan phase in the late Pliocene. Some of this disagreement reflects theoretical and methodological differences between research traditions and some is more directly evidential. Here we present a framework for describing and interpreting Oldowan variation and apply it to three Pliocene assemblages (EG-10, EG-12, and OGS-7) from Gona, all dated to c. 2.6 million years (Ma). Results indicate proficient knapping and a full range of Oldowan reduction strategies in these earliest known occurrences, consistent with the idea of an Oldowan “technological stasis” from 2.6–1.6 Ma. Patterns of variation in raw material selection and predominant reduction strategy at each site clearly indicate the importance of cultural transmission in the Oldowan, but confounding ecological and economic variation continue to render interpretation in terms of multiple tool making traditions or species inappropriate. We propose that cultural transmission and ecological adaptation should be recognized as complementary, rather than mutually exclusive, mechanisms in future attempts to explain Oldowan technological variation.     

The dating and interpretation of a Mode 1 site in the Luangwa Valley, Zambia

Flake based assemblages (Mode 1) comprise the earliest stone technologies known, with well-dated Oldowan sites occurring in eastern Africa between ∼ 2.6-1.7 Ma, and in less securely dated contexts in central, southern and northern Africa. Our understanding of the spread and local development of this technology outside East Africa remains hampered by the lack of reliable numerical dating techniques applicable to non-volcanic deposits. This study applied the still relatively new technique of cosmogenic nuclide burial dating (¹⁰Be/²⁶Al) to calculate burial ages for fluvial gravels containing Mode 1 artefacts in the Luangwa Valley, Zambia. The Manzi River, a tributary of the Luangwa River, has exposed a 4.7 m deep section of fluvial sands with discontinuous but stratified gravel layers bearing Mode 1, possibly Oldowan, artefacts in the basal layers. An unconformity divides the Manzi section, separating Mode 1 deposits from overlying gravels containing Mode 3 (Middle Stone Age) artefacts. No diagnostic Mode 2 (Acheulean) artefacts were found.Cosmogenic nuclide burial dating was attempted for the basal gravels as well as exposure ages for the upper Mode 3 gravels, but was unsuccessful. The complex depositional history of the site prevented the calculation of reliable age models. A relative chronology for the full Manzi sequence was constructed, however, from the magnetostratigraphy of the deposit (N>R>N sequence). Isothermal thermoluminescence (ITL) dating of the upper Mode 3 layers also provided consistent results (∼78 ka). A coarse but chronologically coherent sequence now exists for the Manzi section with the unconformity separating probable mid- or early Pleistocene deposits below from late Pleistocene deposits above. The results suggest Mode 1 technology in the Luangwa Valley may post-date the Oldowan in eastern and southern Africa. The dating programme has contributed to a clearer understanding of the geomorphological processes that have shaped the valley and structured its archaeological record.     

Current research on the late Pliocene and Pleistocene deposits north of Homa Mountain, southwestern Kenya

The late Pliocene and Pleistocene sediments of the Homa Peninsula in southwestern Kenya are richly fossiliferous, preserve Early Stone Age archaeological traces and provide one of the few paleoanthropological data sets for the region between the branches of the East African Rift Valley. This paper presents preliminary results of our ongoing investigation of late Pliocene and Pleistocene deposits at the localities of Rawi, Kanam East, Kanam Central and Kanjera. While fossils have been collected from the peninsula since 1911, little systematic effort has been made to place them into a broader litho-and chronostratigraphic framework. This project has conclusively demonstrated that fossils occur in good stratigraphic context at all of the study localities and that claims of sediment slumping (Boswell, 1935) have been greatly overstated (Behrensmeyer et al., 1995; Plummer & Potts, 1989). A provisional chronostratigraphic framework based on magneto- and biostratigraphy is presented here. We have revised the Plio-Pleistocene stratigraphy of the Rawi and Kanam gullies to include three formations: the Rawi, Abundu and Kasibos Formations. Based on magneto- and biostratigraphy, these formations are dated between approximately three and one m.y.a. (Gauss Chron-Jaramillo Subchron) (Cande & Kent, 1995). The Apoko Formation unconformably overlies the others and may be middle to late Pleistocene in age. All formations contain rich patches of fossils, and Acheulean artifacts have been surface collected from the Abundu and Kasibos Formations. Deposition of the fossil- and artefact-bearing sediments at Kanjera North began in the early Pleistocene and continued into the middle Pleistocene. Deposition at Kanjera South began over one million years earlier than previously thought, at approximately 2.2 m.y.a., and continued into the Olduvai Subchron (1.770-1.950 m.y.a.; Cande & Kent, 1995). Excavations have recovered Oldowan artefacts in association with well-preserved fossil fauna near the base of the sequence, the oldest archaeological traces yet known from southwestern Kenya.     

Mio-pliocene faunal exchanges and african biogeography: the record of fossil bovids

The development of the Ethiopian biogeographic realm since the late Miocene is here explored with the presentation and review of fossil evidence from eastern Africa. Prostrepsiceros cf. vinayaki and an unknown species of possible caprin affinity are described from the hominid-bearing Asa Koma and Kuseralee Members (～5.7 and ～5.2 Ma) of the Middle Awash, Ethiopia. The Middle Awash Prostrepsiceros cf. vinayaki constitutes the first record of this taxon from Africa, previously known from the Siwaliks and Arabia. The possible caprin joins a number of isolated records of caprin or caprin-like taxa recorded, but poorly understood, from the late Neogene of Africa. The identification of these two taxa from the Middle Awash prompts an overdue review of fossil bovids from the sub-Saharan African record that demonstrate Eurasian affinities, including the reduncin Kobus porrecticornis, and species of Tragoportax. The fossil bovid record provides evidence for greater biological continuity between Africa and Eurasia in the late Miocene and earliest Pliocene than is found later in time. In contrast, the early Pliocene (after 5 Ma) saw the loss of any significant proportions of Eurasian-related taxa, and the continental dominance of African-endemic taxa and lineages, a pattern that continues today.     

Raw material selectivity of the earliest stone toolmakers at Gona, Afar, Ethiopia

Published evidence of Oldowan stone exploitation generally supports the conclusion that patterns of raw material use were determined by local availability. This is contradicted by the results of systematic studies of raw material availability and use among the earliest known archaeological sites from Gona, Afar, Ethiopia. Artifact assemblages from six Pliocene archaeological sites were compared with six random cobble samples taken from associated conglomerates that record pene-contemporaneous raw material availability. Artifacts and cobbles were evaluated according to four variables intended to capture major elements of material quality: rock type, phenocryst percentage, average phenocryst size, and groundmass texture. Analyses of these variables provide evidence of hominid selectivity for raw material quality. These results demonstrate that raw material selectivity was a potential component of Oldowan technological organization from its earliest appearance and document a level of technological sophistication that is not always attributed to Pliocene hominids.     

Middle Pliocene vegetation: reconstructions, paleoclimatic inferences, and boundary conditions for climate modeling

The general characteristics of global vegetation during the middle Pliocene warm period can be reconstructed from fossil pollen and plant megafossil data. The largest differences between Pliocene vegetation and that of today occurred at high latitudes in both hemispheres, where warming was pronounced relative to today. In the Northern Hemisphere coniferous forests lived in the modern tundra and polar desert regions, whereas in the Southern Hemisphere southern beech apparently grew in coastal areas of Antarctica.Pliocene middle latitude vegetation differed less, although moister-than-modern conditions supported forest and woodland growth in some regions now covered by steppe or grassland. Pliocene tropical vegetation reflects essentially modem conditions in some regions and slightly cooler-than-or warmer-than- modern climates in other areas. Changes in topography induced by tectonics may be responsible for many of the climatic changes since the Pliocene in both middle and lower latitudes. However, the overall latitudinal progression of climatic conditions on land parallels that seen in the reconstruction of middle Pliocene sea-surface temperatures.Pliocene paleovegetational data was employed to construct a 2 ° × 2 ° global grid of estimated mid-Pliocene vegetational cover for use as boundary conditions for numerical General Circulation Model simulations of middle Pliocene climates. Continental outlines and topography were first modified to represent the Pliocene landscape on the 2 ° × 2 ° grid. A modern 1 ° × 1 ° vegetation grid was simplified and mapped on this Pliocene grid, and then modified following general geographic trends evident in the Pliocene paleovegetation data set.     

Research on late Pliocene Oldowan sites at Kanjera South, Kenya

The late Pliocene is notable for the appearance of two new hominid genera as well as the first archaeological sites, generally attributed to the Oldowan Industrial Complex. However, the behavioral ecology of Oldowan hominids has been little explored, particularly at sites older than 2.0 Ma. Moreover, debates on Oldowan hominid foraging ecology and behavior have centered on data from only two regions, and often from single site levels. Here we describe the preliminary results of our investigation of Oldowan occurrences at Kanjera South. These occurrences preserve the oldest known traces of hominid activity in southwestern Kenya, and unlike most of the Oldowan sites in the 2.0-2.5 Ma time interval, artefacts are found in spatial association with a well-preserved fauna. In 1996 and 1997, this project initiated the first excavation program for Kanjera South. Magneto- and biostratigraphy indicate that deposition began approximately 2.2 Ma, substantially earlier than previously thought. At Excavation 1, artefacts were found in spatial association with a taxonomically diverse faunal assemblage in Beds KS-1 and KS-2. Excavation 2 yielded a partial hippopotamus axial skeleton with artefacts in KS-3. Cores from both sites were incidentally flaked and represent a Mode I lithic technology indistinguishable from the Oldowan. Approximately 15% of the artefacts were manufactured from non-local raw materials, indicating a flow of resources into the area. Stable isotopic analysis of KS-1 and KS-2 pedogenic carbonates suggests that the Excavation 1 assemblages formed in a relatively open (>75% C4 grass) habitat. The Excavation 1 and 2 faunas contain a high proportion of equids relative to Oldowan accumulations from Bed I Olduvai Gorge, Tanzania. Beds KS-1 and KS-2 thus preserve traces of Oldowan hominid activities in a more open setting than has been previously documented.     

A new large Pliocene colobine species (Mammalia: Primates) from Asa Issie, Ethiopia

The Colobinae (Mammalia: Primates) are relatively unknown from the African middle to late Miocene. When they appear in the Pliocene they are unambiguous and already fairly diverse taxonomically, geographically, and ecologically. Discoveries from Pliocene sediments in eastern and southern Africa document a radiation of large-bodied colobines very different from those known today. Paleontological research in Ethiopia has recently led to the discovery and identification of another large-bodied colobine species from the early Pliocene site of Asa Issie, discovered in 2000. This new colobine is larger than but morphologically very similar to its sister taxon Kuseracolobus aramisi, an older taxon also described from the Middle Awash. This new species has significant implications for our understanding of the Pliocene colobine adaptive radiation.     

泥河湾盆地麻地沟E5旧石器地点的遗址成因与石器技术

麻地沟E5旧石器地点位于泥河湾盆地东端岑家湾台地古人类活动集中区,埋藏于泥河湾层灰白色-灰黄色细砂和粉砂层内。遗址发现于2007年,中国科学院古脊椎动物与古人类研究所于2014年对其进行考古发掘,揭露面积22m~2,出土143件石制品、8件动物碎骨和235件砾石。遗址成因(标本分布状况、原料与个体大小、风化磨蚀状况和标本空间聚集特征等)分析表明,遗址形成受到明显的水流搬运和改造。石制品原料均来自遗址周围1km~2范围内,岩性以硅质白云岩、燧石和火山岩为主;锤击法剥片和修理石器,剥片和修理随意,石制品形态无定型,显示旧石器时代早期奥杜威(Oldowan)工业特点。初步古地磁年代测定显示古人类在该遗址活动的时间大致为1.20 Ma。该遗址的发掘对研究泥河湾盆地早更新世遗址成因与古人类的生存方式具有重要意义。     

Geological setting of Pliocene rifting and deposition in the Afar Depression of Ethiopia

Tectonic extension of the Afar amounts to 10–30 km since 1–2 Ma and to 40–60 km since 3–5 Ma ago, or rougly 1–2 cm per year. Active faulting, volcanism, and development of the Pliocene and younger Afar grabens with their rich hominid and other vertebratefaunas, have been controlled by ENE and WNW oriented lineaments or ancient zones of structural weakness. These lineaments also controlled the alignment of the Gulf of Aden and Red Sea, respectively.Crustal stresses resulting from the late Miocene (Messinian) salinity crisis in the Mediterranean basin may have triggered renewed tectonic movements along certain pre-existing lineaments such as the Levant Shear. Separation of Africa from Arabia (and hence the inception of Afar) was controlled by tectonic events far removed from NE Africa, including possible Miocene fusion of the Indo-Arabian plates.During the early Pliocene, the Ethiopian uplands were far lower (possibly by 1000 m) and the southern Afar-Middle Awash region was higher, so that topographic and climatic contrasts between plain and plateau were less pronounced and the Afar climate was less arid than today.There was a major change from lacustrine to fluviatile deposition in the Middle Awash valley soon after 4·0–3·8 Ma ago, caused by extensional tectonics. Within the topographic constraints imposed by volcano-tectonic activity, regional climatic oscillations have controlled the detailed pattern of Pliocene (and later) sedimentation in Afar.     

Raw material selectivity in Late Pliocene Oldowan sites in the Makaamitalu Basin, Hadar, Ethiopia

We report the results of an analysis of raw material selection patterns in the assemblages from two Late Pliocene in situ archaeological localities in the Makaamitalu Basin (Hadar, Ethiopia). While the same local conglomerate was used as a raw material source for both archaeological occurrences, different selection criteria are identified. At A.L. 894, selection for quality is subtle and the clearest selection is against non-homogeneous raw materials. In the A.L. 666 assemblage, higher-quality raw materials were selected and some rare raw materials reached the locality from unknown sources. A comparison between the Makaamitalu and other Oldowan assemblages reveals an overall shift toward higher complexity of both selectivity and transport behaviors from ca. 2.0 Ma onward, contrasting a typo-technological conservatism that pertains until ∼1.6 Ma. It is hypothesized that an increase in complexity of behaviors related to raw material selection and acquisition involved changes in the intensity and fidelity of technological knowledge transmission.     

Fingerprinting facies of the Tuff IF marker,with implications for early hominin palaeoecology,Olduvai Gorge,Tanzania

The top of Tuff IF marks the upper boundary of the Plio-/Pleistocene Bed I succession exposed in Olduvai Gorge, NE Tanzania, a tephrostratigraphic interval that is well known for remarkable Oldowan archaeological and vertebrate fossil assemblages, including early hominins. Geochemically, Tuff IF is characterized by a relatively consistent silica-undersaturated trachytic to phonolitic composition that relates both in terms of numerical ages (1.79 Ma) and compositional constraints to Olmoti volcano, located 22–38 km E of Olduvai Gorge.Measured Tuff IF sections are characterized by a thick unwelded pyroclastic flow in proximal settings and a succession of surges and ashfalls that interfingers with fluvio-lacustrine deposits of the Olduvai Basin in the medial to distal settings. Only in those areas with a relatively low topographic gradient, in distal reaches and beyond the toe of an Olmoti-sourced volcaniclastic fan, did the Tuff IF marker develop a typical threefold subdivision comprising: (1) primary surges and minor fallout, (2) reworked pumice units as lateral correlatives of proximally emplaced pyroclastic flows and (3) a succession of mass flows in conjunction with aeolian and fluvially reworked units.The pyroclastic marker unit is thus highly heterogeneous with regard to its vertical and lateral facies architecture and this has immediate effects on its preservation potential and in situ burial of fossils and stone artifacts. Even though the volcanically-related environmental perturbations were probably more severe at the eastern lake margin, the evidence for at least temporary freshwater sources and trees suggests environments conducive to hominin activities, but not during emplacement of Tuff IF pyroclastic flows and surges. This inference is supported by the presence of rich Oldowan stone artifact assemblages immediately preceding and following the deposition of Tuff IF, but only extremely sparse archaeological traces from one site in Tuff IF, restricted to the upper, fluvially reworked portion of Tuff IF.Tuff IF facies record the maximum of a regressive (drying) cycle that correlates with regional marine arid indicators. An ecological crisis covering ～ 2000–3500 a of time during Tuff IF deposition resulted not only from the lethal effects of explosive volcanism but also from its coincidence with a pronounced period of climate induced drought. These combined effects appear to have made at least the eastern basin uninhabitable by hominins and other vertebrates for most of the time during which Tuff IF accumulated.     

First porcupine fossils (Mammalia,Rodentia) from the late Miocene of NW Iran,with notes on late Miocene–Pliocene dispersal of porcupines

This paper describes the first fossil porcupine remains from Iran. Four upper cheek teeth and two fragmentary lower incisors present sufficient characters for identification as Hystrix aryanensis, a species previously known from the late Miocene locality of Molayan (Afghanistan) estimated at ca. 7–8 Ma. The dental features of porcupines are discussed to show their systematic value and highlight evolutionary trends in late Miocene and Pliocene porcupines. This study also discusses the dispersal history of fossil porcupines in relation to paleobiogeographic provinces and environmental changes during late Miocene to late Pliocene time.     

A new species of Kolpochoerus (Mammalia: Suidae) from the Pliocene of Central Afar, Ethiopia: Its Taxonomy and Phylogenetic Relationships

Kolpochoerus (Mammalia: Suidae) is a suine genus represented by a number of species from Plio-Pleistocene sites in Africa. While the general trends in Kolpochoerus evolution are broadly known, gaps in the fossil record preclude an understanding of the details of its evolutionary tempo and mode. Here, we describe a new species, Kolpochoerus millensis, based on new fossil material from the Woranso-Mille and Gona sites in the Central Afar region of Ethiopia and dated to 3.5–3.8 million years ago (Ma). Third molars of K. millensis are metrically and morphologically intermediate between the early Pliocene K. deheinzelini and earliest late Pliocene K. afarensis. It appears that K. deheinzelini, K. millensis, and K. afarensis are temporally disjunct and phenetically distinguishable parts of a single evolving lineage. The recognition of these chronospecies provides additional evidence for anagenetic evolution. It demonstrates clearly the presence of transitional forms in the fossil record. The extensive and well-dated Kolpochoerus fossil record serves as one of the best documented examples of the occurrence of phyletic evolution. Moreover, K. millensis is one of the best biochronological markers in eastern Africa for the time between 3.5 and 3.8 Ma.     

Stratigraphy, artefact industries and hominid associations for Sterkfontein, member 5

A revised stratigraphy for the early hominid site of Sterkfontein (Gauteng Province, South Africa) reveals a complex distribution of infills in the main excavation area between 2.8 and 1.4 m.y.a, as well as deposits dating to the mid to late Pleistocene. New research now shows that the Member 4 australopithecine breccia (2.8-2.6 Ma) extends further west than was previously thought, while a late phase of Member 4 is recognized in a southern area. The artefact-bearing breccias were defined sedimentologically as Member 5, but one supposed part of these younger breccias, the StW 53 infill, lacks in situ stone tools, although it does appear to post-date 2.6 Ma when artefacts first appear in the archaeological record. The StW 53 hominid, previously referred to Homo habilis, is here argued to be Australopithecus. The first artefact-bearing breccia of Member 5 is the Oldowan Infill, estimated at 2-1.7 Ma. It occupies a restricted distribution in Member 5 east and contains an expedient, flake-based tool industry associated with a few fossils of Paranthropos robustus. An enlarged cave opening subsequently admitted one or more Early Acheulean infills associated in Member 5 west with Homo ergaster. The artefacts attest to a larger site accumulation between ca. 1.7 and 1.4 Ma, with more intensive use of quartzite over quartz and a subtle but important shift to large flakes and heavier-duty tools. The available information on palaeoenvironments is summarized, showing an overall change from tropical to sub-tropical gallery forest, forest fringe and woodland conditions in Member 4 to more open woodland and grassland habitats in the later units, but with suggestions of a wet localized topography in the Paranthropus -bearing Oldowan Infill.     

Depositional environments, archeological occurrences and hominids from Members E and F of the Shungura Formation (Omo basin, Ethiopia)

The several known archeological occurrences from the Shungura Formation (Members E and F) are discussed within the context of their particular depositional and paleoenvironmental settings, and the broader context of the Pliocene of the Turkana Basin. These assemblages date from 2·3–2·4 Ma, and are among the oldest yet known in Africa. In situ archeological assemblages are restricted to the distal edge of fluviatile levees, behind gallery forests and at the edge of open savanna. Extensive and rather dense scatters of slightly disturbed quartz artifacts are linked to large expanses of braided stream situations. Lag components are derived from stream beds. There is no available evidence of occupation either under riverine or deltaic forest settings or on dryland. Hominid skeletal remains from this interval, none of which occur at or are directly associated with such archeological occurrences, are attributed to two hominid taxa. Most specimens represent a “robust” australopithecine and are attributed here to A. aethiopicus. A few hominid teeth are attributed to aff. Homo sp.     

A Review of the Fossil Record of the Genus Itea (Iteaceae, Saxifragales) with Comments on its Historical Biogeography

Itea is a genus of about 20 species of trees and shrubs that are today native to southeastern North America, eastern Asia, and eastern Africa. In this paper, I review the fossil record of Itea, which is based on four types of fossils: diporate, psilate pollen attributed to Itea or the dispersed pollen genus Iteapollis; carpofossils representing fruits and seeds attributed to Itea europaea; flowers preserved in amber and assigned to Adenanthemum iteoides; and leaf impressions attributed to Itea. The distributions of these fossils indicate that Itea was present in western North America from the early Eocene to Miocene, in eastern North America beginning no later than the early Miocene, and in western Eurasia from the late Eocene to Pliocene. Only one datapoint is known from eastern Asia; it is early Miocene in age. Based on the fossil record, it can be inferred that Itea crossed between continents over both the Bering Land Bridge and North American Land Bridge, and that it reached Africa from Europe via Anatolia. Thus, it is predicted that the sole extant North American species, I. virginica, may be most closely related to the sole extant African species, I. rhamnoides. The potential application of Itea fossils to calibrating phylogenetic trees generated from molecular sequence data is also discussed.     

A new skeleton of Theropithecus brumpti (Primates: Cercopithecidae) from Lomekwi, West Turkana, Kenya

A relatively complete skeleton of the fossil papionin, Theropithecus brumpti, from the site of Lomekwi, west of Lake Turkana, Kenya, is here described. The specimen, KNM-WT 39368, was recovered at the site of LO 5 (3°51′N and 35°45′E), from sediments dated to approximately 3·3 Ma. The skeleton is that of an old adult male and preserves a number of articulated elements, including most of the forelimbs and tail. The cranial morphology is that of a large, early T. brumpti, exhibiting a deep mandible with a deeply excavated mandibular corpus fossa, and mandibular alveoli and cheek teeth arrayed in a reversed Curve of Spee. The forelimb skeleton exhibits a unique mixture of characteristics generally associated with a terrestrial locomotor habitus, such as a narrow scapula and a highly stable elbow joint, combined with those more representative of habitual arborealists, such as muscle attachments reflecting a large rotator cuff musculature and a flexible shoulder joint. The forelimb of KNM-WT 39368 also presents several features, unique toTheropithecus , which represent adaptations for manual grasping and fine manipulation. These features include a large, retroflexed medial humeral epicondyle (to which large pronator, and carpal and digital flexor muscles attached) and proportions of the digital rays that denote capabilities for precise opposition between the thumb and index finger. Taken together, these features indicate that one of the earliest recognized representatives of Theropithecus exhibited the food harvesting and processing anatomy that distinguished the genus through time and that contributed to its success throughout the later Pliocene and Pleistocene. Based on the anatomy of KNM-WT 39368 and the known habitat preference of T. brumpti, the species is reconstructed as being a generally terrestrial but highly dexterous, very large-bodied, sexually dimorphic, and possibly folivorous papionin. T. brumpti was adapted for propulsive quadrupedal locomotion over generally even ground, and yet was highly adept at manual foraging. The estimate of 43·8 kg body mass for KNM-WT 39368 renders unlikely the possibility that the species, or at least adult males of the species, were highly arboreal. T. brumpti, as represented by KNM-WT 39368, is seen as a large, colorfully decorated, and basically terrestrial papionin that was restricted to riverine forest habitats in the Lake Turkana Basin from the middle to latest Pliocene.     

An analysis of two Myosorex species (Soricidae) from the Early Pliocene site of Langebaanweg (West coast, South Africa) using geometric morphometrics, linear measurements, and non-metric characters

This represents the first study of the soricid community from the Early Pliocene site of Langebaanweg (LBW) which is represented by four species, two of which belong to the soricid genus Myosorex (Forest shrew). Using geometric morphometrics the two Myosorex fossil species are compared with each other, and the extant western and eastern Cape species: Myosorex varius, M. cafer and M. longicaudatus. In addition, the results from an analysis of a number of non-metric characters and linear measurements on the fossil species are presented. The mandibles of the fossil species are characterized by extremely broad teeth, ascending rami which are relatively gracile and differ in shape and size from modern species, and the existence of a number of primitive features of the mandibles and teeth. This evidence is interpreted as indicating that the two LBW Myosorex species represent an archaic lineage now extinct in southern Africa.     

A revision of the fossil Canidae (Mammalia) of north‐western Africa

Abstract: The fossil record of the Canidae in North‐western Africa begins near the Miocene–Pliocene boundary with a form close to Nyctereutes, a genus best known in the late Pliocene of Ahl al Oughlam. This site yields two other canids. Vulpes hassani sp. nov. is a small fox, probably ancestral to the modern V. rueppelli, recorded from the Middle Pleistocene onwards. Lupulella paralius sp. nov. is a primitive jackal that probably belongs to the clade of modern African jackals. In the middle Pleistocene, the most common canid is Lupulella mohibi sp. nov., remarkable by its Nyctereutes‐like dentition and primitive skull‐features. These are all endemic forms, but V. vulpes and C. aureus, of northern origin, appear in the course of the middle Pleistocene. Lycaon has a sparse record in the middle and late Pleistocene.