东海陆架西南部台北坳陷古近纪孢粉植物群的时空分布

详细叙述东海陆架西南部台北坳陷古近纪不同凹陷区各时期孢粉植物群展布情况。区内古近纪孢粉植物群可分为早古新世、晚古新世、早始新世和中始新世4个发育阶段,早古新世孢粉植物群仅见于坳陷西南的局部地区,称Abietineaepollenities/Cedripites-Parcisporites-Momipites leffingwellii组合,晚古新世孢粉植物群分4种类型,椒江凹陷称Taxodiaceaepollenites-Engelhardtioidites-Momipites leffingwellii组合,丽水-福州凹陷称Taxodiacenepollenites-Myricaceoipollenites-Momipites leffingwellii组合,闽江凹陷南部称Taxodiaceaepollenites-Momipites leffingwellii组合,闽江册陷北部称Lygodiumsporites-Cicatricosisporites dorogensis-Tiliaepollenites-Bombacacidites组合。早始新世孢粉植物群具南北差异,北部以海金沙科和椴科孢粉植物发育为特征,南部以海金沙科、椴科和杨海科孢粉植物群发育为特征,以上均指示为湿热的亚热带气候。中始新世本区孢粉植物群比较一致,称Cicatricosisporties dorogensis-Ephedripites(D)-Nitrariadites组合,指示为受到干旱气候影响的亚热带气候。区内缺失渐新统,故无渐新世孢粉记录。     

中国早第三纪孢粉植物群纲要

简要地叙述中国大陆及陆架区早第三纪孢粉植物群的纵向演变和横向分异，将纵向演变归纳为６个主要的孢粉植物群发展时期。即：１）早古新世榆科花粉发育期；２）晚古新世正型粉扩展期；３）早始新世榆科－桦科－胡桃科花粉繁盛期；４）中始新世壳外科花粉发育期；５）晚始新世蒺藜科花粉扩展期；６）渐新世松科花粉发育期。据横向地区差异，将中国大陆及陆架区划分为７个早第三纪孢粉植物地理区，为：１）东北暖温带至亚热带湿生孢粉植物区，２）华北暖温带至亚热带中生孢粉植物区，３）西北暖温带至亚热带旱生孢粉植物区，４）中南亚热带中生孢粉植物区，５）东海南部陆架亚热带湿生孢粉植物区，６）南方亚热带、热带中生至湿生孢粉植物区，７）西藏亚热带中生孢粉植物区。对以上各期的孢粉植物区的主要特征和各地理区范围，代表性的孢粉和特征分别进行叙述，并以此对中国早第三纪的主要含孢粉地层列表进行了对比。     

东海陆架西南部台北坳陷新近纪孢粉植物群演替

对东海陆架西南部台北坳陷13口钻井孢粉分析表明,本地区新近纪植物群经历了3个发展阶段,即：1)早中新世早期松科花粉优势期;2)早、中中新世阿丁枫科花粉繁盛期;3)晚中新世至早上新世草本被子植物发展期。各时期孢粉植物群在横向上几乎没有大的区别,表明新近纪整个坳陷内构造与沉积环境及其变迁具有很好的一致性。植物群演替反映古气候从早中新世早期的湿润温凉到早、中中新世的暖湿,最后(晚中新世至早上新世)又趋温凉的变化历程。     

藏南江孜地区古近纪甲查拉组孢粉组合及古环境分析

甲查拉组位于藏南特提斯喜马拉雅北带的江孜盆地,是江孜地区的最高海相地层,被视为新特提斯洋关闭时代的主要证据。通过对甲查拉组剖面的孢粉分析,自下而上划分为3个孢粉组合带:1 Ulmipollenites minor+Ulmoideipites-Myricipites组合带;2 Quercoidites-Caryapollenites组合带;3 Abiespollenites-Betulaceoipollenites+Betulaepollenites组合带。甲查拉组孢粉植物群揭示出其沉积自晚古新世开始,其沉积结束可能延续到早渐新世。推测晚古新世—早始新世为温暖湿润的暖温带或亚热带气候,始新世中晚期为温暖、潮湿的亚热带气候,早渐新世为温和湿润的暖温带气候。     

东海陆架盆地西南部始新世孢粉

通过对东海陆架盆地西南部台北拗陷始新统之瓯江组、温州组和平湖组的孢子和花粉的研究,本区始新统可分为两个孢粉组合:Taxodiaceaepollenites-Caryapollenites triangulus-Aquilapollenites spinulosus组合(早始新世)和Retitricolpites-Retitricolporites组合,后者又分为两个亚组合,分别代表中始新世和中始新世晚期或/至晚始新世早期的孢粉植物群。孢粉组合反映早始新世的植物群主要由杉科、胡桃科和完斗科组成,指示暧湿的、近现今的中南亚热带气候;中晚始新世孢粉植物群具有明显的现代性,并有较多南亚热带成份加入,指示更趋暖热的气候环境,其后半期有较多指示干旱气候的分子出现,表明曾一度受到干旱气候的影响。     

拉萨地区林周盆地设兴组上部(古近系)年代研究的新进展及意义

拉萨地区林周盆地典中那玛剖面设兴组上部新发现的孢粉组合以落叶、阔叶植物为主体,主要为桦科的Alnipollenites,Betulaepollenites,Carpiniptes,山毛榉科的Quercoidites,胡桃科的Juglanspollenites,榆科的Ulmipollenites,椴科的Tiliapollenites等,孢粉化石多为古近纪常见分子,未发现白垩纪的特征分子。古近纪早期大量出现的三孔沟,网面三孔沟等花粉少量出现;孢粉组合更接近于古近纪中晚期的面貌,其时代可能属于晚始新世。因此,设兴组上部的年代可能是始新世晚期。由此推论,设兴组和林子宗群之间的角度不整合不能代表白垩纪古近纪之间的构造运动,而是代表始新世晚期后的构造运动。同时,林周盆地可能沉积有古近纪的河湖相地层。     

山东青东凹陷西部古近纪孢粉组合特征及意义

通过对山东青东凹陷西部古近纪地层孢粉研究,共建立4个孢粉组合:Ⅰ.Ephedripites-Inaperturopollenites-Ulmipollenites minor,Ⅱ.Pinaceae-Ulmoideipites tricostatus-Taxodiaceaepollenites,Ⅲ.QuercoiditeshenriciTaxodiaceaepollenites hiatus,Ⅳ.Alnipollenites-Juglanspollenites-Polypodiaceaes porites。根据与渤海湾地区的孢粉资料对比,确定4个组合的时代分别为早始新世晚期—中始新世早期、中始新世早—中期、中始新世中—晚期、中始新世晚期—晚始新世。该区古近纪植被类型由落叶、阔叶和灌木混交林逐步转变为落叶、常绿阔叶和针叶混交林,古气候经历了干热的亚热带气候到湿热的亚热带气候再到亚热带型温暖气候的变化。沙四段上部到沙三段下部沉积时期是研究区烃源岩最为发育时期。     

云南吕合地区晚第三纪孢粉植物群

吕合地区晚第三纪孢粉植物群研究表明该植物群属北极第三纪植物区系 ,孢粉组合以被子植物为主 ,温带成分如Alnus、Betula、Carpinus、Corylus、Quercus、Castanea、Ulmus等 ,亚热带、热带成分如Cyclobalanopsis、Castanopsis、Liquidambar、Carya、Davidia、Ilex、Palmae等。吕合附近海拔较高的地带分布松科Abies、Tsuga、Pinus等针叶树种。吕合孢粉组合中温带成分比热带、亚热带成分花粉类型多 ,每种类型的花粉含量也多。植被类型为常绿落叶阔叶混交林 ,反映温暖湿润的亚热带气候类型。植物群对比揭示吕合植物群时代为中新世中晚期_上新世早期。     

西藏新生代秋乌组孢粉化石的发现及其初步研究

本文首次报道西藏雅鲁藏布江缝合带附近秋乌组含煤沉积中的孢粉化石。孢粉植物群主要由Quercoidifes,具孔类及三孔沟类花粉组成,包括12科26属及一些未定类群。组合中产有少量的Graminidites及Tsugaepollenites。植物群的总体特征显示其时代可能为渐新世至早中新世。秋乌组沉积早期的环境比较温暖。     

滇东南普阳盆地含煤地层孢粉组合特征及其年代

云南地处青藏高原东南缘, 生物多样性丰富, 季风气候特征明显。新生代剧烈的构造活动形成了众多山间盆地, 其间保存了大量精美的植物化石, 是探讨新生代以来植被、植物多样性和地球环境演变的理想地区。但是长期以来, 由于地层年代学证据的缺乏, 这些新生代沉积盆地的地质年代还存在很大争议。本研究通过对滇东南地区富宁县普阳盆地的含煤地层开展深入的孢粉学研究, 探讨盆地含煤地层年代及其古气候演化过程。剖面下部煤层孢粉组合以杉粉属(Taxodiaceaepollenites)为主, 指示以杉科为主的湿润沼泽森林, 冬春季相对湿润; 剖面上部孢粉组合指示以常绿栎类为主的亚热带常绿‒落叶阔叶混交林, 生长山核桃粉属(Caryapollenites)、冬青粉属(Ilexpollenites)、胡桃粉属(Juglanspollenites)、枫香粉属(Liquidambarpollenites)等亚热带常见树种, 气候温暖湿润, 季节分明。结合新发现的哺乳动物化石证据和周边地区不同地质时代的孢粉组合, 普阳盆地含煤地层的沉积时代应为晚始新世; 同时, 孢粉组合也表明滇东南地区植被现代化面貌至少在晚始新世就已经开始出现。     

云南元谋盆地上新世甘棠组植物和孢粉组合及其古气候意义

云南元谋盆地湾堡甘棠组植物群的叶化石和孢粉化石合计107种类型,分属86属,47科。植物叶化石包括18科,24属,35种。除一种裸子植物外,其它均为被子植物。被子植物中,榆科、桦木科、豆科无论属种或者化石数量都是最多;杨柳科、槭科、蔷薇科次之;壳斗科、樟科、杨梅科、鼠李科、忍冬科、胡颓子科、杜鹃科、香蒲料、禾本科均有代表。甘棠组的孢粉植物群也相当丰富,合计97个类型,分属72属,44科。以松科、榆科、禾本科及蕨类植物的水龙骨科最为丰富,胡桃科、壳斗科、金缕梅科及蕨类植物的凤尾蕨属也很常见。被子植物花粉占孢粉总数的50%--60%,以禾本科和榆属花粉含量最高,多在10%以上。裸子植物绝大部分为松科花粉,占孢粉总数的20%-30%。孢子以水龙骨科和凤尾蕨属为主,二者各占孢粉总数的5%-10%。发现于元谋盆地湾堡甘棠组的植物叶化石以及孢子花粉组合在成分上都是混合的类群。组合中既有相当多的湿润亚热带常绿阔叶林成分,也有耐干旱的胡颓子、杨梅、旱蕨等灌木、小乔木或草本分子,同时还有含量很高的、在半干旱环境下也能生存的禾本科、榆科花粉。据此推测甘棠组的植物群和孢粉组合反映了上新世时元谋地区的植被已经分化,盆地的周围山地生长有松科、壳斗科、樟科、胡桃科、金缕梅科为主的常绿和落叶阔叶树木或林块,可能有少许杜鹃科小乔木混杂其中,属于地带性植被。而在盆地内则是以禾本科、蔷薇科等为主,杂以大戟科、豆科、忍冬科、杨梅科等灌木或草本植物以及榆科、桦木科等树木组成的稀树灌丛草原型植被。盆地内的植被所反映的气候和周边地区相比,显得明显干旱。因此推测元谋盆地在2-3百万年前的中晚上新世已经发育成为金沙江流域的干热河谷盆地之一。     

The first Loranthaceae fossils from Africa

An ongoing re-investigation of the early Miocene Saldanha Bay (South Africa) palynoflora, using combined light and scanning electron microscopy (single grain method), is revealing several pollen types new to the African fossil record. One of the elements identified is Loranthaceae pollen. These grains represent the first and only fossil record of Loranthaceae in Africa. The fossil pollen grains resemble those produced by the core Lorantheae and are comparable to recent Asian as well as some African taxa/lineages. Molecular and fossil signals indicate that Loranthaceae dispersed into Africa via Asia sometime during the Eocene. The present host range of African Loranthaceae and the composition of the palynoflora suggest that the fossil had a range of potential host taxa to parasitise during the early Miocene in the Saldanha Bay region.     

Palynoflora of the Liushagang formation (Eocene-Early Oligocene) in the northern part of South China Sea

In the northern part of South China Sea, including Tonkin Gulf, Hainan lsland, Leizhon Peninsula and some basin of Guangdong Province, Paleogene deposits are composed of three formations: the Weizhon, the Liushagang and the Changliu formations arranged in descending order. The paper on the palynoflora of the Weizhon Formation (early and middle Oligocene) is in press^[2]. This paper deals only with the palynoflora of the Liushagang Formation, with may be divided into four main stages: The first stage is represented by Monocolpollenites tranquillus and Crassoretitriletes sp., assigned to early Eocene The second stage is characterized by Salixipollenites, Momipites triletipollenites and Operculumpollis. Its age is middle Eocene. The third stage is dominated by some species of Quercoidites and Ulmipollenites and also characterized by the presence of Platycaryapollenites and Prominangularia dogyingensis, This sporo-pollen assemblage suggests a late Eocene in age. The fouth stage is marked by profusion of some alga of brockish water, such as Rugasphera corrugia, Granodiscus gronulatus and some pollen types of Liquidambarpollenites minutus, Multiporopollenites puctatus and Tricolporopollenites minutus. The age of the last stage is assigned to early oliocene. In generaly, the palynoflora of the Liushagang Formation is quite different from that of the Weizhou Formation. The main types of spores and pollen are common with those found in Europe and North America of the same age, while the Weizhou Formation has many elements common both in this region and Borneo. During Eocene and early Oligocene this area was of continental phase with brackish basins. At the beginning the climate was rather moist and hot, but then itbecame moist and warmtemperate.     

Arctic plant diversity in the Early Eocene greenhouse

For the majority of the Early Caenozoic, a remarkable expanse of humid, mesothermal to temperate forests spread across Northern Polar regions that now contain specialized plant and animal communities adapted to life in extreme environments. Little is known on the taxonomic diversity of Arctic floras during greenhouse periods of the Caenozoic. We show for the first time that plant richness in the globally warm Early Eocene (approx. 55-52 Myr) in the Canadian High Arctic (76° N) is comparable with that approximately 3500 km further south at mid-latitudes in the US western interior (44-47° N). Arctic Eocene pollen floras are most comparable in richness with today's forests in the southeastern United States, some 5000 km further south of the Arctic. Nearly half of the Eocene, Arctic plant taxa are endemic and the richness of pollen floras implies significant patchiness to the vegetation type and clear regional richness of angiosperms. The reduced latitudinal diversity gradient in Early Eocene North American plant species demonstrates that extreme photoperiod in the Arctic did not limit taxonomic diversity of plants.     

四川被子植物区系特征的初步研究

四川地区被子植物区系188科1493属8711种及1252变种,全球50％以上的科都有代表,与毗邻地区及世界植物区系有着广泛而密切的联系。区系古老而原始,珍稀孑遗类群丰富;单型或少种的科属众多,含100种以上的大科和大属也十分繁荣;含5种以下的属占76.89%,但占总属数约22.64%的多种属和大属是区系的主体,包含区系总种数的63.28%。科的分布类型有明显热带性质,属、种的分布类型反映出典型的亚热带属性。优势现象明显,特有类群和表征类群丰富。该地区被子植物区系对中国乃至东亚都有代表性。     

Combined LM and SEM study of the middle Miocene (Sarmatian) palynoflora from the Lavanttal Basin,Austria: part IV. Magnoliophyta 2 – Fagales to Rosales

An ongoing investigation of the middle Miocene (Sarmatian) palynoflora from the Lavanttal Basin continues to show that it contains an extremely rich assemblage of angiosperm taxa. The Fagales to Rosales pollen record documented here contains 34 different taxa belonging to the Betulaceae (Alnus, Betula, Carpinus, Corylus, Ostrya), Fagaceae (Castanea, Fagus, Quercus Groups Cerris, Ilex, Cyclobalanopsis, Quercus/Lobatae), Juglandaceae (Engelhardioideae, Carya, Juglans, Pterocarya), Myricaceae (Morrella vel Myrica), Cannabaceae (Celtis), Elaeagnaceae (Elaeagnus), Rhamnaceae, Rosaceae (Prunus) and Ulmaceae (Cedrelospermum, Ulmus, Zelkova). Two of the pollen types represent extinct genera, Trigonobalanopsis and Cedrelospermum, and are also reported for the first time from the Lavanttal Basin along with pollen of Rhamnaceae and Prunus. The different types of Quercus pollen are now affiliated with Groups Cerris, Cyclobalanopsis, Ilex and Quercus/Lobatae based on sculpturing elements observed using scanning electron microscopy (SEM). Köppen signatures of potential modern analogues of the fossil Fagales and Rosales suggest a subtropical (Cfa, Cwa) climate at lower elevation and subsequent subtropical to temperate climate with altitudinal succession (Cfa → Cfb/Dfa→ Dfb; Cwa → Cwb → Dwb) in the Lavanttal area during accumulation of the palynoflora. Most of the fossil taxa have potential modern analogues that can be grouped as nemoral and/or merido-nemoral vegetation elements, and the diversity of Fagales indicates a varying landscape with a high variety of niches.     

EXTINCT TRANSITIONAL FAGACEAE FROM THE OLIGOCENE AND THEIR PHYLOGENETIC IMPLICATIONS

Fruits, catkins, and associated leaves of at least two extinct trigonobalanoid taxa have been discovered at an Oligocene fossil plant locality rich in fagaceous remains. These fossils exhibit a mosaic of fruit and pollen characters found in the two extant subfamilies Castaneoideae and Fagoideae of Fagaceae. Comparison with cladograms based on modern taxa suggests that these extinct taxa were similar to the ancestors of subfamily Fagoideae and may have been intermediate between Fagus and the modern trigonobalanoid genera. Pollen types isolated from the fossil staminate catkins provide unique character states that are transitional between modern pollen types in Fagaceae and are important in understanding the evolution of exine micromorphology within the family. This analysis provides a striking example of the use of character data from fossils to determine character-state adjacency prior to polarization of characters using outgroup comparison. Because of the mosaic nature of their character complexes, these fossils support monophyly in both the family Fagaceae and the subfamily Fagoideae. In addition, the occurrence of trigonobalanoid fossils in the Oligocene of North America has interesting biogeographic implications and provides insights into the nature of North American Fagaceae during the Tertiary.     

江汉盆地渔洋组上部的古新世孢粉组合

20世纪70至80年代,几乎所有关于江汉盆地跑马岗组和渔洋组的古生物文献都将其时代确定为晚白垩世。李伟同在深入地研究了跑马岗组标准剖面上的轮藻化石后提出跑马岗组大部分属于古新统。取自层位与跑马岗组相当的渔洋组上段上部的岩芯样品经分析后获得了丰富的孢粉化石。这一组合总的面貌与王大宁等获得的晚白垩世渔洋组的第三组合区别明显,而与古近纪的新沟嘴组的第四组合相似处很多。在该孢粉植物群中,常见于当地的晚白垩世分子,如希指蕨孢属、克拉梭粉属、江汉粉属、刺参粉属、鹰粉属等没有或极少见到。而在本组合常见的南岭粉属、五边粉属、漆树粉属等多出现在我国南方古新世地层中。因此,其时代很有可能为古新世。如果是这样,渔洋组上部有可能跨越了白垩系和古近系界线。从当前孢粉植物群含有很多亚热带成分以及麻黄粉属以及胡颓子粉属、忍冬粉属等看,它反映了江汉盆地当时偏干旱的亚热带气候。