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1.
    
Cruziana reticulata is an arthropod-related ichnospecies that is characterized by a conspicuous net-like scratch pattern whose initial formation and later preservation require the presence of consolidated substrates in shallow marine fine-grained bottoms. There are two scenarios in which epifaunal to shallow infaunal benthic organisms may access firm siliciclastic substrates: first, by exposure of the compacted bottom after erosion of the upper, water-saturated and usually soft portion of sediment column; and, second, by primary fast substrate stabilization in the absence of biogenic sediment mixing. Whereas the first mode occurs throughout the Phanerozoic, the latter is only prevalent in marine bottoms during the Early Palaeozoic mainly as a consequence of poorly developed infaunal bioturbation. However, by eradicating burrowing organisms, mass extinctions are known (i.e. the end-Permian extinction) to ‘reset’ intensity of ecospace utilization, which entails the return to this anachronistic style of trace fossil preservation in younger times. An earliest Carboniferous ichno-assemblage dominated by Cruziana reticulata from Morocco attests the spread of firm substrates in the aftermath of the Hangenberg Event – a major extinction at the end of the Devonian period. Since evidence for erosion is lacking in accompanying sedimentary rocks and the ichno-assemblage shows characteristics of opportunistic exploitation of the nutritious muddy seabed, we favour the interpretation of this assemblage to represent a post-extinction ichnofauna. It shows that other taxonomically less severe mass extinctions may also exhibit transient but severe ecological effects in open marine ecosystems such as the collapse of vital sediment mixing. Such post-extinction effects may distort the perception of the stratigraphical record as a firmground ichno-assemblage often taken as evidence for submarine erosion and are crucial in highlighting sequence boundaries. We, thus, encourage appreciating macroevolutionary framework of respective ichnofaunas.  相似文献   

2.
Limestone clasts from the beach at Marloes Sands, southwest Wales, contain slender, straight to sinuous borings cross-cut by younger, clavate borings. The former were probably produced by sipunculids or polychaetes; the latter preserve shells of the boring bivalve Gastrochaena dubia (Pennant). Unusually, the calcareous linings of the clavate bivalve borings extend into many of the slender worm borings. Such linings are considered part of the hard parts of the producing bivalve, but the chance association of the two morphologies of borings has led to the lining becoming intimately associated with both of them. The modified linings of the bivalve borings have a similar morphology to the crypt of certain clavagellid bivalves, perhaps presenting an analogue for the morphology of a pre-clavagellid, boring ancestor.  相似文献   

3.
The first arthropod trackways are described from the Middle Cambrian Burgess Shale Formation of Canada. Trace fossils, including trackways, provide a rich source of biological and ecological information, including direct evidence of behaviour not commonly available from body fossils alone. The discovery of large arthropod trackways is unique for Burgess Shale-type deposits. Trackway dimensions and the requisite number of limbs are matched with the body plan of a tegopeltid arthropod. Tegopelte, one of the rarest Burgess Shale animals, is over twice the size of all other benthic arthropods known from this locality, and only its sister taxon, Saperion, from the Lower Cambrian Chengjiang biota of China, approaches a similar size. Biomechanical trackway analysis demonstrates that tegopeltids were capable of rapidly skimming across the seafloor and, in conjunction with the identification of gut diverticulae in Tegopelte, supports previous hypotheses on the locomotory capabilities and carnivorous mode of life of such arthropods. The trackways occur in the oldest part (Kicking Horse Shale Member) of the Burgess Shale Formation, which is also known for its scarce assemblage of soft-bodied organisms, and indicate at least intermittent oxygenated bottom waters and low sedimentation rates.  相似文献   

4.
    
Random shell sections of the Pliensbachian (Early Jurassic) larger bivalve Opisoma from columns within the Main Post Office building of Ferrara, northern Italy, have been discovered to bear neat clavate-shape boreholes. These boreholes belong to the ichnogenus Gastrochaenolites Leymerie and represent bivalve borings. Opisoma is a subordinate component of the Lithiotis fauna characterised by aberrant shells thriving in tropical lagoonal settings which were widespread throughout the Tethyan and Panthalassa coasts. Although the Lithiotis fauna is well known in the palaeontological literature, no bivalve boring have been so far been formally described. The uniqueness of the morphology, size and substrate of these borings merits the designation of the new species Gastrochaenolites messisbugi ichnosp. nov. which thereby represents the first ichnospecies described from this fauna. The morphology of the boreholes and the included bivalves allows the boring activity to be ascribed to a mytilid bivalve. Palaeoecological and taphonomic analyses allowed the presence of the boreholes to be correlated to the Opisoma mode of life (epifaunal, free-living form) as well as to generally low sedimentation rates and seasonal mesotrophic conditions during an overall oligotrophic regime.

http://zoobank.org/urn:lsid:zoobank.org:act:CCF77B32-D459-4305-BC86-93F228852E50  相似文献   


5.
    
The Val Gardena Formation of the Dolomites region in northern Italy preserves the most significant assemblage of Late Permian tetrapod footprints in the world. More than 120 years of collecting resulted in about 900 publicly accessible specimens from the study area. This huge amount of data is comprehensively revised in the light of recent advances in the study of Late Palaeozoic – Early Mesozoic tetrapod ichnofossils. According to our analyses, the Val Gardena Sandstone Formation includes tracks that can be assigned to cf. Batrachichnus isp. (temnospondyl amphibian), Capitosauroides isp. (amphibian), Dicynodontipus isp. (cynodont therapsid), Dolomitipes accordii n. igen. n. comb. (dicynodont therapsid), cf. Dromopus isp. (neodiapsid), Pachypes dolomiticus (pareiasaurian parareptile), Paradoxichnium problematicum (archosauromorph neodiapsid), Procolophonichnium tirolensis n. comb. (procolophonoid parareptile), cf. Protochirotherium isp. (archosauriform neodiapsid) and Rhynchosauroides pallinii (neodiapsid). The ichnoassociation is dominated by tracks of neodiapsid and parareptilian tetrapods, whereas synapsid and anamniote tracks are rather minor components. It includes 10 out of 12 tetrapod ichnogenera known from Lopingian deposits and thus it constitutes a reference for the Paradoxichnium biochron. It shows striking similarities with other low-latitude non-aeolian contemporaneous ichnoassociations of Europe and North Africa, differences may be linked to the palaeoenvironment. Moreover, it shows a clear Triassic affinity.

The new ichnogenus Dolomitipes was registered in Zoobank.org. urn:lsid:zoobank.org:act:5B4D871C-D16A-4E93-8211-CEE08019BA60  相似文献   


6.
Erratum     
Gerhard Cadee 《Ichnos》2013,20(1):47-55
Abstract

Bored relict log-grounds are locally abundant in hybridized glauconitic fossiliferous muddy sands and associated carbonate concretions in the Eocene Tallahatta Formation, southern Alabama. Log-ground ichnofabrics are dominated by commonly stenomorphic, calcite-lined, variably filled clavate borings assigned to Teredolites longissimus and attributed to teredinid bivalves. Differences in axial morphology and occurrence of accessory tube features (e.g., retrusive calcite caps, concamerations, etc.) within and between borings (1) reflect ontogeny and variable behavioral responses to substrate overcrowding and/or degradation and (2) help to assess previously proposed models for teredinid bivalve life histories. Wood preservational state and significant textural variability among tube-filling sediments reflect extended residence time of log-grounds at or near the sediment-water interface. Like similar accumulations in other marine shelf sequences, the abundance of Teredolites-bearing log-ground accumulations in the Tallahatta can be attributed to both an influx pulse of xylic material and sediment starvation associated with transgression. The low ichnologic diversity (T. longissimus and very rare Thalassinoides) of these and other marine log-grounds may reflect a limited medley of wood-inhabiting tracemakers, but also likely reflects poor substrate preservation associated with physical reworking, biochemical degradation, and the bioerosion process itself.  相似文献   

7.
    
The Paleocene (Danian) Clayton Formation of western Alabama, USA, includes multiple marine shelf parasequences, each comprising a relatively thick marl, capped by a thin limestone, the latter variably reflecting marine flooding episodes. The marls host relatively large firmground burrow systems that penetrate 50–60 cm beneath, and are cast by, superjacent limestones. Excavation of two partially exposed burrow systems – one beneath a highstand parasequence-bounding flooding surface and the other beneath an overlying coplanar sequence boundary/transgressive surface (SB/TS) – reveals complex, primarily horizontal, irregularly branching networks. The former, allied with Thalassinoides paradoxicus, lacks wall bioglyphs, whereas the latter, allied with Spongeliomorpha iberica, is characterized by pervasive, mainly rhombohedral wall bioglyphs that reflect a relatively more firm substrate. Contrasts between these burrow systems are consistent with sequence stratigraphical context and inferred differences in the mechanism and magnitude of depositional hiatuses responsible for firmground development. Both excavated burrow systems likely represent cumulative structures produced by multiple organisms over extended periods of time. The cumulative nature and potential taphonomic biases associated with these and comparable burrow systems in the stratigraphical record preclude confident interpretation of tracemakers and their behaviours. The Clayton burrow systems likely were produced by one or more species of decapod crustacean that engaged in suspension-feeding, surface detritus feeding, gardening or some combination thereof.  相似文献   

8.
Middle Eocene, non-marine sediments from southern England contain examples of Teredolites borings in two contrasting palaeoenvironmental settings, viz.: (A) as in situ borings in an allochthonous lignite in an abandoned river channel and (B) as bored logs in cross-bedded fluvial sandstones of probable point-bar origin. The lignite is 0.30 m thick, of which the upper 0.20 m is intensely bored. Rounded pebbles of ?charcoal at its base also show small borings. A log in a fluvial sandstone shows densely-packed. radial club-shaped borings. now filled with sandstone. The wood substrate has subsequently been oxidized away. To our knowledge, this is the first detailed account of Teredolites from an ancient freshwater setting.  相似文献   

9.
10.
An occurrence of "Glossifungites saxicava”; Lomnicki (=Rhizocorallium jenense Zenker) is redescribed from its type area in Lviv, Ukraine, after more than 100 years since its first report. This trace fossil densely penetrates a Miocene transgressive surface cut in Cretaceous marlstones. Its occurrence is restricted to the top layer of the marlstones, where it was formed in a firmground typical of the Glossifungites ichnofacies. Most probably, the trace fossil was produced during a period of non‐deposition shortly before and at the beginning of deposition of Miocene strata by opportunistic suspension‐feeding organisms, possibly crustaceans.  相似文献   

11.
Joseph Barratt was a British-born and educated physician who settled in the United States in 1819. He had a great interest in natural history, collecting both plants and insects and studying geology, mycology, ornithology, chemistry, meteorology, Native Americans, and local history. He was apparently a man of great energy and ambition but one who could not focus to see a project to completion. Barratt was active in the early history of the discovery of vertebrate footprints in the Newark Supergroup in the eastern United States but latter developed some very strange theories regarding the age and significance of these deposits. In his latter years, Barratt's mental state deteriorated and he became even more obsessed with trying to publicize his outlandish theories. Dr. Joseph Barratt is remembered in ichnology for basically two things; he sold Edward Hitchcock a superb specimen of vertebrate tracks; and he may have the most elaborate ichnological tombstone ever constructed.  相似文献   

12.
The boring Teredolites longissimus Kelly and Bromley is recorded infesting silicified logs in the Fossil Wood Member, Kangilia Formation (Lower Danian, Paleocene) of West Greenland. There are two morphologies of T. longissimus, probably representing two different episodes of invasion of the wood. The initial borings of Morphology 1 are large, deformed to an elliptical section, and confined to a direction of boring parallel to sub-parallel to the wood grain. Morphology 2 borings are less common, narrower and shorter, sinuous, markedly crosscut the grain of the wood and, where preserved intact, retain their circular cross-section. They were a later infestation than Morphology 1. Following the infestation by Morphology 1, the logs were deformed, probably by weight of overburden, and then reworked, when some of them were infested by Morphology 2 producers. Final burial was followed by brittle collapse of some Morphology 2 borings and lithification before further deformation could occur. The occurrence of T. longissimus in deeper water deposits close to a land mass with a steep slope indicates that transport in the marine realm was minimal before waterlogging.  相似文献   

13.
    
We describe six proviverrine species from the Early Eocene of France. Three species are new: M inimovellentodon russelli sp. nov. from Mutigny [mammal palaeogene (MP)8 + 9], B oritia duffaudi sp. nov. from La Borie (MP8 + 9), and L eonhardtina godinoti sp. nov. from Grauves (MP10). We describe new specimens and propose new generic combinations for three species from MP10: Protoproviverra palaeonictides, Matthodon menui, and Oxyaenoides lindgreni. We also propose a new generic combination for the primitive Eoproviverra eisenmanni (MP7). Matthodon menui was previously considered as a possible oxyaenodontan, but the new fossils clearly support its reference to Hyaenodontida. Leonhardtina godinoti and Ma. menui are the oldest occurrences for these genera, which were previously unknown before the Middle Eocene. Moreover, the discovery of the proviverrine Mi. russelli in Mutigny implies that the Proviverrinae dispersed in Northern Europe between biozone Palaeocene‐Eocene (PE) III (Abbey Wood) and biozone PE IV (Mutigny). This also supports a homogenization of the European faunas during the Early Eocene. The dispersal is concomitant with the disappearance of the oxyaenodontans, arfiines, and sinopines (Hyaenodontida) from Europe. The proviverrines may have filled the ecological niches left vacant by the disappearance of the other carnivorous mammals. With 20 genera and over 30 species, proviverrines were successful in Europe. We performed the first phylogenetic analysis comprising almost all the Proviverrinae. Our analyses indicate that the Proviverrinae diversified greatly during the Early Eocene Climatic Optimum and show a general trend towards specialization throughout the Eocene. © 2014 The Linnean Society of London  相似文献   

14.
U-shaped, pouch-like burrows with parallel limbs, covered with short scratches arranged in sets, occur in the thalweg of the Oh?e river in NW Czech Republic. Similar, but smaller burrows with rare scratches, not arranged in sets, occur in the thalweg of the Drw?ca river in N Poland. Probably, they are produced by larvae and/or nymphs of Palingenia and Polymitarcis (Ephoron), respectively. In both localities, they burrowed in firmground surfaces at shallow depths. The burrowed surfaces were emerged during low water levels. A review of recent mayfly burrows shows that they are 1) U-shaped pouches with parallel limbs and septum, which may be covered with short scratches and are oriented perpendicular to the bottom, irrespective of its inclination, or 2) wide U-shape burrows with divergent limbs, which may be branched. In the fossil record, the ichnogenera Fuersichnus, Asthenopodichnium, and Rhizocorallium are partly ascribed to mayfly burrows, but their comparison to the recent burrows shows that such interpretations are somewhat problematic. The mayfly burrows are potentially good indicators of aquatic, non-marine, well oxygenated, clean water environments.  相似文献   

15.
    
Fossilized footprints contain information about the dynamics of gait, but their interpretation is difficult, as they are the combined result of foot anatomy, gait dynamics, and substrate properties. We explore how footprints are generated in modern humans. Sixteen healthy subjects walked on a solid surface and in a layer of fine‐grained sand. In each condition, 3D kinematics of the leg and foot were analyzed for three trials at preferred speed, using an infrared camera system. Additionally, calibrated plantar pressures were recorded. After each trial in sand, the depth of the imprint was measured under specific sites. When walking in sand, subjects showed greater toe clearance during swing and a 7° higher knee yield during stance. Maximal pressure was the most influential factor for footprint depth under the heel. For other foot zones, a combination of factors correlates with imprint depth, with pressure impulse (the pressure‐time integral) gaining importance distally, at the metatarsal heads and the hallux. We conclude that footprint topology cannot be related to a single variable, but that different zones of the footprint reflect different aspects of the kinesiology of walking. Therefore, an integrated approach, combining anatomical, kinesiological, and substrate‐mechanical insights, is necessary for a correct interpretation. Am J Phys Anthropol, 2010. © 2009 Wiley‐Liss, Inc.  相似文献   

16.
A new ichnotaxon is recognized in the Hidden Lake Formation, Upper Cretaceous of James Ross Island, Antarctica. Fuersichnus striatus consists of horizontal to subhorizontal, isolated or loosely clustered, U‐shaped, curved to banana‐like burrows, characterized by distinctive striations parallel to the trace axis. It is interpreted as a dwelling structure probably produced by crustaceans or polychaetes. This recording of Fuersichnus extends its stratigraphic range from the Triassic‐Jurassic to the Cretaceous and its environmental setting from nonmarine to marine environments. F. striatus typified consolidated, but unlithified substrates. Accordingly, it must be considered a member of the Glossifungites ichnofacies.  相似文献   

17.
    
Animal bioerosion trace fossils upon mineral substrates are analyzed from the point of view of the Seilacherian ethological classification. Several of the currently accepted ethological classes: cubichnia, fugichnia, repichnia, fodinichnia, agrichnia, calichnia and aedificichnia are not represented in these substrates. This fact points out a lower behavioral diversity of hard substrate trace fossils when compared with soft sediment trace fossils. Bioerosion traces can be classified in just five classes: domichnia, pascichnia, equilibrichnia, praedichnia and fixichnia. Fixichnia is here erected to gather superficial etching scars resulting from the anchoring of fixation of sessile epiliths by means of a soft or skeletal body part. Praedichnia and fixichnia are exclusive of the bioerosion realm.  相似文献   

18.
    
Abstract

The ichnogenus Kouphichnium and associated ichnofossils attributed to xiphosuran activity are here re-examined from samples collected from the Pennsylvanian-age Steven C. Minkin Fossil Site at the Union Chapel Mine, in Walker County, Alabama, USA. The large sample size offers an unique opportunity to evaluate some Kouphichnium ichnospecies. Thus, the morphological variability resulting from the taphonomic, ethological, taxonomic variability and underprint fallout are evaluated using this large sample set. Three morphotypes have been segregated from the material previously assigned to K. aspodon discovered at this and adjacent sites. Ichnospecies of Kouphichnium identified at the Union Chapel Mine site include: K. lithographicum, K. aspodon and two new ichnospecies (K. atkinsoni and K. minkinensis). Additionally, Kouphichnium-like traces that are associated with “jumper” traces have been previously misinterpreted as Kouphichnium and Selenichnites, respectively, and are excluded from this study, leaving them in open nomenclature as they will be the subject of a subsequent publication. We here redescribe the holotype of K. aspodon and designate lectoparatypes to better define the ichnospecies. New trace makers for some Kouphichnium ichnospecies are hypothesized, in contrast to the traditional xiphosuran attribution.  相似文献   

19.
Finds of fossil wood with bivalve wood borings (Teredolites clavatus and T. longissimus) occur in various facies and presumed sedimentary settings of the platform, shallow-marine Bohemian Crectaceous Basin. The basin comprises areas with sandy-dominated sediments, with marl and clay-dominated sediments, areas with predominat sandy-marly rocks, and finally areas dominated by calcareous nearshore sediments. Teredolites clavatus is common in fossil wood of sandstones, originating in beach or deltaic settings; marl and clay-dominated rock frequently bear wood fragments densely bored by Teredolites longissimus. When accompanied by evidence of marine environments as body fossils, glauconite or typical trace fossils, most of the wood fragments are bored. The presence/absence of borings in wood fragments can be considered the most reliable and easily useable criterion of distinction of marine settings in sandy sediments of the margin of the Bohemian Cretaceous Basin.  相似文献   

20.
    
We report a Holocene human and animal footprint site from the Namib Sand Sea, south of Walvis Bay, Namibia. Using these data, we explore intratrail footprint variability associated with small variations in substrate properties using a “whole foot” analytical technique developed for the studies in human ichnology. We demonstrate high levels of intratrail variability as a result of variations in grain size, depositional moisture content, and the degree of sediment disturbance, all of which determine the bearing capacity of the substrate. The two principal trails were examined, which had consistent stride and step lengths, and as such variations in print typology were primarily controlled by substrate rather than locomotor mechanics. Footprint typology varies with bearing capacity such that firm substrates show limited impressions associated with areas of peak plantar pressure, whereas softer substrates are associated with deep prints with narrow heels and reduced medial longitudinal arches. Substrates of medium bearing capacity give displacement rims and proximal movement of sediment, which obscures the true form of the medial longitudinal arch. A simple conceptual model is offered which summarizes these conclusions and is presented as a basis for further investigation into the control of substrate on footprint typology. The method, model, and results presented here are essential in the interpretation of any sites of greater paleoanthropological significance, such as recently reported from Ileret (1.5 Ma, Kenya; Bennett et al.: Science 323 (2009) 1197–1201). Am J Phys Anthropol 151:265–279, 2013. © 2013 Wiley Periodicals, Inc.  相似文献   

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