Primary homology assessment of structures in the female atrial system among species of the Polycystididae (Rhabditophora, Eukalyptorhynchia)

Abstract. The female atrial system of members of the Polycystididae has been studied at the light microscopical level and compared among the constituent taxa. Based on the criteria of position and conjunction, hypotheses of homology are put forward and compared with the assessments of homology found in earlier literature. Contradictory terminology is synonymized and adapted to fit new findings that recognize homology among structures in different taxa. Based on differences in position and structure, 2 types of female duct are recognized: female duct type I and female duct type II. The term “female bursa” is restricted to a sperm resorbing organ at the proximal end of the female duct type I and/or at the end of a common oviduct. Some species have bundles of glands at the place where the oviduct(s) enter the female duct type I, which are considered homologous among these species. Different types of seminal receptacles are recognized. The term “insemination duct” is restricted to the ducts which in some species, in addition to the oviducts, connect the ovaries with the female duct type I. A single muscular duct, the common oviduct, connects the joined oviducts with the female duct type I in species of Duplacrorhynchus and is considered homologous with the similar duct present in some other species. A uterus is always present, entering the common genital atrium through its frontal wall, or entering a female duct type II. This morphological review of the female atrial system reveals a mosaic distribution of characters if applied to previous phylogenies that are based primarily on the male reproductive system.     

Revision of Trigonostomum Schmidt, 1852 (Platyhelminthes, Typhloplanoida, Trigonostomidae) with the description of seven new species

A morphological and taxonomic account of the genus Trigonostomum is provided. All known species are discussed and briefly re-described where necessary. Seven new species are described: T. franki from Curaçao, Florida (USA), the East African Coast and New Caledonia, T. nataschae from the French sub-Antarctic island Kerguelen, T. spinigerum from New Caledonia and T. watsoni from the Australian East Coast and New Caledonia. T. tori and T. galapagoensis , both formerly enclosed in T. setigerum , are considered new species, while T. australis also belongs to the same species group. Proxenetes denhartogi is transferred to Trigonostomum . Based on a comparison of old and new material, T. marki is synonymized with T. penicillatum , while T. prytherchi and T. divae are regarded as junior synonyms of T. lilliei ; T. intermedium and T. quadrifolium are considered synonyms of T. coronatum . Three species are considered species inquirendae : T. brunchorsti , T. piriforme and Marinellia lingulifera . Similarities and differences of the 17 valid species are discussed and summarized in an identification key.  © 2004 The Linnean Society of London, Zoological Journal of the Linnean Society , 2004, 141 , 271–296.     

A cladistic re-assessment of the Polycystis species complex (Polycystididae,Eukalyptorhynchia)

The taxonomy of the genus Polycystis Kölliker, 1845 sensu Schockaert &; Karling, 1975 is discussed. Character analysis reveals the monophyletic state of the three species groups these authors defined within it. As no synapomorphies for the three groups can be found, the genus is split in three genera: Polycystis Kölliker, 1845, Macrorhynchus Graff, 1882 and Paulodora Marcus, 1948. The systematic position of Macrorhynchus yaganus (Marcus, 1954) Evdonin, 1977 is also discussed, resulting in the erection of the new genus Marcusia gen. n.     

Primary Homology Assessment in the Male Atrial System of the Polycystididae (Platyhelminthes: Eukalyptorhynchia)

A comparative lightmicroscopical study of the male atrial system of the Polycystididae is presented. Firstly, the different glandular structures associated with the male atrial system are compared. This has lead to the recognition of eight different types of glandular vesicles, four of which are considered prostate, the other four accessory. Secondly, the different hard structures that can be found in the male atrium are discussed and compared. These hard structures can consist of numerous hard spines (armed cirrus), but mostly the hard structure is a single- or double-walled tubiform stylet, or a plate-shaped stylet. Different types of stylets can be recognised, some of which are associated with a prostate vesicle and therefore considered prostate stylets. Using the criteria of position and conjunction, conjectures of homologies are put forward. These primary homologies are compared with homology assessments found in earlier literature.     

Revision of the European species of the genus Trachelipus Budde-Lund, 1908 (Crustacea: Isopoda: Oniscidea)

The European species of Trachelipus Budde-Lund, 1908 are revised. The generic diagnosis is extended. The species T. aegaeus (Verhoeff, 1907), T. arcuatus (Budde-Lund, 1885), T. ater (Budde-Lund, 1896), T. camerani (Tua, 1900), T. dijfkilis (Radu, 1950), T. nodulosus (C. L. Koch, 1838), T. palustris (Strouhal, 1936), T. rathkii (Brandt, 1833), T. ratzeburgii (Brandt, 1833), T. razzautii (Arcangeli, 1913), T. rhinoceros (Budde-Lund, 1885), T. squamuliger (Verhoeff, 1907), T. trilobatus (Stein, 1859), (77 troglobius Tabacaru & Boghean, 1989) and 7T vespertilio (Budde-Lund, 1896) are recognized as valid. Except for T. troglobius , redesriptions, figures and distribution maps are given. A key to species is provided. Nineteen names are new synonyms of the above mentioned ones; nine names become nomina dubia.     

Two new species of the genus Gieysztoria (Rhabdocoela: Dalyelliidae) from China

Two new turbellarian species, Gieysztoria shantouensis Zhang, Li Wang, sp. nov. and G. huizhouensis Zhang, Wu Wang, sp. nov. are described, which were collected from ditches in Shantou and Dongjiang River, Huizhou of Guangdong Province, respectively. G. macrovariata(Weise, 1942) was firstly recorded in China. All specimens examined were deposited in National Zoological Museum, Institute of Zoology, Chinese Academy of Sciences, China.     

Revision of the lygaeid genus Nysius (Heteroptera: Lygaeidae: Orsillinae) of Japan,with description of a new species

Japanese species of the lygaeid–orsilline genus Nysius were revised, and five species including a new species, N. hidakai, were recognized. Male and female genitalia of all the species were illustrated, and a key to species of Japanese Nysius was provided. The DNA barcoding information of each species except N. expressus were recorded.     

Callionymus omanensis,a new species of dragonet from Oman,north‐western Indian Ocean (Teleostei: Callionymidae)

A new species of deep‐living dragonet Callionymus omanensis from Oman is described on the basis of a single male specimen collected in a trawl from 500 m depth off the coast of Oman. The new species is characterized within the subgenus Bathycallionymus by having a small branchial opening; head short (3·9 in proportion to standard length); eye large (2·4 in proportion to head length); preopercular spine with a long, upcurved main tip, with a small antrorse barb and a larger antrorse spine, and with a strong antrorse spine laterally at the preopercular‐spine base, ventral margin smooth; first dorsal fin slightly higher than second dorsal fin (male); second dorsal fin distally straight; 17 pectoral fin rays; distal end of caudal fin slightly pointed, with two median unbranched rays bearing short filaments; first dorsal fin with basal black spot reaching from first to fourth membranes, third membrane with an ocellated distal black blotch; second dorsal fin with vertical dark grey bars; distal three‐fourths of anal fin black; upper half of caudal fin with oblique dark grey bars; pelvic fin dark grey, second ray basally with a black blotch. The new species is compared with similar species. Revised keys to callionymid species of the western Indian Ocean and the Red Sea, as well as species of the subgenus Bathycallionymus, are presented.     

Notes on some corydiid species from China,with the description of a new genus (Blattodea: Corydioidea: Corydiidae)

This article presents the diversity of cockroach family Corydiidae from China. Epipolyphaga wukong Qiu, Che et Wang, n. gen., n. sp., is described. Habitus and detailed characters of the new taxa are illustrated. A comparison between the new genus and the morphologically similar genus Eupolyphaga Chopard is given. The genus Tivia Walker and the species Eucorydia westwoodi (Gerstaecker) are newly recorded from China. The Corydioidea chapter in Cockroaches of Southeastern China by Liu et al. is revised. A key to genera and a checklist of Chinese Corydiidae are provided.     

Revision of fossil species of Deinodryinus, with description of a new species (Hymenoptera, Dryinidae)

Deinodryinus veltenisp. n. is described from middle Eocene Baltic amber. The species differs from other fossil Palaearctic species of Deinodryinus Perkins owing to the shape of the antenna (clavate, with distal part very thickened), the large compound eyes, and the distal part of the stigmal vein much longer than the proximal part. A revision and a key to the fossil Palaearctic species of Deinodryinus Perkins, 1907 is presented.     

铬黄圆痕叶蝉属的订正及圆痕叶蝉族巴西二新属二新种描记（半翅目：叶蝉科）

基于比较形态学对曾归入圆痕叶蝉亚科铬黄圆痕叶蝉属Chromagallia的8个有效种（C. saucia (St?l), C. flavofasciata (St?l), C. longistilata (Coelho & Dutra), C. carvalhoi Gon?alves et al., C. lamasi Gon?alves et al., C. lanceolata Gon?alves et al., C. paraguayensis Gon?alves et al.和C. zanolae Gon?alves et al.）进行了订正,明确了该属的范围仅限于具有黄斑的3个种（C. flavofasciata (St?l), C. longistilata (Coelho & Dutra) 和C. rodriguesoi sp. nov.）,对该属进行了重新描述,并修订了鉴别特征。此外,对模式种C. flavofasciata 进行了重新描记,首次提供了C. longistilata的雌性生殖器图,并作了描记。把曾归入铬黄圆痕叶蝉属Chromagallia具有红斑的6个种移出并新建了2个新属：Rubragallia 和Neorubragallia,其中Rubragallia 包括R. saucia (St?l) n. comb.和R. paraguayensis (Gon?alves et al.) n. comb., Neorubragallia包括N. lamasi (Gon?alves et al.) n. comb., N. lanceolata (Gon?alves et al.) n. comb., N. zanolae (Gon?alves et al.) n. comb., N. carvalhoi (Gon?alves et al.) n. comb. 和N. mervini sp. nov.。文中提供了3个属的分种检索表,并对不同种的分类地位及3个属的划分进行了讨论。     

First record of Isocapnia (Plecoptera: Capniidae) from China with description of a new species

Summary

The winter stonefly genus Isocapnia Banks, 1938??? is mainly distributed in western North America and eastern Asia, but has never been reported from China. In this paper, we report, describe and illustrate a new species of this genus, I. anguis n. sp. from Sichuan Province of China. This is the first record for the genus Isocapnia from China, extending the distribution of Isocapnia into the southwestern China. An identification key to all the Isocapnia species from Asia is provided.???     

脊颜茎蜂属评述暨一新种(膜翅目:茎蜂科)

基于模式标本和新材料研究了茎蜂科脊颜茎蜂属,修订了该属属征和主要鉴别特征.该属唇基上区具锐利中脊,下唇须第4节短,明显宽于下颚须第4节,左上额粗短,中部显著弯折,雄虫多数鞭分节腹侧钝齿状突出,雌虫部分锯刃具多列亚基齿等特征,与茎蜂科其余各属均不相同.描述了脊颜茎蜂属1新种,采集自江西和湖南南部山区.编制了种检索表.     

Review of the Chinese species of the genus Varma Distant (Hemiptera,Fulgoromorpha, Tropiduchidae), with description of two new species

Two new species of Varma Distant, 1906, V. falcata Chang & Chen, sp. n. (China: Guizhou) and V. lobata Chang & Chen, sp. n. (China: Guizhou) are described and illustrated. The female genitalia of four speices including two known species are described and illustrated for the first time. The diagnostic characters of this genus are redefined. A checklist to the species of Varma in China is given. The Keys on male and female genitalia to the Chinese species of Varma are provided.     

Systematic revision of Myristicaceae (Magnoliales) in Madagascar, with four new species of Mauloutchia

The systematics of Malagasy Myristicaceae is revised to take into account new collections made since the work by Capuron in the 1970s, as well as improved knowledge from fieldwork and two scanning electron microscopy studies by the author. Four new species are described: Mauloutchia annickiae Sauquet, M. capuronii Sauquet, M. echinocarpa Capuron ex Sauquet, and M. sambiranensis (Capuron) Sauquet. In addition, basic information is given for each remaining Malagasy species of the family (synonymy, type specimen, updated distribution and main distinctive features). According to this treatment, Malagasy Myristicaceae now consist of four endemic genera and 15 species: Brochoneura Warb. (three species), Doyleanthus Sauquet (one species), Haematodendron Capuron (one species) and Mauloutchia (Baill.) Warb. (ten species). Two identification keys to these species are provided: one based primarily on fruit characters and one based primarily on male flower and inflorescence characters. Putative phylogenetic relationships among these species are also indicated, based on a previous combined morphological and molecular study by the author.  © 2004 The Linnean Society of London, Botanical Journal of the Linnean Society , 2004, 146 , 351–368.     

龙州蝗属分类研究及一新种记述（直翅目：斑腿蝗科）

系统研究了龙州蝗属,共记述10种,其中包括1新种,即贵州龙州蝗Longzhouacris guizhouensis sp.nov.,该新种近似于斑角龙州蝗L.annulicornis Lu,LiYou,2000,主要区别为:1)前胸背板沟前区长为沟后区长的2.16倍;2)前翅到达第3腹节背板中部;3)后足跗节爪间中垫长为爪长的1.5倍;4)雄性尾片三角形,顶尖;5)触角基半黄褐色,端半暗褐色;6)前翅前半黑褐色,后半淡褐色;7)后足胫节黄绿色。文中附有分种检索表。新种的模式标本保存于陕西师范大学动物研究所昆虫标本室。     

Phylogeny and systematics of the Acrapex apicestriata (Bethune-Baker, 1911) species complex (Lepidoptera,Noctuidae, Noctuinae,Apameini, Sesamiina) with the description of eight new species from the Afrotropics

Twelve morphologically similar species of Acrapex Hampson 1894, (Lepidoptera, Noctuidae, Noctuinae, Apameini, Sesamiina), from Western, Central and Eastern Africa are reviewed. Eight of these species are new to science and are described: Acrapex akunamatata n. sp. and A. incrassata n. sp. from Kenya; A. gracilis n. sp., A. iringa n. sp., A. lukumbura n. sp. and A. rungwe n. sp. from Tanzania; A. soyema n. sp. from Ethiopia; and A. zoutoi n. sp. from Benin. All 12 species belong to a species complex that we hereby define as the Acrapex apicestriata group. Host-plants for three of the new species are recorded: Setaria incrassata (Hochst.) Hack. for Acrapex incrassata; Cymbopogon pospishilii (K. Schum.) C.E. Hubb. for A. rungwe; and Andropogon perligulatus Stapf. for A. zoutoi. We also conducted molecular phylogenetic analyses (using maximum likelihood and Bayesian inference) on a six gene multimarker molecular dataset (four mitochondrial and two nuclear gene fragments; 4581 nucleotides in length) consisting of 15 Acrapex species (including seven species from the apicestriata group) and four outgroups species from the subtribe Sesamiina (from genera Busseola Thurau 1904, Sciomesa Tams & Bowden 1953, Pirateolea Moyal, Le Ru, Conlong, Cugala, Defabachew, Matama-Kauma, Pallangyo & Van den Berg 2010 and Sesamia Boisduval & Guenée 1852). Both maximum likelihood and Bayesian inference analyses yield a similar and well-supported topology, which supports the monophyly of the apicestriata group.