Population dynamics of perch, Perca fluviatilis L. in Slapton Ley, Devon

Perch, Percafluviatilis L. were studied in Slapton Ley which is a eutrophic freshwater lagoon in south Devon, England. The perch were caught in traps from February 1970 to June 1971. The traps selected males during the spawning season. The water temperature did not affect the numbers of trapped fish directly, but the sight of perch in the traps appeared to attract others. Sex ratios were found to be equal in the first 2 years of life but older year classes were predominantly female. The population consisted of young individuals. Males were found to mature in their first year and most females matured in their second. The fecundity varied from an average of 1000 eggs in perch n i the 9.75 cm length group to 30,500 eggs in fish in the 27.75 cm group. A few shoals of perch moved throughout the Ley although the majority of fish stayed in the same area. Population estimates showed a range from 10,000 to 95,000 for fish 9.75 cm and above. The large variations in population estimates and survival rates were attributed to the necessity of using traps which had variable catches and to the harmful effects of handling the perch. Chironomids were important in the diet of perch of all sizes. Plankton and Asellus were other important foods, the latter especially during the winter months. Perch 11.5 cm to 13.9 cm selected the widest range of food organisms. Only perch 14.0 cm and above ate fish.     

The relative performance of pheromone and light traps in monitoring the seasonal activity of both sexes of the eastern hemlock looper, Lambdina fiscellaria fiscellaria

The seasonal flight activity of both sexes of the eastern hemlock looper, Lambdina fiscellaria fiscellaria Guenée (Lepidoptera: Geometridae) was studied during two consecutive years in Quebec and Newfoundland, using light (L), pheromone (P) and combined light and pheromone (LP) traps. Moth density significantly affected the performance of the different traps, with P traps being more effective at low than high density. However, P trap catches decreased just prior to the onset of female captures, probably as a result of competition between traps and virgin calling females. Nearly all females caught in L and LP traps were already mated and even the first females caught had laid at least half of their egg complement. In Quebec under warm nights, the pattern of male activity occurred at different times, with peak P catches being later in the scotophase than those of L traps, but overall similar numbers of males were caught in both traps. In contrast, under cool nights, males were caught early in the night in both P and L traps, suggesting a strong competition effect between traps, although more males were caught in P than L traps overall. In Newfoundland, the pattern of male captures in L and P traps was similar at both high and low temperatures, so competition between trap types would always be high. Under these conditions P traps were more effective than L traps. Irrespective of the region, year or temperature, significantly more males were captured in LP, with the effect of L and P being additive. In both regions, females responded similarly to L and LP traps with peak activity occurring early in the night. Captures of females were lower than those of males under cool temperatures, suggesting that the temperature threshold for flight is higher for females. The use of L and P traps simultaneously and/or in combination is discussed in relation to integrated pest management programs and ecological considerations.     

Monitoring populations of the carrot fly, Psila rosae, with sticky and water traps

The relative, effectiveness of Rebell®, small cylinder, large cylinder, windmill and water traps, the five types of trap used currently for monitoring populations of the carrot fly, Psila rosae, was assessed in nine field experiments, three in south west Lancashire, four in the Fens (Suffolk, Norfolk, Cambridgeshire), one in East Suffolk and one at Wellesbourne, Warwickshire. Regression analysis of the numbers of flies caught on each type of trap against the numbers caught on the Rebell® trap indicated that each trap samples a constant proportion of the fly population relative to the other traps. Therefore, provided the fly population was sufficiently large for insects to be caught on the least effective traps, any of the five traps would monitor adequately fluctuations in carrot fly populations. However, the Rebell® trap caught 4–17 times as many flies/trap and 5–7 times as many flieshnit area of trap as any of the other traps tested.
Operators considered the Rebell® trap to be the easiest to use. However, it was more expensive than any of the other traps tested.     

A yardstick for measuring populations of small rodents

(1) Six-monthly trappings (summer and winter) were done between 1952 and 1969 of Wood mice (Apodemus sylvaticus) and Bank voles (Clethrionomys glareolus) in Wytham Woods, near Oxford. One hundred and fifty Longworth live traps were used on two 2-acre grids in oak-ash-sycamore woodland. (2) The traps were pre-baited for 48 hours and then set. The catch was examined on the evening of the same day and on the following morning. The total of animals caught during the 24 hours was used as an index to the actual numbers present. Animals retrapped in the morning (identified by a simple method of marking) were omitted from the total. (3) Between 1969 and 1972 the same procedure was used but trapping, evening and morning, was continued for 3-4more days until some 75–80% of the catch consisted of marked animals. (4) This enabled estimates to be made of the actual numbers of voles and mice present at each trapping and so to assess the proportion of the total present which was caught ring the first 24 hours. (5) These estimates were made by (Method la) a straight Lincoln Index, (Method lb) a Lincoln Index method as modified by Hayne (1949) and (Method 2) by Hayne's trap-out method. Over six trappings the estimates by these methods showed reasonable agreement. (6) Comparison of the index trapping (i.e. the numbers caught during the first 24 hours) with the estimates of the total numbers present showed that, on all occasions, more than half of the mice and voles present entered the traps during the first 24 hours. With Bank voles there was a tendency for about half of them to be trapped during this time during the winter and about three-quarters of them during the summer. (7) With Wood mice the figures are more variable but, again, never less than half were trapped during the first 24 hours and, on some occasions, nearly the whole of the population. (8) These results show that reasonably accurate estimates of the numbers of these rodents can be achieved within 3–4 days' trapping (estimates that can be cross-checked by different methods) by using a high density of traps (see (1) above). Furthermore, trapping for 24 hours only will always catch at least half of the population present and, on occasions, considerably more than half.     

Effect of hunger on yellow water trap catches of hoverfly (Diptera: Syrphidae) adults

1 An experiment was conducted in a winter wheat field using yellow water traps at crop height and at ground level, near to and distant from flowers, to test the hypothesis that such traps are seen as a source of food by flower‐feeding adult hoverflies and are therefore likely to selectively trap hungry individuals. Hoverflies caught in each trap were counted and identified and the amount of pollen in their guts was assessed. Ratios of numbers of hoverflies seen in the wheat crop to numbers caught in nearby traps were compared for the different treatments. 2 Most hoverflies were caught in crop‐high traps but they included a high proportion of individuals with empty guts. The taxa were: Episyrphus balteatus (De Geer), Metasyrphus corollae (F.) (these species together accounted for over 90% of the individuals trapped), with Sphaerophoria spp., Syrphus spp., Scaeva pyrastri (L.), Melanostomini and unidentified others making up the rest. 3 Significantly fewer hoverflies were captured in low than in high traps. The ratios of numbers trapped to numbers observed, in flower and in no flower treatments would be expected to be the same if the traps were not selective. This was supported for low traps. With high traps, however, there was a highly significant difference between the ratios (71.34 and 126 : 8, respectively). 4 Flies captured in high traps had less pollen in their gut than those captured in low ones. At each distance, more E. balteatus captured in high traps were in pollen category 1 (< 20 grains) than in any other category. The opposite state was seen in low trap catches, where most flies were in category 5 (> 5000 grains). Median pollen categories were 2 (21–200 grains) and 4 (1501–5000 grains) for flies caught in high and low traps, respectively. 5 The ecological selectivity of traps according to their height and the physiological condition of the targeted individuals is a problem likely to affect many trapping systems apart from the one described in this paper.     

Effects of habitat and food resources on morphology and ontogenetic growth trajectories in perch

Studies on resource polymorphism have mainly been considered at the end stage of ontogeny, whereas many species undergo diet changes as they grow. We conducted a field survey to analyze the role of adaptive variation during ontogeny in Eurasian perch (Perca fluviatilis). We caught perch from the littoral and pelagic zones of a lake to investigate whether perch differ in morphology and diet between these habitats. We also investigated whether there were any differences in morphological trajectories during the ontogeny of perch collected from the two habitats. We found that perch caught in the littoral habitat, independently of size, had a deeper body, larger head and mouth and longer fins than perch caught in the pelagic zone. Macroinvertebrates and fish dominated the diet of littoral perch, whereas the diet of the pelagic perch consisted mainly of zooplankton and to some extent fish. Independently of size, the more streamlined individuals had a larger proportion of zooplankton and a smaller proportion of macroinvertebrates in their diet than the deeper-bodied individuals, indicating a relation between diet and morphology. Some morphological characters followed different ontogenetic trajectories in the two habitats; e.g. the changes to a deeper body and a larger head were faster in the littoral than in the pelagic perch. The relationship between the length of perch and the size of the mouth and fins also differed between perch from the two habitats, where the increase in the length of the pelvic fin and the area of the mouth increased faster with size in the littoral perch. Our findings show that variation in morphology between habitats differs during ontogeny in a way that corresponds to functional expectations for fish species that occupy these habitats.     

Effect of trap type, trap color, trapping location, and pheromone dispenser on captures of male Palpita unionalis (Lepidoptera: Pyralidae)

Field studies were conducted to evaluate the influence of trap design, trapping location, type of pheromone dispenser, and trap color on the capture of Palpita unionalis (Hübner) males, in olive groves. The experiments were carried out in two regions, Alexandria (northern Greece) and Oropos (central Greece), where olives are cultivated. In both regions, the majority of the males (> 70% of the total) were caught from late autumn to early winter, whereas < 1% was caught during July and August. Among the trap types used, the Funnel was significantly more attractive than Delta, Pherocon 1C, and Pherocon II traps. More males were caught in traps placed at the periphery of the groves than those placed in the center. Among the four colored traps tested, white traps were the most effective. However, a significant difference in trap catches was found between white and brown traps. Traps baited with red rubber septa captured more males than those baited with the white one. The use of these parameters in monitoring and managing P. unionalis is discussed.     

The destruction of an endemic species flock: quantitative data on the decline of the haplochromine cichlids of Lake Victoria

Synopsis The Lake Victoria fish fauna included an endemic cichlid flock of more than 300 species. To boost fisheries, Nile perch (Lates sp.) was introduced into the lake in the 1950s. In the early 1980s an explosive increase of this predator was observed. Simultaneously, catches of haplochromines decreased. This paper describes the species composition of haplochromines in a research area in the Mwanza Gulf of Lake Victoria prior to the Nile perch upsurge. The decline of the haplochromines as a group and the decline of the number of species in various habitats in the Mwanza Gulf was monitored between 1979 and 1990. Of the 123+ species originally caught at a series of sampling stations ca. 80 had disappeared from the catches after 1986. In deepwater regions and in sub-littoral regions haplochromine catches decreased to virtually zero after the Nile perch boom. Haplochromines were still caught in the littoral regions where Nile perch densities were lower. However, a considerable decrease of species occurred in these regions too. It is expected that a remnant of the original haplochromine fauna will survive in the littoral region of the lake. Extrapolation of the data of the Mwanza Gulf to the entire lake would imply that approximately 200 of the 300+ endemic haplochromine species have already disappeared, or are threatened with extinction. Although fishing had an impact on the haplochromine stocks, the main cause of their decline was predation by Nile perch. The speed of decline differed between species and appeared to depend on their abundance and size, and on the degree of habitat overlap with Nile perch. Since the Nile perch upsurge, the food web of Lake Victoria has changed considerably and the total yield of the fishery has increased three to four times. Dramatic declines of native species have also been observed in other lakes as a result of the introduction of alien predators. However, such data concern less speciose communities and, in most cases, the actual process of extinction has not been monitored.     

Aphis migration and the efficiency of the trapping method

Reproducible results were obtained when adhesive traps for aphides were placed at the same height in different parts of a potato field. The total catches in different localities in different seasons can therefore justifiably be compared. During the summer dispersal flights, different species flew at different heights, so traps at one height only are not suitable for comparing the relative abundance of different species. There was less variation in numbers caught on traps at 3–4 ft. above the ground than on traps at 5–6 ft. or at ground-level.
During the first fortnight of July an average of 200 aphides, 1.6% of which were Myzus persicae , was caught on traps in the position of potato plants. Most of these were not potato aphides, but it is suggested that some of them might act as vectors of potato viruses. The number of aphides caught was roughly proportional to the volume of free air space around the traps, and as the foliage increased in density the numbers caught on lower traps decreased in proportion.
Wind-speed records within and above the crop are discussed and it is shown that conditions are often suitable for voluntary aphis flight in and near the crop on relatively windy days. Records of catches on the half-traps facing the wind and on the other halves showed that voluntary flight decreased with increasing height.
Brilliant yellow traps, coated with adhesive grease, caught more aphides than white traps, which in turn caught more than black.     

The effectiveness of traps used currently for monitoring populations of the cabbage root fly (Delia radicum)

Six different types of traps were compared for capturing adults of Delia radicum. Cone traps caught so few flies that it seems inadvisable for them to be used for monitoring populations of this pest. All the other traps tested were suitable for monitoring D. radicum populations. When expressed on a trap for trap basis, the large 1800 cm² Canadian traps caught most flies. When expressed as the numbers of flies caughthnit area of trapping surface, most flies were caught in the water traps. Similar numbers of D. radicum, D. platura, syrphids, blowflies and a tachinid, Eriothrix rufomaculatus were caught per unit area on each of three different vertical sticky traps. Although water traps have the advantage that they catch about 5 times as many females D. radicum/unit area as the most effective vertical sticky traps of a comparable size, their disadvantage is that they catch about 10 times as many beneficial syrphids. The reasons why water traps are so effective against D. radicum are discussed, with details of how to convert water-trap data to sticky-trap equivalents, and vice versa.     

Dispersal of adult caddisflies (Trichoptera) into forests alongside three New Zealand streams

The abundance and taxonomic richness of adult caddisfly faunas were determined at varing distances (up to 200 m) away from three North Island, New Zealand, streams to help define appropriate forested riparian zone widths for adult aquatic insects. Adults were collected using sticky traps and ultraviolet light traps on four occasions over summer. Light traps were more effective at catching caddisflies than sticky traps, but both methods gave similar patterns of declining abundance and taxonomic richness with distance from the stream edge. Abundances of total caddisflies at 20 m were <21% of those caught at the stream edge for both trapping techniques. The same trends were evident for abundances of most common species in light traps, whereas bimodal peaks were evident with distance from the channel edge for percentage females of three common species. More than 30% of species caught at the stream edge was found in light traps at least 70 m into forest at all sites. A faster rate of decline for abundance than richness with distance away from the stream reflected the relatively large distances travelled by representatives of many species. Similar results from the three sites for numbers and species richness indicate that the main area of activity for adult Trichoptera in forested riparian zones was within 30 m of the stream edge at these sites.     

Seasonal abundance and spatio-temporal distribution of dominant xylem fluid-feeding hemiptera in vineyards of central Texas and surrounding habitats

A survey of xylem fluid-feeding insects (Hemiptera) exhibiting potential for transmission of Xylella fastidiosa, the bacterium causing Pierce's disease of grapevine, was conducted from 2004 to 2006 in the Hill Country grape growing region of central Texas. Nineteen insect species were collected from yellow sticky traps. Among these, two leafhoppers and one spittlebug comprised 94.57% of the xylem specialists caught in this region. Homalodisca vitripennis (Germar), Graphocephala versuta (Say), and Clastoptera xanthocephala Germar trap catches varied significantly over time, with greatest counts usually recorded between May or June and August and among localities. A comparison of insect counts from traps placed inside and outside vineyards indicated that G. versuta is always more likely captured on the vegetation adjacent to the vineyard. C. xanthocephala was caught inside the vineyard during the summer. Between October and December, the natural habitat offers more suitable host plants, and insects were absent from the vineyards after the first freezes. H. vitripennis was caught in higher numbers inside the vineyards throughout the grape vegetative season. However, insects were also caught in the habitat near the affected crop throughout the year, and residual populations overwintering near vineyards were also recorded. This study shed new light on the fauna of xylem fluid-feeding insects of Texas. These results also provide critical information to vineyard managers for timely applications of insecticides before insect feeding and vectoring to susceptible grapevines.     

The Food and Feeding Chronology of Yellow Perch (Perca flavescens) in Lake Washington

The food habits of yellow perch (Perca flavescens) in Lake Washington, Seattle, U.S.A. were studied over a period of nine months. Altogether 549 yellow perch were examined for the stomach contents. The daily activity pattern of the yellow perch in Lake Washington appears to correspond roughly with the feeding pattern. The maximum numbers of perch caught by the nets during 24 hour study periods correspond well with degree of fullness of stomachs noted for the different times of the day. The food consisted mainly of cottids, mysid shrimps (Acanthomysis awatchensis) and chironomid pupae and larvae. There was no difference in the food consumed between the different sexes and between the different size groups. Maximum feeding by perch during the day was observed just before dark and the daily ration was calculated to be about 1.4% of their wet body weight.     

Prey attraction by larvae of the New Zealand glowworm, Arachnocampa luminosa (Diptera: Mycetophilidae)

Abstract. The hypothesis that bioluminescence produced by larvae of the New Zealand glow-worm, Arachnocampa luminosa , attracts prey was tested experimentally in Reserve Cave, Waitomo, New Zealand, and in its bush-clad entrance over a total of 200 days during winter, spring, and summer. We compared catches on transparent adhesive traps placed either over glowworms or over areas from which glowworms had been removed. Adhesive traps over glowworms caught significantly more invertebrates per trap per day than did control traps. Glowworms in bush attracted greater numbers and types of invertebrates than did glowworms in the cave. Diptera predominated in both bush (86% of the total catch) and cave (89%). Also caught were small numbers of Araneae, Coleoptera, Hymenoptera, Orthoptera, Trichoptera, Gastropoda, Acarina, and Neuroptera—listed in order of abundance—but no adults of A. luminosa were caught. Glowworms under adhesive traps survived with little or no food for up to 78 days.     

Some factors affecting the efficiency of water-traps for capturing cabbage root flies

Several factors influencing the efficiency of water-traps in capturing cabbage root flies were studied at Wellesbourne in 1971 and 1972. In both the laboratory and field, approximately twice as many flies were caught in fluorescent as in non-fluorescent yellow traps. Depending upon trap density, addition of a source of the attractant allylisothiocyanate (ANCS) increased the numbers of females captured by approximately twofold in fluorescent traps and from two- to sevenfold in non-fluorescent traps. Traps were equally efficient irrespective of whether the ANCS was renewed every 2, 3, 4 or 5 days. On the first day of trapping, the number of flies caught per unit area was linearly related to the square root of the number of traps in that area. On the following days the rate was probably in equilibrium with the combined effect of immigration and the rate of development of responsive flies in the trapping zone. Most males were caught 30 cm above the soil surface and most females at soil level. Traps 120 cm above the soil surface caught few flies. Populations of marked flies were released into large field cages containing both a section of hedgerow and a plot of cauliflowers. Even after a week, only 81 % of the males and 55 % of the females had been recaptured from the most responsive of these captive populations. Furthermore, only 30 % of females were recaptured when they were more than 8 days old, the age at which most probably enter the new host-crop.     

Trapping oestrid parasites of reindeer: the response of Cephenemyia trompeand Hypoderma tarandi tobaited traps

Abstract. At 340–360 km North of the Arctic Circle in Norway, Hypoderma tarandi (L.) and Cephenemyia trompe (Modeer) females were caught in baited traps from 10 July to 21 August. During three summers, adverse climatic conditions inhibited flight activity of these oestrids on 56–68% of the days. Flies were not caught prior to or after these dates, nor at winds above 8 m/s, temperatures below 10C, light intensities below 20,000 lux, or during periods of rain or snow. C0₂-baited insect flight traps caught significantly more H. tarandi females than non-baited traps. However, neither a white reindeer hide or reindeer interdigital pheromone glands enhanced the attraction of C0₂to H. tarandi or C. trompe. Hypoderma tarandi females also were attracted to mobile people, but not to stationary individuals. There were no significant differences in the numbers of C.trompe or H.tarandi caught in C0₂-baited traps in a birch/willow woods, on the treeless vidda (=tundra-like biome), or at woodsrvidda ecotone sites. Flies were caught in traps on days when the nearest reindeer herds were 25–100 km away. Significantly more H.tarandi and C.trompe were caught from 09.30 to 14.30 hours than from 14.30 to 19.30 hours; no flies were caught from 20.00 to 07.00 hours (Norwegian Standard Time = NST). Because CO_z-baited traps caught from hundreds to thousands of mosquitoes, blackflies and Culicoides midges, when climatic conditions inhibited oestrid activity, reindeer aggregations and movements attributed to insect attacks during warm sunny days may be largely in response to attacks by H.tarandi and C.trompe.     

Trap type, chirality of alpha-pinene, and geographic region affect sampling efficiency of root and lower stem insects in pine.

Root and lower stem insects cause significant damage to conifers, vector phytopathogenic fungi, and can predispose trees to bark beetle attacks. The development of effective sampling techniques is an important component in managing these cryptic insects. We tested the effects of trap type and stereochemistry of alpha-pinene, in combination with ethanol, on catches of the root colonizing weevils (Coleoptera: Curculionidae) Hylobius spp. [mostly Hylobius pales (Herbst)], and Pachylobius picivorus (Germar), the root colonizing bark beetle (Coleoptera: Scolytidae) Hylastes porculus Erickson, and the lower stem colonizing bark beetle Dendroctonus valens (LeConte). We tested for inter-regional differences by conducting similar field assays in the northern (Wisconsin) and southern (Louisiana) United States. The more effective trap type varied with region. Root weevils were caught primarily in pitfall traps in Wisconsin, whereas they were caught mostly in lower stem flight traps in Louisiana. In Wisconsin, root colonizing bark beetles were also caught primarily in pitfall traps, but lower stem colonizing bark beetles were caught primarily in lower stem flight traps. The root feeding weevils preferred (-) over (+)-alpha-pinene in both regions. Some exceptions relating to trap type or gender occurred in southern populations. The two root and lower stem colonizing bark beetles in Wisconsin showed no preference between (+) and (-)-alpha-pinene in combination with ethanol. No bark beetles were caught in the south. Our results suggest that modifying trap type and enantiomeric ratios of monoterpenes for different insect groups and in different regions can improve sampling efficiency for these important pests.     

Seasonal synchrony between pheromone trap catches of the bean bug, <Emphasis Type="Italic">Riptortus pedestris</Emphasis> (Heteroptera: Alydidae) and the timing of invasion of soybean fields

Seasonal catches of the bean bug, Riptortus pedestris (Fabricius), captured in traps containing the synthetic pheromone, were investigated under different field conditions from 2005 to 2007. In soybean fields, the number of bugs attracted to the pheromone traps increased after flowering and peaked 9–13 days after flowering. After these attraction peaks, the populations of adult bugs and nymphs increased in soybean fields. In traps located in grassland, however, only small numbers of the bugs were caught during the soybean flowering stages (from mid August to early September). The sex ratio of adults caught in the pheromone traps differed among soybean growth stages. Before flowering, more males were caught than females. After flowering, trapped females increased in number and the proportion of females exceeded 0.5 throughout the flowering periods. These results suggest that attraction to the pheromone may be affected by host plant phenology, and that females, in particular, respond strongly to the pheromone during flowering of the host plant soybean.     

Orientation of field populations ofBrachymeria intermedia [Hym.: Chalcididae], to host and host-habitat cues

Brachymeria intermedia (Nees), a pupal parasitoid of the gypsy moth, was caught in the field using plexiglas traps coated with Tangletrap (2). Significantly greater numbers of females were collected in traps baited with gypsy moth pupae than in traps with male or femaleB. intermedia or traps painted yellow.B. intermedia collected in traps during the first 3 weeks of native gypsy moth pupation were primarily female. A preponderance of males were caught during the fourth week. The hyperparasite,Brachymeria compsilurae (Cwfd) was also trapped in small numbers. Gypsy moth pupae were hung in interior and exterior areas of woodland sites. A greater number ofB. intermedia emerged from pupae in woodland edge sites than from pupae in interior woodland sites.     

Determination of an Optimal Pitfall Trap Size for Sampling Spiders in a Western Australian Jarrah Forest

Pitfall trapping is a sampling technique extensively used to sample surface foraging invertebrates for biological diversity studies and ecological monitoring. To date, very few invertebrate studies have considered what trap size is optimal for sampling spiders. This study presents preliminary findings from a single short sampling period on the role of trap size in sampling spiders in a Western Australian Jarrah forest. Four different trap diameters (4.3, 7.0, 11.1 and 17.4 cm) were examined (4 trap sizes × 15 replicates = 60 traps). Two-way ANOVAs revealed no significant interaction effects between trap size or the spatial positioning of transects within the study site along which the pitfall traps were arranged. Post-hoc tests revealed abundance, family richness and species richness increased with increasing trap sizes for traps 7.0 cm. No significant differences in these dependent variables occurred between 4.3 and 7.0 cm traps, or for species richness between 11.1 and 17.4 cm traps. Determination of an optimal trap size was undertaken by bootstrapping and calculating species accumulation curves for increasing numbers of traps used. Three different criteria were considered: equivalent number of traps (15), standardized sampling intensity (cumulative trap circumference, approximately 207 cm) and standardized cumulative handling time (approximately 1 hour 17 minutes). The largest trap size (17.4 cm) was most efficient in terms of number of traps and trap circumference. For the same number of traps, it caught 19 species whereas all other trap sizes caught ten species. At the standardized circumference, it caught seven species whereas all other trap sizes caught five. For handling time, however, the two largest trap sizes (17.4 and 11.1 cm) were optimal. Both caught nine species whereas all other traps caught 相似文献