Root Reinforcement by Hawthorn and Oak Roots on a Highway Cut-Slope in Southern England

Highway embankments and cutting slopes in the United Kingdom, particularly in the South East of England, are often constructed of or within stiff over-consolidated clays. These clays are prone to softening with time leading to shallow slope failures and costly repairs. Reinforcement by natural vegetation is potentially a cost-effective method of stabilising these types of slopes over the medium–long term. However, there is a lack of information on how natural vegetation reinforces and stabilises clay slopes. To investigate this problem, the potential reinforcement of selected oak (Quercus robur L.) and hawthorn (Crataegus monogyna Jacq.) roots was assessed by conducting in situ root pull-out experiments on a London Clay cutting in south-east England. Pull-out tests were carried out using specifically designed clamps and either a hand pull system with a spring balance and manual recording of force for oak roots or a jacking system with electronic data logging of applied force and displacement for hawthorn roots. Oak roots had a mean pull-out resistance of 7 MPa and that of hawthorn roots was 8 MPa. The electronic data logging of applied force (pull-out resistance) and displacement of the hawthorn roots provided additional data on the failure of branched roots which could be correlated with variations in root morphology. The failure of the roots can be categorised into three modes: Type A: single root failure with rapid rise in pull-out resistance until failure occurs; Type B: double peak failure of a forked or branched root and Type C: stepped failure with multiple branches failing successively. The different types of root–soil bonds are described in relation to root anchorage and soil stability.     

The influence of plant diversity on slope stability in a moist evergreen deciduous forest

The influence of plant diversity on slope stability was investigated at early phases of succession in a mixed forest in Sichuan, China. The first phase comprised big node bamboo (Phyllostachys nidularia Munro) only. In the second phase, bamboo co-existed with deciduous tree species and in the third phase, deciduous species existed alone. Root density at different depths and root tensile strength were determined for each species. The factor of safety (FOS) was calculated for slopes with and without vegetation for each succession phase. For phase 2, FOS was determined for different species mixtures and positions. In phase 3, simulations were performed with a single tree at the top, middle or toe of the slope. Due to its shallow root system, bamboo contributed little to slope stability. In simulations with the tree at the top or middle of the slope, FOS decreased because tree weight added a surcharge to the slope. FOS increased with the tree at the bottom of the slope. Different mixtures of species along the slope had no influence on FOS. Differences in root tensile strength between species played a small role in FOS calculations, and tree size and density were the most important factors affecting slope stability, excluding hydrological factors.     

3-D numerical investigations into the shear strength of the soil–root system of Makino bamboo and its effect on slope stability

This study attempts to quantify the reinforcement effect of the Makino bamboo (Phyllostachys makinoi Hayata) root system on the stability of slopeland through numerical analyses and in situ tests. Based on the field surveys of Makino bamboo root morphology, a three-dimensional (3-D) numerical model of the soil–root system consisting of the reverse T-shape tap root and hair roots was developed and successfully applied to the finite element simulations of in situ pull-out tests. In the simulations, the soil mass was simulated by a soil element with a perfect elastic–plastic (or Mohr–Coulomb) material model whereas the root system was simulated by a ground anchor element with a linear elastic material model. In addition, a mechanical conversion model with simple mathematical form, which enables a direct transformation of the ultimate pull-out resistance into the shear strength increment of soil–root system was proposed. The conversion model offered a convenient way to quantify the reinforcement effect of the Makino bamboo root system required for the 3-D slope stability analyses. The numerical results indicated that the shear strength increment of the Makino bamboo soil–root system ranged from 18.4 to 26.3 kPa and its effect on the slope stability was insignificant when compared with those adverse influence factors such as the steep slope angle (=50–70°), shallow root depth (=0.8–1.0 m) and large growth height (>10 m) of the Makino bamboo forest slopeland. It can be also speculated that the tension cracks widespread over the slope surface due to the wind loading acting on the bamboo stems and the sequential rainwater infiltration is the dominating factor in the collapse failure of slopeland. For a Makino bamboo forest slopeland with medium slope (25° < slope angle β < 40°), the reinforcement effect of the Makino bamboo root system can mobilize its maximum stabilization capacity when compared with those of slopeland with mild (β < 25°) and steep slopes (β > 40°). Conclusively, the contribution of the Makino bamboo root system to the stability of slopeland is not as significant as expected.     

The Mechanics of Anchorage in Wheat Triticum aestivum L.: I. THE ANCHORAGE OF WHEAT SEEDLINGS

Model of the mechanics of uprooting lead to the identificationof ‘optimal’ anchorage systems which can withstanda given upward force at a minimum construction cost. Such systemshave many downward-pointing fibrous roots which are strengthenedprogressively towards the base. A study of the anchorage systemof 7- and 21-d-old wheat (Triticum aestivum L.) plants showedthat the plants possessed five seminal roots, of which onlythree pointed vertically. Each root was well suited for anchorage,being convered in root hairs and strengthened progressivelytowards the base by lignification of the stele. Strength andstiffiness of roots but not their mass per unit length increasedwith age. There was little interaction between roots when plantswere uprooted; the three vertical roots broke while the twohorizontal ones pulled out, as occurred when roots were pulledout singly, Uprooting forces increased with age and the rootsystem could withstand uprooting forces greater than those requiredto pull out upper leaves, so reducing the chances of the plantbeing uprooted by a herbivore, By 3 weeks a stiff adventitiousroot system, which would later help prevent the wheat lodging,was developing.     

Quantifying the role of vegetation in slope stability: A case study in Tuscany (Italy)

Vegetation significantly affects hillslope hydrological and mechanical properties related to shallow landslide triggering. In view of the complexity of soil plant hydrological interactions, the quantification of root mechanical reinforcement remains a challenge. Herein we present a back analysis of mechanical stability criteria related to a well-characterized vegetated shallow landslide in Italy, focusing on the quantification of lateral and basal root reinforcement. Lateral root reinforcement is included in slope stability estimates by adding a stabilizing force proportional to the scarp surface and root distribution. This stabilizing force is added to the force balance equation for the infinite slope model for different landslide shapes and dimensions. To quantify root reinforcement, we use the Wu model and the fiber bundle model (WM and FBM, respectively). Implementation of the latter model allows the quantification of the stress–strain behaviour of a bundle of roots for different root distributions and mechanical properties. Results of these models are compared highlighting key differences between the two approaches. Calculations using the FBM can explain the overestimation of lateral root reinforcement using WM and the commonly observed overestimation in the factor of safety. The model also quantifies the displacement-dependent behaviour of root reinforcement on vegetated slopes. Lateral root reinforcement can strongly influence the stability of slopes up to a certain area (1000–2000 m²). The magnitude of this stabilizing effect depends on parameters such as inclination, soil mechanical properties, and root distribution.     

Observation and Simulation of Root Reinforcement on Abandoned Mediterranean Slopes

The mechanics of root reinforcement have been described satisfactorily for a single root or several roots passing a potential slip plane and verified by field experiments. Yet, precious little attempts have been made to apply these models to the hillslope scale pertinent to landsliding at which variations in soil and vegetation become important. On natural slopes positive pore pressures occur often at the weathering depth of the soil profile. At this critical depth root reinforcement is crucial to avert slope instability. This is particularly relevant for the abandoned slopes in the European part of the Mediterranean basin where root development has to balance the increasing infiltration capacity during re-vegetation. Detailed investigations related to root reinforcement were made at two abandoned slopes susceptible to landsliding located in the Alcoy basin (SE Spain). On these slopes semi-natural vegetation, consisting of a patchy herbaceous cover and dispersed Aleppo pine trees, has established itself. Soil and vegetation conditions were mapped in detail and large-scale, in-situ direct shear tests on the topsoil and pull-out tests performed in order to quantify root reinforcement under different vegetation conditions. These tests showed that root reinforcement was present but limited. Under herbaceous cover, the typical reinforcement was in the order of 0.6 kPa while values up to 18 kPa were observed under dense pine cover. The tests indicate that fine root content and vegetation conditions are important factors that explain the root reinforcement of the topsoil. These findings were confirmed by the simulation of the direct shear tests by means of an advanced root reinforcement model developed in FLAC 2D. Inclusion of the root distribution for the observed vegetation cover mimics root failure realistically but returns over-optimistic estimates of the root reinforcement. When the root reinforcement is applied with this information at the hillslope scale under fully saturated and critical hydrological conditions, root pull-out becomes the dominant root failure mechanism and the slip plane is located at the weathering depth of the soil profile where root reinforcement is negligible. The safety factors increase only slightly when roots are present but the changes in the surface velocity at failure are more substantial. Root reinforcement on these natural slopes therefore appears to be limited to a small range of critical hydrological conditions and its mitigating effect occurs mainly after failure.     

Influence of slope on root system anchorage of Pinus yunnanensis

In southwestern China, Yunnan pines (Pinus yunnanensis) have been extensively cultivated on barren hills for reforestation and ecological engineering. The objective of this work is to study the influences of slope gradient on anchorage of root systems of P. yunnanensis. Pulling experiments were carried out at low (5–6°), moderate (25–26°) and high gradient (42–43°) by using selected 15-year-old P. yunnanensis. The results showed that the anchorage resistances induced by the slope gradient were significantly (P < 0.01) different from each other and followed in the order of high slope > moderate slope > low slope. Anchorage strength of upslope-grown roots increased with size and length of first-order lateral roots as well as the number of second-order lateral roots. Contributions of upslope-grown roots for preventing plants from overturning varied considerably between those from different slopes. The contribution of roots growing on upslope side on high slope to the anchorage of root system reached 50%, whereas those on moderate and low slopes were about 44% and 37%, respectively. It was concluded that the anchorage resistance was closely related to slope gradient and root distribution, while upslope-grown roots were positively related to anchorage resistance.     

The influence of slope on <Emphasis Type="Italic">Spartium junceum</Emphasis> root system: morphological,anatomical and biomechanical adaptation

Root systems have a pivotal role in plant anchorage and their mechanical interactions with the soil may contribute to soil reinforcement and stabilization of slide-prone slopes. In order to understand the responses of root system to mechanical stress induced by slope, samples of Spartium junceum L., growing in slope and in plane natural conditions, were compared in their morphology, biomechanical properties and anatomical features. Soils sampled in slope and plane revealed similar characteristics, with the exception of organic matter content and penetrometer resistance, both higher in slope. Slope significantly influenced root morphology and in particular the distribution of lateral roots along the soil depth. Indeed, first-order lateral roots of plants growing on slope condition showed an asymmetric distribution between up- and down-slope. Contrarily, this asymmetric distribution was not observed in plants growing in plane. The tensile strength was higher in lateral roots growing up-slope and in plane conditions than in those growing down-slope. Anatomical investigations revealed that, while roots grown up-slope had higher area covered by xylem fibers, the ratio of xylem and phloem fibers to root diameter did not differ among the three conditions, as also, no differences were found for xylem fiber cell wall thickness. Roots growing up-slope were the main contributors to anchorage properties, which included higher strength and higher number of fibers in the xylematic tissues. Results suggested that a combination of root-specific morphological, anatomical and biomechanical traits, determines anchorage functions in slope conditions.     

A generic 3D finite element model of tree anchorage integrating soil mechanics and real root system architecture

Understanding the mechanism of tree anchorage in a forest is a priority because of the increase in wind storms in recent years and their projected recurrence as a consequence of global warming. To characterize anchorage mechanisms during tree uprooting, we developed a generic finite element model where real three-dimensional (3D) root system architectures were represented in a 3D soil. The model was used to simulate tree overturning during wind loading, and results compared with real data from two poplar species (Populus trichocarpa and P. deltoides). These trees were winched sideways until failure, and uprooting force and root architecture measured. The uprooting force was higher for P. deltoides than P. trichocarpa, probably due to its higher root volume and thicker lateral roots. Results from the model showed that soil type influences failure modes. In frictional soils, e.g., sandy soils, plastic failure of the soil occurred mainly on the windward side of the tree. In cohesive soils, e.g., clay soils, a more symmetrical slip surface was formed. Root systems were more resistant to uprooting in cohesive soil than in frictional soil. Applications of this generic model include virtual uprooting experiments, where each component of anchorage can be tested individually.     

Modulated dispersion of activation and repolarization by premature beats in patients with cardiomyopathy at risk of sudden death

Premature beats can trigger ventricular arrhythmias in heart disease, but the mechanisms are not well defined. We studied the effect of premature beats on activation and repolarization dispersion in seven patients with cardiomyopathy (57 ± 10 yr, left ventricular ejection fraction 31 ± 7%). Activation time (AT), activation-recovery interval (ARI), and total repolarization time (TRT) were measured from 26 unipolar electrograms during right ventricle (RV) endocardial (early) to left ventricle epicardial (late) activation in response to RV apical extrastimulation (S1S2). Early TRT dispersion increased significantly with shorter S1S2 (1.0 ± 0.2 to 2.3 ± 0.4 ms/mm, P < 0.0001), with minimal change in late TRT dispersion (0.8 ± 0.1 to 1.0 ± 0.3 ms, P = 0.02). This was associated with an increase in early AT dispersion (1.0 ± 0.1 to 1.5 ± 0.2 ms/mm, P = 0.05) but no change in late AT dispersion (0.6 ± 0.1 to 0.7 ± 0.2 ms/mm, P = 0.4). Early and late ARI dispersion did not change with shorter S1S2. AT restitution slopes were similar between early and late sites, as was slope heterogeneity. ARI restitution slope was greater in early vs. late sites (1.3 ± 0.6 vs. 0.8 ± 0.6, P = 0.03), but slope heterogeneity was similar. With shorter S1S2, AT-ARI slopes became less negative (flattened) at both early (-0.4 ± 0.1 to +0.04 ± 0.2) and late (-1.5 ± 0.2 to +0.3 ± 0.2) sites, implying less activation-repolarization coupling. There was no difference in AT-ARI slopes between early and late sites at short S1S2. In conclusion, high-risk patients with cardiomyopathy have greater TRT dispersion at tightly coupled S1S2 due to greater AT dispersion and activation-repolarization uncoupling. Modulated dispersion is more pronounced at early vs. late activated sites, which may predispose to reentrant ventricular arrhythmias.     

Structural development and stability of rice Oryza sativa L. var. Nerica 1

The structural development of glasshouse-grown rice Oryza sativa L. var. Nerica 1 was studied in relation to its stability against lodging. The morphology and mechanical properties of both the stem and roots were examined from tillering, 4 weeks after transplantation up to maturity, together with plant weight distribution and anchorage strength. The "factors of safety" against root and stem failure were subsequently calculated throughout development. Rice plants showed similar morphology to wheat, although they possessed around twice as many tillers per plant and 10 times as many coronal roots. The mechanics of anchorage were also similar. The strength and rigidity of individual tillers increased throughout development as the plants grew taller and heavier and were around 15 times greater than in wheat. By contrast, individual root bending strength, the number of roots, and the anchorage strength levelled off earlier, and anchorage strength was only around twice that in wheat. Consequently, while the self-weight safety factor against stem failure was much higher than in wheat, increasing until late on in development from around 30 to 150, the self-weight safety factor against root anchorage failure was similar to wheat, decreasing from around 15 to 5. Consequently, plants subjected to anchorage tests always failed in their root system rather than their shoot system. The results suggest that, in the field, rice plants would be more likely to undergo root lodging than stem lodging, and that breeding efforts to reduce the incidence of lodging should act to strengthen the rather weak coronal roots.     

Vegetation Succession and its Consequences for Slope Stability in SE Spain

The effect of land abandonment as a result of changing land-use policies is becoming more and more important throughout Europe. In this case study, the role of vegetation succession and landslide activity on steep abandoned slopes was investigated. The influence of vegetation succession on soil properties over time, as well as how developing root systems affect soil reinforcement was determined. The study was carried out in the Alcoy basin in SE Spain, where the marl substratum is prone to landsliding along steep ravines. The bench-terraced slopes have been abandoned progressively over the last 50 years and show various stages of revegetation. The study was carried out at two scales; at the catchment scale long-term evolution of land-use, vegetation succession and slope failure processes were investigated. At a more detailed scale, vegetation cover, soil properties and rooting effects on soil strength were determined. Results showed that the soil has changed over a period of 50 years with respect to soil properties, vegetation cover and rooting, which is reflected in the activity of geomorphological processes. Vegetation succession progressively limits surface processes (sheet wash and concentrated overland flow) over time, whereas slopes affected by mass wasting processes increase in number. The spatial heterogeneity of infiltration increases over time, leading to increased macro-pore flow towards the regolith zone, enhancing the potential risk of fast wetting of the regolith directly above the potential plane of failure, as was concluded from rainfall simulations. In situ experiments to determine soil shear strength in relation to rooting indicated that roots contributed to soil strength, but only in the upper 0.4 m of the soil. Most failures however, occur at greater depths (1.0–1.2 m) as anchorage by deeper roots was not effective or absent. The observed initial increase in mass wasting processes after land abandonment can therefore be explained in two ways: (1) the limited contribution of anchorage by root systems at potential slip planes which cannot counterbalance the initial decline of the terrace walls, and (2) the fast transfer of rainfall to the potential slip plane by macro-pores enhancing mass movements. However, after approximately 40 years of abandonment, mass wasting processes decline.     

Model and whole-plant studies on the anchorage capabilities of bulbs

Though the resistance to uprooting of cylindrical roots and root systems has been extensively investigated, almost no research has been performed on the factors that influence the uprooting resistance of bulbs. However, engineers have modelled bulb-like foundations and have investigated their resistance to upward movements. This study combined engineering theory with practical biology, using model bulbs of different shapes and sizes, embedding them at different depths in different soil media, and pulling them out while recording the uprooting force. Uprooting resistances of the models was compared to those of real onion and garlic bulbs with and without their root systems. Cone shaped models resisted uprooting best at all embedment depths and in both soil types, always followed by bulb shaped and cylindrical models. These results are explicable in terms of engineering theory. Cones resisted uprooting best because their maximum diameter is embedded deepest. A bulb shape is an ideal compromise as it has no sharp edges, and also allows easy downward movement. In sand uprooting resistance increased faster with depth than with bulb diameter, whereas in agricultural soils, the uprooting force was proportional both to the depth and the diameter of the model. The tests on the plants showed that real bulbs anchor plants by similar mechanisms and amounts to the models. The bulbs accounted for between 15% and 50% of the uprooting resistance of the plant, so they can make an important contribution to anchorage, particularly towards the end of the season.     

Linking plant morphological traits to uprooting resistance in eroded marly lands (Southern Alps, France)

In marly catchments of the French Southern Alps, soils are subjected to harsh water erosion that can result in concentrated flows uprooting small plants. Evaluating and predicting plant resistance to uprooting from simple plant traits is therefore highly important so that the most efficient plant strategy for future restoration of eroded slopes can be defined. Twelve species growing on marly land were studied. For each species, in-situ lateral uprooting tests were conducted and morphological plant traits were measured on small plants at the early stages of their development. The results show that maximum uprooting force was most positively correlated with stem basal diameter. Resistance to uprooting depends on a combination of several traits. Tap root length, the proportion of fine lateral roots and root topology were the best predictors of anchorage strength.     

The Anchorage of Leek Seedlings: The Effect of Root Length and Soil Strength

The mechanical behaviour of single roots being extracted fromsoil was modelled as a process in which tension is transferredfrom the upper regions of the root to the soil via shear. Quantitativepredictions were made about the extraction forces and the shapeof the uprooting curves, and these were tested using leek radiclesof different lengths in soil of two different strengths. Results of uprooting tests were qualitatively similar to thepredictions. The pullout resistance rose with root length, untilthe breaking strength of the root was reached, at around 30mm: longer roots all broke before the tip was stressed. In wholeroot systems, therefore, failure will occur proximally beforethe line distal roots are mechanically stressed, so these canhave no anchorage function. Resistance to an upward force will be most economically achievedby having many strengthened proximal root axes, as in the adventitiousroot systems of grasses, sedges and stoloniferous dicots. Allium porrum, root, anchorage, shear, tension, soil     

广西西南部熊猴对石山森林的利用

栖息地利用直接反映灵长类适应环境或应对环境变化所采取的行为策略,是了解和评估物种适应能力与进化潜力的重要途径.石山森林是一类特殊的生境,栖息在石山森林中的灵长类动物可能展现出独特的适应策略.为了解熊猴(Macaca assamensis)对石山森林的利用策略,2012年9月至2013年8月采用瞬时扫描法对广西弄岗国家级...     

The anchorage mechanics of deep rooted larch, Larix europea L. japonica

The anchorage of deep rooted 16-year-old larch trees, Larixeuropea japonica, has been studied by combining winching testswith analyses of strain around the base of the trunk and rootsystem and mechanical tests on individual roots. These showedthat anchorage is provided by the laterals which emerge fromaround the stem base, sinker roots which emerge along theirlength, and tap roots positioned directly underneath the bole.During anchorage failure the leeward laterals are bent and eventuallybreak close to their base, whilst the windward laterals arepulled out of the ground, with their sinker roots intact. Afterinitially being confined by the soil and bending, the tap rootrotates in the soil. Anchorage failure is similar when the soilis dry as when it is wet, but failure occurs closer to the trunk.Strain measurements along the lateral roots revealed that thestresses were highest close to the trunk and that these regionsof the roots contribute most to tree stability. The two major components of anchorage were found to be the resistanceof leeward laterals to bending and the resistance of tap rootsand windward sinkers to uprooting. Bending tests on leewardlaterals revealed that they provide around 25% of tree anchorage.Almost 75% of the anchorage strength must, therefore, be providedby the windward sinkers and tap roots. Anchorage strength ofroots was positively correlated to their cross-sectional area.The vertical orientation of the sinkers makes the anchoragesystem of larch more efficient than the plate system formedby Sitka spruce on waterlogged soils and means that no root-soilplate is formed. Key words: Anchorage, root architecture, sinker roots, root bending strength, windthrow     

Sensorimotor function of the upper-airway muscles and respiratory sensory processing in untreated obstructive sleep apnea

Numerous studies have demonstrated upper-airway neuromuscular abnormalities during wakefulness in snorers and obstructive sleep apnea (OSA) patients. However, the functional role of sensorimotor impairment in OSA pathogenesis/disease progression and its potential effects on protective upper-airway reflexes, measures of respiratory sensory processing, and force characteristics remain unclear. This study aimed to gain physiological insight into the potential role of sensorimotor impairment in OSA pathogenesis/disease progression by comparing sensory processing properties (respiratory-related evoked potentials; RREP), functionally important protective reflexes (genioglossus and tensor palatini) across a range of negative pressures (brief pulses and entrained iron lung ventilation), and tongue force and time to task failure characteristics between 12 untreated OSA patients and 13 controls. We hypothesized that abnormalities in these measures would be present in OSA patients. Upper-airway reflexes (e.g., genioglossus onset latency, 20 ± 1 vs. 19 ± 2 ms, P = 0.82), early RREP components (e.g., P1 latency 25 ± 2 vs. 25 ± 1 ms, P = 0.78), and the slope of epiglottic pressure vs. genioglossus activity during iron lung ventilation (-0.68 ± 1.0 vs. -0.80 ± 2.0 cmH(2)O/%max, P = 0.59) were not different between patients and controls. Maximal tongue protrusion force was greater in OSA patients vs. controls (35 ± 2 vs. 27 ± 2 N, P < 0.01), but task failure occurred more rapidly (149 ± 24 vs. 254 ± 23 s, P < 0.01). Upper-airway protective reflexes across a range of negative pressures as measured by electromyography and the early P1 component of the RREP are preserved in OSA patients during wakefulness. Consistent with an adaptive training effect, tongue protrusion force is increased, not decreased, in untreated OSA patients. However, OSA patients may be vulnerable to fatigue of upper-airway dilator muscles, which could contribute to disease progression.     

The mechanics of anchorage in seedlings of sunflower, Helianthus annuus L.

Forces applied to plants will subject many of the roots to tension, which must be transferred to the soil via shear if uprooting is to be prevented. The stress distribution will depend on the relative stiffnesses of the earth and root, and the mode of failure will depend on the relative strength of the soil and of the root soil bond. This study of the anchorage of sunflower radicles combined uprooting tests performed by a tensile testing machine with mechanical tests on the roots and soil.
The maximum extraction force increased with length to an asymptotic value and was reached at a very low displacement. Root hairs and soil particles covered the tapered top 20 mm of extracted root, but the lower cylindrical region was bare. The soil was stiffer than the root, so shear stress was initially concentrated at the top of the root, soil strength over the top 20 mm resisting uprooting. Lower regions of the root were stressed later, their sparser root hairs being sheared off, and resist uprooting only by friction. In a further lest upper and lower regions of radicles were uprooted separately. As predicted, the upper region generated much greater resistance to uprooting per unit length, and at much lower displacements than the lower region.
The top of the radicle is well adapted for anchorage, the profuse root hairs and mucigel it produces glueing the root to the soil. The lower regions are thus protected from damage.     

秦岭大熊猫栖息地巴山木竹生物量

巴山木竹是秦岭大熊猫冬、春季食物的主要来源,其生长状况、数量和空间分布格局与大熊猫的生存与活动有着密切的关系。为了探讨不同立地因子对巴山木竹生物量的影响,确定巴山木竹生长的主导因子和适宜立地条件,在秦岭大熊猫栖息地内选取54块样地,对不同立地条件下巴山木竹生物量进行了测定。通过统计分析,结果表明:由于不同地区气候和土壤条件的差异,导致了巴山木竹生物量亦存在较大差别;巴山木竹生物量在其分布区内随海拔的升高而呈规律性变化,1500—1700 m为最适分布区域,其生物量最大;坡度越大生物量越小,陡坡不适宜竹林的生长;上坡位不利于竹林生长,而中、下坡位生物量没有显著差异;坡向对生物量的影响不明显,均适宜巴山木竹生长;在落叶阔叶和针阔混交林中,当林冠郁闭度>0.6时,竹林生长减弱。在这些立地因子中,影响巴山木竹生物量积累的主导因子是坡位和林冠郁闭度。