Les Ostracodes et le Quaternaire d’Aigion (golfe de Corinthe, Grèce)

The Upper Pleistocene and the Holocene of Aigion have delivered an abundant microfauna with 35 ostracode species. The Pleistocene from Aigion borehole generally provides Ostracodes from oligohaline environment with Cyprideis torosa, Candona angulata and Tyrrhenocythere amnicola while, in the Holocene, marine infralittoral species dominate with Cytheridea neapolitana, Carinocythereis whitei, Loxoconcha ovulata and Cytherois frequens. The marine sedimentation occurred at depth from some meters to some tens of meters. In the Aghios Constantinos section, the lagoonal marls are characterised by Euxinocythere schuldtae and a dwarf species of Xestoleberis. Then, a drastic environmental change occurs around the Pleistocene-Holocene boundary and presumably affected the whole Corinth gulf.     

Les industries acheuléennes de la France septentrionale

The excavations of several open air sites in Northern France and, especially in the Somme basin, have given numerous data concerning the age and the environment of the human occupations for the period from MIS 12 to MIS 8. The quality of the raw material and the function of the different sites have a great effect on the composition of the lithic assemblages as it has been seen for the different sites located near the confluence of the Somme and Avre (Cagny-la Garenne, Cagny-l’Épinette, Ferme de l’Épinette, Gentelles). The characteristics of the lithic industries of Gouzeaucourt show the complexity of the transition from Lower to Middle Palaeolithic.     

Les faunes d’ammonites du Pliensbachien inférieur des séries dauphinoises de la Durance (Jurassique inférieur, Barrage de Serre-Ponçon, Hautes-Alpes, France)

The Liassic ammonite faunas and especially those of the Early Pliensbachian are of major importance for the understanding of the stratigraphic framework of the “Dauphiné” zone of the southern subalpine ranges (“dôme de Remollon”). Until now, only listed by the authors, these faunas are, for the first time, described and illustrated in the present paper. With about twenty species, these faunas cover the whole of the Early Pliensbachian. There are poorly diversified and of clearly Euroboreal affinities. Nearly all of the species are abundant and common in most of the NW European fossiliferous localities. Despite of a paleogeographical location fairly close to the Alpine zones (Southern Alps and Austroalpine units) with rather diversified faunas of mainly Mediterranean affinities, none Mediterranean taxa are present in the studied area. Unfortunately, the current stratigraphic and paleogeographic knowledge do not allow to ascertain if this reflects a separation due to paleoenvironmental or true paleogeographic barriers. In this latter case, the general geologic implications would be remarkable.     

Le Paléolithique moyen et supérieur de la Hongrie du nord : nouvelles investigations dans la région d’Eger

The region of Eger town at the foot of the Bükk Mountains in Northeastern Hungary has particular importance for the understanding the Middle and Upper Palaeolithic periods of the Carpathian Basin. One of the key-problems is the association of carinated tools and foliate points in the lithic assemblages of the sites. New investigations at the Egerszalók-K?vágó site give fundamental data for the animated discussion on the relation between Aurignacian and Szeletian.     

Paléogéographie des chasseurs de l’Europe du Paléolithique supérieur : répartition et spécialisation des sites

During the Upper Paleolithic, Europe was occupied by human populations mainly living from hunting of big terrestrial mammals. A repartition of fauna and of archeological sites was done with a database: a repartition of Upper Paleolithic sites in central and western Europe. This study shows that the european space was structured following several factors; climate, environment, cynegetic behaviours… We observe a partition of Europe with a mediterranean area with mainly deer, and ibex and an atlantic area with reindeer. Equally, most of the occupations are localized in the “aquitano-cantabric horn” around the Gulf of Gascony. The occupation of Europe undergone a pulsation with a retreat of occupations toward the west and the south during the glacial maximum, then a colonization of North Europe during the Tardiglacial climatic amelioration.     

Des steppes aux déserts, à l’Aurignacien. Les hommes modernes venus d’Asie

From east to west, Aurignacian populations and civilizations moved into Europe during dry and cold climatic conditions. The zones of origin in the Middle East at that time had a steppic environment with large fauna that were rich caloric resources. Alongside many other examples, this new cultural unit has strict parallels that are clearly evidenced at Jabrud (Syria) and Yafteh (Iran). Now a desert region, during the Aurignacian this vast geographic and ethnic region extended on both sides of the Central Zagros Mountains, towards Central Asia where immense steppes seem having formed the epicenter.     

Les ammonites traditionnellement regroupées dans la sous-famille des Bouleiceratinae (Hildoceratidae, Toarcien) : aspects phylogénétiques, biogéographiques et systématiques

The genera usually considered into the Bouleiceratinae subfamily belong in fact to different lineages of the Hildoceratidae family. An approach using paleogeographical, stratigraphical and morphological arguments on one hand and a cladistic approach on the other hand have allowed to separate 3 clusters with different paleogeographical and stratigraphical localisations. These are:

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the Bouleiceratinae s.s. (genera Bouleiceras, Nejdia and Kohaticeras) from lower Toarcian;

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the Leukadiellinae (genera Leukadiella and Renziceras) from middle Toarcian;

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and the Paroniceratinae (genera Frechiella, Paroniceras and Oxyparoniceras) from middle and upper Toarcian. These latters are mainly present and the Tethyan domain, but are also known in the NW of Europe and especially in France where new populations have been studied in the Causses, in the centre-ouest and in the Lyon area. New taxa have been erected here: Oxyparoniceras (Neoparoniceras) nov. sub. gen., O. (N.) brocardi nov. sp. and O. (N.) sciaui nov. sp.

     

Puycelci, nouveau site à vertébrés de la série molassique d’Aquitaine. Densité et continuité biochronologique dans la zone Quercy et bassins périphériques au Paléogène

After the previous lacking evidence allowed the interpretation of a non-deposition episod, the newly recorded mammal fauna from Puycelci (Tarn, SW France, close South to the Quercy paleokarstic area) now shows the MP 26 standard level (Late Oligocene, Early Chattian) being represented within the Eastern Aquitaine molassic basin filling. According to the evolutionary stage of known mammal lineages, the most significant Puycelci fossils are the Issiodoromys pauffiensis rodent and the Metriotherium cf. sarelense nov. sp. (described in this paper) artiodactyl ungulate. This new dating exemplifies the improvement of the mammalian biochronologic record as regards both the Quercy paleokarst and the surrounding lacustrine Tertiary basins W, S, and S-E to the Quercy, as well as, directly E and N-E, the smaller tectonic Tertiary basins within the crystalline Massif Central basement. The updated record shows the strong extension in time of the paleokarstic data, down to the Middle and Early Eocene, and up to the Early Miocene. Regarding the classically reported Late Eocene to Late Oligocene span, as well in the paleokarst and the peripheral basins, the available biochronological data now attains a valuable density level, the variation of which is significant. On one side there are periods (Late Eocene, Early Oligocene, Early Late Oligocene) with high biochronologic density, allowing improved time resolution using evolutionary stages within mammalian lineages (e.g. numerical ages). Such periods corresponds to almost continuous sedimentary processes, with only possible short breaks. On the other side, there are periods with unreported data, shared in the paleokarst and peripheral basins (Late Oligocene as best example). Such periods may well correspond to some non-deposition episods as once alleged by geologists. Then the surrounding biochronologic data would bear constraining value regarding these episods.     

Biostratigraphie,paléoécologie et zones d’acmé des foraminifères planctoniques au passage Crétacé-Paléogène dans la Téthys (Tunisie et Espagne) et l’Atlantique (France)

Biostratigraphical high-resolution analyses and quantitative data confirm that deposition is continuous across the K-Pg transition in several sections in Tunisia (El Kef, stratotype section) and Spain (Agost and Caravaca sections) located in the Tethyan realm and the Bidart sections in the Atlantic realm, without any relevant hiatus. The Upper Maastrichtian assemblages of planktic foraminifera from these sections are largely dominated by small biserial heterohelicids. They are associated to common species having planispiral test (i.e. globigerinelloids), trochospiral test (i.e. hedbergellids, rugoglobigerinids globotruncanids), to rare triserial heterohelicids (i.e. guembelitriids) and trochospiral species showing tubulospines (i.e. schackoinids). Stratigraphical ranges of these diverse taxa through the late Maastrichtian in the Tethyan and Atlantic realms show very few changes in the planktonic foraminiferal assemblages and most of the species are present in the Abathomphalus mayaroensis biozone. By our high-resolution sampling and the intensive research for the A. mayaroensis index species in the uppermost Maastrichtian samples, we confirm that this species is omnipresent up to the top of the Maastrichtian. Therefore, A. mayaroensis is present in almost all samples which are late Maastrichtian in age, but this species became very scarce in the uppermost Maastrichtian samples. This scarceness could be due to a climate cooling. A sharp decrease in relative abundance of the deep dwellers species, like as Abathomphalus intermedius and A. mayaroensis as well as in other keeled globotruncanids is observed at the studied sections from the Tethyan realm (indicative of low latitude) across the latest Maastrichtian. At the K/Pg boundary, all the globotruncanids disappeared. They are considered specialists living in tropical-subtropical deep seawater habitat. At this boundary, large and ornate heterohelicids also disappeared. Therefore, all the studied sections show that about 90 % of the Maastrichtian species became extinct according to a catastrophic mass extinction pattern. Only about 10 % crossed the K/Pg boundary and survived during the earliest Danian. The minor difference in the number of disappeared taxa is related to their latitude location or environment paleodepth. The changes in the species relative abundance, observed in the successive planktic foraminiferal assemblages, make it possible to recognize the Acme-stage 0 typical of the upper Maastrichtian interval. It is characterized by the highest species richness of Globotruncanids and heterohelicids specialists of tropical to subtropical marine conditions, the Acme-stage 1 typical of the Guembelitria cretacea Zone, and in particular of the Hedbergella holmdelensis Subzone dominated by “opportunists” species belonging to Guembelitria, the Acme-stage 2 which corresponds to the Pv. eugubina Zone dominated mainly by specimens belonging to Palaeoglobigerina and Parvularugoglobigerina and the Acme-stage 3 which characterizes mainly the Ps. pseudobulloides Zone dominated by biserial species belonging to Chiloguembelina and Woodringina.     

Les ostracodes du Néogène supérieur du littoral nord-ouest marocain entre Tanger et Asilah

Ostracodes populations have been studied in 19 samples from the upper Neogene of the Atlantic coast of NW of Morocco, between Tangier and Asilah. One hundred and two species belonging to 53 genera have been indentified. Analysis of the species assemblages allowed to reconstruct the palaeoenvironmental evolution of these deposits. This begins with the transgression of the upper Tortonian which corresponds to a relatively deep circalittoral environment. The series continues by deposits showing a general regressive character, punctuated by few transgressives pulses. Shallowing starts at the end of the upper Tortonian with the deposition of calcarenites in massive banks, in an infralittoral exposed environment. This facies pass towards the top to a detrital facies, assigned to the Messinian and deposited under high energy conditions in a shallow-marine environment. The lower Pliocene transgression shows a more generalized character. The maximum deepening corresponds to the grey clay facies deposited in the external infralittoral environment, relatively protected and rich in phytal biotopes. The more recent deposits are shallower, from coastal environments, and show a more limited extension.     

Stratigraphie, paléoécologie et foraminifères du paléogène des Alpes Maritimes et des Alpes de Haute-Provence (Sud-Est de la France)

Paleogene sedimentation in the Alpes Maritimes and Alpes de Haute Provence regions is composed of one complete sedimentary sequence. This sequence begins with the continental Formation from Microcodium and continues with the Cerithium layers, the Calcaires Nummulitiques and the Marnes Bleues, which are overlaid by the turbidites of the Grès d’Annot Formation. Sedimentation starts in the Nummulites perforatus zone close to the base of ‘the Truncorotaloides rohri zone (P 14) and ends in the Cassigerinella chipolensis-Pseudohastigerina micra zone (P 18) and the NP21 zone in the upper part of the Grès d’Annot. More biomarkers were used in order to define a more detailed local biozonation (biozones AMP 1 to AMP 7). Four local zones were also defined by the last occurrence of Nummulites millecaput and N. perforatus-N. ptukhiani and then by the first appearance of N. retiatus (AMGF 1-4). The evolution of benthic foraminiferal assemblages shows an increase in bathymetry from the internal platform in the Cerithium layers to the calcareous platform with large foraminifers, then to the external platform and the deep offshore environment of the Marnes Bleues and Grès d’Annot. Low faunal diversity in the Grès d’Annot together with the predominance of agglutinated species indicates a poorly oxygenated, organic rich and turbidite environment. Seven steps (SD 1 to SD 7) in the Eocene marine transgression are shown from East to West by detailed local biozonation and sequence analysis. Grès d’Annot sedimentation is also diachronous, beginning within the Truncorotaloides rohri zone (P 14, AMP 1) in the East and ending within the Pararotalia opima opima zone (P 20, middle Rupelian) in the West (Barrême). Small foraminifer Paleogene fauna from the Alpes Maritimes and Alpes de Haute Provence was studied from 400 samples. It is composed of 378 species. Two new taxa are proposed: Fissurina niceana n. sp. and Globocassidulina alpina n. sp. The species from the Escarène and Gorbio neighborhood described by M. von Hantken (1884) were re-examined.     

Mise en évidence d’une flore oxfordienne dans le Sud-Est de la Tunisie : intérêts stratigraphique et paléoécologique

The Late Jurassic - Early Cretaceous series encountered in South-East Tunisia, formely considered globally as Purbecko-Wealdian in age, has the most complete palaeobotanical record of all North-Gondwana. Thanks to the stratigraphical results of the last years and to new field researches, it was possible, for the first time, to evidence an Oxfordian flora at the base of this series (Bir Miteur Formation). This flora, constituted of impressions and axes of Pteridophytes and Gymnosperms, is encountered within different types of plant-bearing levels. The study of deposit facies and sequential analysis of this Oxfordian formation allowed the identification of corresponding sedimentary environments and to infer their palaeoecological meaning. The discovery of exceptional specimens provided new informations about the general morpho-anatomy of Alstaettia, a very peculiar tree fern of a now extinct type. This Oxfordian flora is an interesting key-point in the evolution of North-Gondwanian flora at the Jurassic/Cretaceous transition.     

La stratigraphie et le Paléolithique du complexe saalien dans la région de la Saale et de l’Elbe

Some sediment sequences were known in the Elbe-Saale region. They enable a subdivision of the Saale complex, that is from Neumark Nord (Geisel valley), from Weimar-Ehringsdorf and Bad Kösen-Lengefeld (Saale valley). According to it, there are two interglacials between the Saalian groundmoraine (s.str.) and the Eemian. They are characterized by a strong subcontinental climatic influence, which could not be observed in the other interglacials of the Elbe-Saale region. The interglacials are connected with find horizons from the Middle Palaeolithic. They are concisely described here with their inventories.     

Néandertal et le feu au Paléolithique moyen ancien. Tour d’horizon des traces de son utilisation dans le Nord de la France

This article aims at drawing up balance sheet of remains of fire use by the first Neanderthals of Northern France, during the second part of the Saalian (MIS 8 to 6). This overview reminds us the rarity of fire testimonies during Early Middle Palaeolithic (300–130 ky BP) on the scale of North-Western Europe. For Northern France, only the sites of Biache-Saint-Vaast and Therdonne present remains of combustion. At Biache-Saint-Vaast, it is not less than six levels, which present clues of combustion: burnt flint and faunal remains and sometimes charcoals. At Therdonne, besides burnt numerous flint and some rare faunal remains were brought to light during the excavation of level N3 several rich zones in organic residues and micro-charcoals. All the datas collected concerning the clues of combustion at Biache-Saint-Vaast and Therdonne is compiled, analyzed and interpreted. This approach permits to establish the fire use or its absence in saalian occupations of Neanderthals of Northern France and to discuss modalities of its use, particularly at Therdonne. To conclude, fire status and its implications in first Neanderthals occupations are briefly discussed.     

Stratégie de débitage et mode de façonnage des industries du Paléolithique inférieur en Chine et en Europe entre 1 Ma et 400 000 ans: Ressemblances et différences de la culture de l’homme de Yunxian et Acheuléen européen

These stone tools from Yunxian with other Lower Paleolithic sites in South China, for example, the site Lantian (Shanxi, China) and the site Bose (Guangxi, China), indicates that there are many common characteristics among these industries. There are choppers, chopping-tools, picks, hand-axes, cleavers and scrapers. Long ago we thought that there are not hand-axes in China. In fact, there are lots of hand-axes in China and these hand-axes existed more earlier than those in Europe. The hand-axe appeared in China before 800,000 years old and it barely arrived in Europe before 600,000 years old. The quaternary deposits of the site of Yunxian Hominid (Hubei, China) are at least dated to 800,000 years old. A large quantity of lower Paleolithic material was discovered. The types of the tools are choppers, picks, scrapers, hand-axes and cleavers. The pebbles tools make up the majority of the implements. Small flake tools are relatively rare. The lithic industries comparison of the two regions has help us to know the cultural evolution in China and Europe.     

Découverte de charophytes de l’Albien dans la Formation Kebar (Tunisie centrale) : implications paléoécologiques et paléobiogéographiques

In Tunisia, in the platform domain, the Aptian-Albian transition is associated with a major stratigraphic gap not precisely dated, which is related to the subaerial exposure of the Aptian Orbata platform. Locally, in central Tunisia (Jebel Kebar), this gap is substituted by the sedimentary records of the Kebar Formation. This formation provided a rich association of fossil charophytes, formed by Atopochara trivolvis trivolvis, Clavator harrisii zavialensis and Sphaerochara verticillata which allows attributing it to the late Aptian–early Albian interval (most probably early Albian). In addition, a new variety, Sphaerochara verticillata var. kebariensis, is described. The charophyte assemblage shows significant biogeographic affinities with a coeval flora found in the Iberian Peninsula. The Kebar Formation was previously regarded as exclusively nonmarine, but the presence of the assemblage of charophytes and associated ostracods described rather indicates a fluctuation between margino-littoral (lagoonal) and continental environments.     

Les ammonites du Bathonien-Callovien du Boulonnais : biodiversité, biostratigraphie et biogéographie

The ammonite fauna of the outcropping Bathonian-Callovian of the Boulonnais (Northern France) is described in detail for the first time. In the middle and Late Bathonian, the fauna is very restricted, reduced to some Perisphinctidae (Procerites) and Clydoniceratidae (Clydoniceras, Delecticeras). The abundance and the biodiversity of assemblages increase during the Callovian. These faunas are typically Subboreal, marked by Gowericeratinae (Kepplerites, Sigaloceras), Kosmoceratinae (Kosmoceras) and Proplanulitinae (Proplanulites), associated with Boreal taxa such as Cadoceratinae (Cadoceras) and Cardioceratidae (Quenstedtoceras), and Submediterranean taxa, Pseudoperisphinctinae (Homoeoplanulites, Poculisphinctes), Peltoceratinae (Pseudopeltoceras) and Macrocephalitinae (Macrocephalites). Representing the only Jurassic onshore outcrops, between the English and Normandy coasts, and the Ardennes area, the interest of these faunas for correlations between the Western and Eastern European (Russian) platforms is emphasized. Palaeobiogeographically, particularly concerning the southward migration/dispersion of the Boreal taxa, the role played by the transgressions/regressions and the resulting paleobathymetric and palaeoclimatic implications is discussed.     

Évolution des caractères technologiques et typologiques des industries lithiques dans la stratigraphie de la Caune de l'Arago

The thick Quaternary deposits of the Caune de l’Arago (Pyrénées-Orientales, France) are dated to between 690 000 and 90 000 years old. At least fifteen different archeostratigraphical units have been identified within these deposits, each corresponding to distinct prehistoric occupations. Numerous stone tools made from several different rock types, have been discovered in each unit. The tools present specific characteristics concerning the choice of raw materials, the typology, and the technology used to produce them. Morpho-technological study of the different components of the assemblage contributes to a better understanding of the debitage methods used for their production. Each raw material is considered individually in order to ascertain its origin in the environment, its typological role and the technology applied during its exploitation. Defining production systems leads to the characterisation of the assemblages from each unit. When compared, they reveal common elements, as well as differences, suggesting evolutionary trends. Some observations are also made concerning the extent to which changing uses of the site may have influenced the general morphology of each assemblage, therefore taking into account exterior factors. Analysis of this rich stone tool assemblage helps to situate the Caune de l’Arago industry within the larger evolutionary context of the Lower Paleolithic in Mediterranean Europe.     

Ostracodes de la Formation de Bhanness,Kimméridgien (Jurassique supérieur) du Liban central : taxonomie,paléoécologie et paléobiogéographie

A microfauna composed of 11 species of ostracodes belonging to 11 genera have been recovered from the Bhanness Formation of Kimmeridgian age, in Central Lebanon. Four species are new and are described herein: Ektyphocythere dahressawanensis nov. sp., Dicrorygma (Orthorygma) libanensis nov. sp., Citrella? elongata nov. sp. and Procytherura inflata nov. sp. From a palaeobiogeographical standpoint, this fauna characterizes the North Gondwana province which differs from the South Gondwana province, particularly by the presence of the genus Schuleridea and the absence of the genus Majungaella.