气候变化和不同强度造林对大兴安岭主要树种林分信息和地上生物量的长期影响

气候变化及相应火干扰在不同尺度上影响着我国大兴安岭地区森林动态,且在未来的影响可能继续加剧。为了提高森林生态功能和应对气候变暖,国家在分类经营基础上全面实施抚育采伐和补植造林,效果较好,但抚育采伐对森林主要树种的长期影响知之甚少,其在未来气候下的可持续性也有待进一步评估,同时,探讨造林措施对未来森林的影响也显得尤为重要。本文运用森林景观模型LANDIS PRO,模拟气候变化及火干扰、采伐和造林对大兴安岭地区主要树种的长期影响。结果表明:1)模型初始化、短期和长期模拟结果均得到了有效验证,模拟结果与森林调查数据之间无显著性差异(P0.05),基于火烧迹地数据的林火干扰验证亦能够反映当前火干扰的效果,模型模拟结果的可信度较高;2)与当前气候相比,气候变暖及火干扰明显改变了树种组成、年龄结构和地上生物量,B1气候下研究区森林基本上以针叶树种为主要树种,A2气候下优势树种向阔叶树转变;3)与无采伐预案相比,当前气候下,抚育采伐使落叶松的林分密度和地上生物量分别降低了(165±94.9)株/hm~2和(8.5±5.1) Mg/hm~2,增加了樟子松、白桦和云杉等树木株数和地上生物量(3.3—753.4株/hm~2和0.2—4.0 Mg/hm~2),而对山杨的影响较小;B1和A2气候下抚育采伐显著改变林分密度,降低景观尺度地上生物量,进而表现为不可持续;4)B1气候下,推荐实施中低强度造林预案(10%和20%强度),在A2气候下,各强度造林均可在模拟后期增加树种地上生物量。     

树种水平的大兴安岭地上生物量变化特征及其与气候和干扰的关系

树种水平地上生物量(每个树种的地上生物量)是衡量森林生态系统结构功能的重要指标。为揭示树种水平森林地上生物量变化机制及其与气候变化和干扰的关系,运用KNN (k-nearest neighbor distance)方法将森林调查数据和MODIS数据相结合,估算了黑龙江大兴安岭2000、2010和2015年树种水平的森林地上生物量,在此基础上运用典型对应分析和随机森林方法,分析了研究区树种水平地上生物量变化特征及其与气候和干扰因素的关系。研究结果表明:2000—2015年期间,研究区总的森林地上生物量增加了8.9%(0.41×10~8 t),其中2010—2015年期间地上生物量的增加速度要明显高于2000—2010年;地上生物量增加最多的树种为白桦(Betula platyphylla Suk.),与2000年相比生物量增加了0.40×10~8 t,其次为樟子松(Pinus sylvestris var.mongolica Litv.)、山杨(Populus davidiana Dode)和蒙古栎(Quercus mongolica Fisch. ex Ledeb.),落叶松(Larix gmelinii(Rupr.) Kuzen)地上生物量下降了0.08×10~8 t,柳树(Chosenia arbutifolia(Pall.) A. Skv.)和云杉(Picea koraiensis Nakai)基本上没有变化;林火、采伐和造林等森林干扰均对树种水平地上生物量影响显著,林火对树种水平地上生物量的影响要高于造林和采伐;气候要素显示出了比干扰要素更为重要的作用,多年平均温度和降水解释了最多的树种水平地上生物量变异。年均温度与阔叶树种的生物量以及林火干扰有显著的正相关性,与总的森林地上生物量呈现出显著的负相关,与落叶松和白桦表现出微弱的负相关,预示着气候变暖将影响该区域的树种组成并降低该区域的森林生产力。     

林火干扰对大兴安岭主要林分类型地上生物量预测的影响模拟研究

林火干扰是北方森林最主要的自然干扰之一,对北方森林地上生物量影响是一个长期的过程。因此,在预测地上生物量动态变化时需要考虑林火的影响。运用空间直观景观模型LANDIS PRO,模拟大兴安岭林区林火对不同树种地上生物量预测的影响。选取研究区5种主要树种林分(兴安落叶松、樟子松、云杉、白桦和山杨),以无干扰情景为参考预案,在验证模型模拟结果的基础上,模拟林火在短期(0—50a)、中期(50—150a)和长期(150—300a)对地上生物量的定量化影响,及其对不同立地类型地上生物量的动态变化。结果表明:(1)基于森林调查数据参数化的2000年森林景观模拟结果能够较好地代表2000年真实森林景观,模拟的2010年森林林分密度和胸高断面积与2010年森林调查数据无显著性差异(P0.05),当前林火干扰机制模拟结果能够较好地与样地调查数据匹配,说明林火模拟能够代表当前研究区林火发生情况;(2)与无干扰预案相比,整个模拟时期内景观水平上林火减少了1.7—5.9 t/hm2地上生物量;(3)与无干扰预案相比,林火预案下主要树种生物量在短期、中期和长期变化显著(P0.05);(4)在不同模拟时期,林火显著地改变了地上生物量空间分布,其中以亚高山区地上生物量降低最为明显。研究可为长期森林管理以及森林可持续发展提供参考。     

采伐强度对长白山森林地上生物量和景观格局的长期影响

采伐是驱动长白山森林景观变化的关键因子。本研究采用空间直观森林景观模型(LANDIS PRO 7.0)模拟长白山露水河林业局在5个不同采伐强度方案下的森林地上生物量和景观格局的长期动态变化。结果表明:1)采伐导致了树种产生不同程度的景观破碎化;2)采伐强度对森林地上生物量具有显著影响,采伐强度增大,地上生物量减小;采伐同样显著降低了不同树种的地上生物量,其中采伐对水曲柳、椴树和云杉的影响较大。因此,在模拟的前100年(2003—2103年),当采伐强度较高时,应优先采伐白桦和山杨,然后是水曲柳、云杉和椴树;当采伐强度较低时,水曲柳、云杉、椴树,白桦和山杨都可作为采伐树种。在模拟的后100年(2103—2203年),由于森林地上生物量呈现减小的变化趋势,应适当减小采伐强度,水曲柳、云杉、椴树,白桦和山杨都可作为采伐树种,从而为当地森林管理部门制定合理的管理措施提供科学的依据。     

呼中林业局森林采伐方式对森林景观格局的长期影响

采用空间直观景观模型(LANDIS)模拟了大兴安岭地区呼中林业局在无采伐和同一采伐量下皆伐、二次渐伐和择伐共4种方案下森林景观300年的长期动态变化。结果表明,与无采伐方案相比较,采伐使落叶松和云杉的分布面积减小,樟子松、白桦的分布面积增大;择伐对本区分布面积最大的2个树种———落叶松和白桦的影响最小。采伐改变了树种年龄结构,降低了过熟林的相对面积;相对于皆伐而言,渐伐和择伐时降低的程度较大。各种采伐方式间落叶松和云杉的聚集度差异较小,白桦和樟子松的聚集度则存在显著差异。     

模拟分类经营对小兴安岭林区森林生物量的影响

运用空间直观景观模型LANDIS 7.0 PRO,模拟了在当前采伐模式和无采伐两个预案下,小兴安岭林区森林生物量及主要树种生物量在2000—2200年间的动态。模拟结果如下:(1)无采伐预案下,森林生物量由最初的93.6 t/hm2逐渐升高,90a后达到最大值258 t/hm2,之后森林生物量在245 t/hm2上下小幅波动;(2)前100a采伐预案会明显降低森林生物量,与无采伐预案相比森林生物量最大可降低21.4 t/hm2,平均减少14.7 t/hm2;后100a采伐对森林生物量的影响逐渐减弱,森林生物量平均减少2.6 t/hm2;(3)当前采伐模式促进保护树种红松和紫椴生长,其生物量分别最大可提高9.0 t/hm2和0.53 t/hm2,占到无采伐预案生物量的56%和15%;(4)采伐预案对云冷杉生物量影响较小,主要降低先锋树种(白桦、山杨)和一些阔叶树种(枫桦、春榆)的生物量。研究结果表明现行采伐模式在未来100 a内会显著影响森林生物量,之后其影响逐渐减小,并且保护政策能提高所保护树种(红松、紫椴)的生物量,但要保持较高的总生物量,仍需要降低目前的采伐强度。     

树种组成和径级结构对热带次生林生物量恢复影响的研究

本研究在海南尖峰岭和吊罗山热带林区海拔245～1255 m范围内根据林分恢复时间设立固定监测样地,探讨了森林地上生物量与树种组成和径级结构的关系.结果显示:海南热带次生林平均地上生物量为(155.38±37.16)×103 kg/hm2,其中低地次生雨林为(137.91±31.02)×103 kg/hm2,山地次生雨林...     

模拟西伯利亚红松种植对大兴安岭中北部森林地上生物量的长期影响

西伯利亚红松是寒温带森林重要建群树种，其在大兴安岭引种种植将直接影响区域森林生态系统的结构和固碳功能。分析引种种植西伯利亚红松对大兴安岭森林地上生物量的长期影响，对厘清区域碳汇及碳中和有重要意义。本研究采用野外调查数据和森林景观模型相结合的方法，量化景观尺度西伯利亚红松不同种植强度对大兴安岭中北部森林及本地主要树种地上生物量动态影响。结果表明：引种种植西伯利亚红松对提高大兴安岭中北部林区森林地上生物量起积极作用，未来可在该区域大面积引种种植西伯利亚红松；不同种植强度预案下森林地上生物量变化趋势相似，均提高了森林地上生物量，其中低强度种植预案下森林地上生物量增幅最大，达8.66 Mg·hm^-2;西伯利亚红松引种种植对本地主要树种的影响存在明显差异，兴安落叶松和云杉地上生物量升高0.19～8.88 Mg·hm^-2,樟子松、山杨和白桦地上生物量降低0.42～12.86 Mg·hm^-2。     

川西亚高山不同森林生态系统碳氮储量及其分配格局

森林采伐和恢复是影响森林碳氮储量的重要因素。以川西亚高山岷江冷杉原始林、粗枝云杉阔叶林、天然次生林和粗枝云杉人工林为研究对象,采用样地调查和生物量实测的方法,研究了不同森林生态系统各组分碳、氮储量及其分配特征。结果表明岷江冷杉原始林、粗枝云杉阔叶林、天然次生林和粗枝云杉人工林生态系统碳储量分别为611.18、252.31、363.07 tC/hm~2和239.06 tC/hm~2;氮储量分别为16.44、12.11、15.48 tN/hm~2和8.92 tN/hm~2。恢复林分与原始林碳储量在土壤—植被的分配格局发生了变化,而氮储量未发生变化。岷江冷杉原始林以植被碳储量为主,恢复林分以土壤为主,氮储量均以土壤为主。乔木层碳储量分别占生态系统总储量的56.65%、17.63%、13.57%和22.05%,土壤层(0—80 cm)分别占32.03%、69.87%、76.20%和72.12%;土壤层氮储量占生态系统总储量的76.80%—92.58%。植物残体碳氮储量分别占生态系统总储量的4.40%—9.83%和2.94%—7.08%,林下植被所占比例最小。空间格局上,岷江冷杉原始林植被部分具有较高的碳储量,应进行保护。3种恢复林分具有较高的碳汇潜力,且地上/地下碳储量较低,表明其碳汇潜力尤其表现在地上部分。天然次生林利于土壤有机碳的积累,而人工林乔木层碳储量较高。     

白龙江上游地区森林植被恢复能力的分析

通过对残存林分的天然更新能力、天然次生群落和人工恢复群落的生长状况调查,分析了白龙江上游地区森林植被的恢复能力。结果表明：在人工恢复植被的过程中,营造于海拔3000m以上的人工云杉(Picea asperata)幼林生长不良,没有成林地分布,而营造于海拔2700～3000m带的人工云杉幼林生长发育良好,并且已成林地的林分结构符合理想同龄林林分结构的分布规律,能够定向培育成森林资源;残存天然林分的更新表现为草类云杉林天然更新良好,残存箭竹云杉林、杜鹃岷江冷杉林(Abies faxoniana)和箭竹岷江冷杉林的天然更新不良;采伐迹地上天然次生群落生长良好,更新及时,正处于正向演替阶段。根据影响林分生长的主要因子(海拔、土壤厚度、坡度、坡向)和林分的实际恢复能力,将该地区划分为3个森林植被恢复类型区,并分别提出了相应的恢复措施。     

Projected climate change effects on Alberta's boreal forests imply future challenges for oil sands reclamation

Climate change will drive significant changes in vegetation cover and also impact efforts to restore ecosystems that have been disturbed by human activities. Bitumen mining in the Alberta oil sands region of western Canada requires reclamation to “equivalent land capability,” implying establishment of vegetation similar to undisturbed boreal ecosystems. However, there is consensus that this region will be exposed to relatively severe climate warming, causing increased occurrence of drought and wildfire, which threaten the persistence of both natural and reclaimed ecosystems. We used a landscape model, LANDIS‐II, to simulate plant responses to climate change and disturbances, forecasting changes to boreal forests within the oil sands region. Under the most severe climate forcing scenarios (representative concentration pathway [RCP] 8.5) the model projected substantial decreases in forest biomass, with the future forest being dominated by drought‐ and fire‐tolerant species characteristic of parkland or prairie ecosystems. In contrast, less extreme climate forcing scenarios (RCPs 2.6 and 4.5) had relatively minor effects on forest composition and biomass with boreal conifers continuing to dominate the landscape. If the climate continues to change along a trajectory similar to those simulated by climate models for the RCP 8.5 forcing scenario, current reclamation goals to reestablish spruce‐dominated boreal forest will likely be difficult to achieve. Results from scenario modeling studies such as ours, and continued monitoring of change in the boreal forest, will help inform reclamation practices, which could include establishment of species better adapted to warmer and drier conditions.     

Carbon and nitrogen cycling in North American boreal forests

A model of boreal forest dynamics was adapted to examine the factors controlling carbon and nitrogen cycling in the boreal forests of interior Alaska. Empirical relationships were used to simulate decomposition and nitrogen availability as a function of either substrate quality, the soil thermal regime, or their interactive effects. Test comparisons included black spruce forests growing on permafrost soils and black spruce, birch, and white spruce forests growing on permafrost-free soils. For each forest, simulated above-ground tree biomass, basal area, density, litterfall, moss biomass, and forest floor mass, turnover, thickness, and nitrogen concentration were compared to observed data. No one decay equation simulated forests entirely consistent with observed data, but over the range of upland forest types in interior Alaska, the equation that combined the effects of litter quality and the soil thermal regime simulated forests that were most consistent with observed data. For black spruce growing on permafrost soils, long-term simulated forest dynamics in the absence of fire resulted in unproductive forests with a thick forest floor and low nitrogen mineralization. Fires were an important means to interrupt this sequence and to restart forest succession.     

The Origin and Dynamics of Subalpine White Spruce and Balsam Fir Stands in Boreal Eastern North America

Associations among the few tree species in the North American boreal landscape are the result of complex interactions between climate, biota, and historical disturbances during the Holocene. The closed-crown boreal forest of eastern North America is subdivided into two ecological regions having distinct tree species associations; the balsam fir zone and the black spruce zone, south and north of 49°N, respectively. Subalpine old-growth stands dominated by trees species typical of the balsam fir forest flora (either balsam fir or white spruce) are found on high plateaus, some of which are isolated within the black spruce zone. Here we identified the ecological processes responsible for the distinct forest associations in the subalpine belt across the eastern boreal landscape. Extensive radiocarbon dating, species composition, and size structure analyses indicated contrasted origin and dynamics of the subalpine forests between the two ecological regions. In the black spruce zone, the subalpine belt is a mosaic of post-fire white spruce or balsam fir stands coexisting at similar elevation on the high plateaus. With increasing time without wildfire, the subalpine forests become structurally similar to the balsam fir forest of the fir zone. These results concur with the hypothesis that the subalpine forests of this area are protected remnants of an historical northern expansion of the fir zone. Its replacement by the fire-prone black spruce forest flora was caused by recurrent fires. In the subalpine belt of the fir zone, no fire was recorded for several millennia. Harsh climate at high altitude is the primary factor explaining white spruce dominance over balsam fir forming a distinct subalpine white spruce belt above the balsam fir dominated forest.     

Spore deposition of wood-decaying fungi: importance of landscape composition

The abundance of many species of wood-decaying fungi has decreased dramatically in Swedish boreal forests over the last century. Therefore, we have investigated the relationship between the spore dispersal of wood-decaying fungi and two key features of landscape composition, namely the amount and age of old forest stands at different spatial scales. Spore deposition was monitored in two regions, using sampling methods based on recording the dikaryotisation of monokaryotic mycelia on nutrient agar and wood discs. The studied species, all mainly confined to Norway spruce in the boreal forest zone, were Fomitopsis pinicola, Fomitopsis rosea , Gloeoporus taxicola , Phlebia centrifuga and Trichaptum laricinum . The study of forestry intensity showed for both regions, that the spore deposition for F. pinicola , F. rosea and G. taxicola was higher in circular plots (radius 2-km) with a high proportion of old Norway spruce forest (>80 yr) than in plots with a lower proportion of old forest. Analysis of the variation in spore deposition of F. rosea and P. centrifuga in relation to the proportion of old spruce forest within 1, 2 and 3 km of the spore sampling point showed that the proportion of old forest within a 3-km radius explained more of the variation than the proportions within 1- and 2-km radii. In addition, the proportion of forest older than 140 yr explained more of the variation than the proportion of younger forests. Thus, the results show that the spore deposition of the studied species strongly depends on the landscape composition at both regional and local scales. Further, the spore deposition at the local scale was best explained by the proportion of >140 yr old spruce forest, which exceeds the common harvest rotation period.     

气候变化、林火和营林措施对寒温带典型森林生态弹性的影响

生态弹性是森林生态系统在遭受外在扰动后恢复到稳定状态的能力,是森林资源可持续发展的重要目标之一,且森林生态弹性对诸如气候变化、林火和营林措施等外部因子的影响较为敏感.探究这些外部因子对森林生态弹性的影响在未来森林生态系统管理方面有重要意义.本研究首先从森林组成、结构和功能等方面选取指标因子并估算了森林生态弹性值,然后运用LANDIS PRO模型,模拟气候变化、林火干扰和营林措施等对寒温带典型森林生态弹性的影响,并探讨了当前抚育采伐方案在未来气候下的可持续性.结果表明: 模型初始化的2000年林分密度和胸高断面积与2000年真实景观较为吻合,模拟的2010年森林景观与野外调查数据无明显差异,基于当前林火干扰状况的模拟结果与火烧迹地调查数据基本匹配,说明林火模块能很好地模拟当前研究区林火发生状况.林火干扰增加30%将会使该区模拟期内景观水平上森林生态弹性提高15.7%～40.8%,而林火干扰增加200%则会降低该区4.4%～24.6%的森林生态弹性.短期和中期林火干扰增加对森林生态弹性的影响大于气候变化的影响.与当前预案相比,B1气候(林火增加30%预案)和A2气候(林火增加200%预案)对整个模拟阶段景观尺度森林生态弹性的影响分别处于-15.9%～38.9%和-60.4%～34.8%范围内.与无采伐预案相比,B1和A2气候下在整个模拟时期内若继续实施当前抚育采伐方案,将不利于景观水平森林生态弹性的提高.在B1气候(林火增加30%预案)下,在各模拟时期内无需实施任何营林措施;而在A2气候(林火增加200%预案)下,建议实施中、高强度种植的营林措施以提升景观水平森林生态弹性.     

气候变化、林火和采伐对大兴安岭森林碳储量的影响

大兴安岭林区林火发生的频率受气候变化的影响将会增加,可能会增加该地区森林生态系统碳损失.本研究通过耦合森林生态系统模型和森林景观模型以模拟未来百年大兴安岭森林碳储量动态变化,量化气候变化、林火和采伐对森林碳储量的影响.结果表明: 虽然采伐和林火会抵消相当一部分由气候变化增加的碳储量,但气候变化仍然能够增加大兴安岭森林碳储量.未来100年该地区森林地上和土壤有机碳储量将会分别增加9%～22%和6%～9%.短期(0～20年)气候变化对大兴安岭森林碳储量的影响大于同期林火的影响,中期(30～50年)和长期(60～100年)气候变化对森林碳储量的影响小于林火和采伐的影响.由于未来大兴安岭地区气候变化及其林火干扰存在不确定性,导致未来该地区森林碳储量存在较大的不确定性.未来100年大兴安岭森林地上碳储量和土壤有机碳储量不确定性分别为12.4%～16.2%和6.6%～10.4%.为准确估算我国北方森林生态系统碳储量,需要考虑种子传播、林火和采伐的影响.     

Damaged forests provide an opportunity to mitigate climate change

British Columbia (BC) forests are estimated to have become a net carbon source in recent years due to tree death and decay caused primarily by mountain pine beetle (MPB) and related post‐harvest slash burning practices. BC forest biomass has also become a major source of wood pellets, exported primarily for bioenergy to Europe, although the sustainability and net carbon emissions of forest bioenergy in general are the subject of current debate. We simulated the temporal carbon balance of BC wood pellets against different reference scenarios for forests affected by MPB in the interior BC timber harvesting area using the Carbon Budget Model of the Canadian Forest Sector (CBM‐CFS3). We evaluated the carbon dynamics for different insect‐mortality levels, at the stand‐ and landscape level, taking into account carbon storage in the ecosystem, wood products and fossil fuel displacement. Our results indicate that current harvesting practices, in which slash is burnt and only sawdust used for pellet production, require between 20–25 years for beetle‐impacted pine and 37–39 years for spruce‐dominated systems to reach pre‐harvest carbon levels (i.e. break‐even) at the stand‐level. Using pellets made from logging slash to replace coal creates immediate net carbon benefits to the atmosphere of 17–21 tonnes C ha^?1, shortening these break‐even times by 9–20 years and resulting in an instant carbon break‐even level on stands most severely impacted by the beetle. Harvesting pine dominated sites for timber while using slash for bioenergy was also found to be more carbon beneficial than a protection reference scenario on both stand‐ and landscape level. However, harvesting stands exclusively for bioenergy resulted in a net carbon source unless the system contained a high proportion of dead trees (>85%). Systems with higher proportions of living trees provide a greater climate change mitigation if used for long lived wood products.