Cereal weeds variation in middle Egypt: Role of crop family in weed composition

Cereals occupies a major part in the diet of humans globally, participating more to our daily protein and calorie intake than any other crop. The present study highlight the weed flora of cereal crops compared to other crops in middle Egypt and their distribution. Ninety-two weed species were recorded in the all studied crops, cereal and other crops; in the studied area belong to 67 genera and 20 families. Egyptian clover; showed the highest numbers of both weed species and genera followed by wheat, on contrast the lowest weed species and genera numbers were recorded associated with Solanaceous crops tomato and pepper. Wheat crops exhibited the highest number of weed species, among cereals, followed by maize crop, while the lowest weed species number was detected in barley crop. Chenopodium murale, Cynodon dactylon, Convolvulus arvensis and Malva parviflora were the most frequent species in winter cereals, while Echinochloa colona, P. oleraceae were the most frequent weeds in summer cereals. Chorological analysis of the recorded weed species showed that cosmopolitan elements showed the highest numbers in total weed flora Differences in weed species compositions were fundamentally influenced by seasonal priority. Based on TWINSPAN and Ward classifications, crop family showed slightly effect as a factor affecting weed composition.     

Weeds flora demographic analysis of date palm orchards in El-Bahariya,El-Frafra and Siwa oases

Oases are a specific ecosystems which date palm Phoenix dactylifera are the mainly crop growing for its fruits and others tree parts. Palm orchards are infested by many pests, including weeds species and exposed within time to the challenging invasive weeds to control. The heavy incidence of weeds negatively affects palm productivity, especially in small plantation and divergent distances, while the established orchards, palm trees shading cause deprives from the process of weeds photosynthesis to be lower competitive. The survey was conducted in El-Bahariya and El-Frafra than Siwa oases of which 375 plots were taken. Vegetative analysis in oases altogether demonstrate 91 weed specimens were encountered of which 18 perennial, 42 were broad leaved, and 30 narrow leaved as well as one parasitic weed species belonging to 22 families, whereas Poaceae is the most plentiful family displayed by 38.5% from all families, occurred in date palm orchards during the study. The geographical differentiation affected coefficient of similarity whereas, it in oases recorded the highest between El-Bahariya and El-Frafra than Siwa and El-Bahariya, while El-Frafra Oasis was the most diverse and richness than the others which have the middle soil properties, fertility, and structure. The correlation analysis of weed flora between both El-Bahariya and El-Frafra was higher and more than El-Bahariya and Siwa oases. Under the unique closed agroenvironment of oases, weeds population dynamics are a stable approximately. Regardless of the diversity of annual weed species, perennials species represent the highest dominant number and the pattern of grown weeds was more dominant than wild desert and some invasive weed species. The control action should be taken of perennial weeds and proactive as well as long term prevention for invasive species.     

Narrow rows reduce biomass and seed production of weeds and increase maize yield

Smallholder farmers in southern African countries rely primarily on cultural control and hoe weeding to combat weeds, but often times, they are unable to keep up with the weeding requirements of the crop because of its laboriousness, causing them to incur major yield losses. Optimisation of crop planting pattern could help to increase yield and suppress weeds and to reduce the critical period of weed control and the weeding requirements to attain maximum yield. Experiments were carried out in Zimbabwe during two growing seasons to assess the effect of maize density and spatial arrangement on crop yield, growth and seed production of weeds and to determine the critical period for weeding. Planting maize at 60 cm row distance achieved higher yields and better weed suppression than planting at 75 or 90 cm row distance. Increasing crop densities beyond the customary three to four plants m⁻² gave modest reductions in weed biomass but also diminished crop yields, probably because of increased competition for water and nutrient resources. Maize planted in narrow rows (60 cm) intercepted more radiation and suffered less yield reduction from delaying hoe weeding than those planted in wider rows (75 or 90 cm), and the duration of the weed-free period required to attain maximum grain yield was 3 weeks shorter in the narrow spacing than that in the 75- and 90-cm row spacings. Weeding was more effective in curtailing weed seed production in the narrow row spatial arrangements than in the wide row planting. The results of these studies show that narrow row spacings may reduce weeding requirements and increase yields.     

Adventive flora of Israel — Phytogeographical,ecological and agricultural aspects

73 adventive wild species occur in the flora of Israel: 39 are of tropical origin, 20 are North American. The geographical distribution pattern suggests that about two thirds of the species reached Israel through neighbouring countries, while only one third arrived directly from their countries of origin. — Three patterns of population dynamics can be recognized: accidental species (7), colonizing species (30), and species penetrating into natural habitats (3). There is insufficient information about the remaining species. — About 20 species are widespread noxious weeds which now have economic significance in Israel, 20 others are known to have similar ecological tendencies in other countries. These species soon may become aggressive weeds in Israel.     

Measuring the ecological preference for growth of 150 of the most influential weeds in weed community structure associated with agronomic and horticultural crops

The phytosociological researches which intent for studying the performance of weeds and the structure of weed assemblages associated with different crops derives their importance mainly from the adverse effect of weeds on crop productivity. Consequently, it is worth questioning about the ecological preferences of the weed growth in response to three main drivers for weed community structure associated with agronomic, and horticultural crops: crop diversification, crop seasonality, and soil type. A study area was selected comprising farmland of Nile Delta and its adjoining east and west territories, Egypt. A total of 555 species were recorded in 30 agroecosystems monitored and depending on species frequency/abundance values, 150 species were designated as the most influential weeds in weed community structure associated with agronomic and horticultural crops. The ecological preference of species for crop seasonality was evident through the results of Agglomerative hierarchical clustering. Three weed assemblage groups (WAG) identified: WAG A associated with winter agronomic crops, WAG B associated with summer agronomic crops, and WAG C associated with perennial agronomic crops and horticultural crops (orchards). Their diversity evaluated at different levels. The growth preference of the 150 species which were assigned as most influential weeds was gauged in response to the three environmental variables. 61 species were faithful to WAG A, 45 to WAG B, and 44 to WAG C. Concerning crop diversification, 34-species were significantly affected and scored coefficient of variation ≥ 100%. As for soil type, indicator species analysis revealed that 66-species show growth preference in fine grained soil while 84-species prefer coarse grained soil. In the three vegetation units (WAG A – C), 12 within-group associations (alliances) were specified of less-common (differential) species. The record of these alliances match to a specific environmental condition (ecological niche) and in them 29 strong indicators are identified. Redundancy analysis was used to extract and summarize the variation in species records in the response matrix (species vs. sites) that can be explained by the three different types of growth preference (explanatory variables), and the partial linear effect of them was evaluated by variation partitioning.     

Impact of weed control on arbuscular mycorrhizal fungi in a tropical agroecosystem: a long-term experiment

Cover crop species represent an affordable and effective weed control method in agroecosystems; nonetheless, the effect of its use on arbuscular mycorrhizal fungi (AMF) has been scantily studied. The goal of this study was to determine root colonization levels and AMF species richness in the rhizosphere of maize plants and weed species growing under different cover crop and weed control regimes in a long-term experiment. The treatment levels used were (1) cover of Mucuna deeringian (Muc), (2) "mulch" of Leucaena leucocephala (Leu), (3) "mulch" of Lysiloma latisiliquum (Lys), (4) herbicide (Her), (5) manual weeding (CD), (6) no weeding (SD), and (7) no maize and no weeding (B). A total of 18 species of AMF belonging to eight genera (Acaulospora, Ambispora, Claroideoglomus, Funneliformis, Glomus, Rhizophagus, Sclerocystis, and Scutellospora) were identified from trap cultures. Muc and Lys treatments had a positive impact on AMF species richness (11 and seven species, respectively), while Leu and B treatments on the other hand gave the lowest richness values (six species each). AMF colonization levels in roots of maize and weeds differed significantly between treatment levels. Overall, the use of cover crop species had a positive impact on AMF species richness as well as on the percentage of root colonized by AMF. These findings have important implications for the management of traditional agroecosystems and show that the use of cover crop species for weed control can result in a more diverse AMF community which should potentially increase crop production in the long run.     

Developing a set of simulation-based indicators to assess harmfulness and contribution to biodiversity of weed communities in cropping systems

Weeds are both harmful for agricultural production and an essential component of biodiversity in agricultural landscapes. Therefore, new cropping systems aiming at both maximising weed-related biodiversity and minimising weed harmfulness are needed. New cropping systems are now increasingly designed with weed dynamics models but these usually only consider weed densities or crop yield losses. The present paper proposed a set of indicators for assessing the impact on crop production and biodiversity of weed communities simulated with a cropping system model. Five harmfulness indicators were developed to take account of the criteria most frequently listed by farmers via an internet survey: (1) crop yield loss, (2) harvest pollution by weed seeds, stems and leaves, (3) harvesting problems due to green weed biomass blocking the combine, and (4) field infestation represented by weed biomass averaged over cropping seasons. A fifth indicator was added, i.e. (5) the increase in crop disease (i.e. take-all disease of cereals) in the presence of weeds. The biodiversity indicators were chosen in collaboration with ecologists. Two indicators reflect the weed contribution to vegetal biodiversity: (1) species richness and (2) Pielou's index for species equitability. Three other indicators were developed to assess weeds as a trophic resource for other organisms in the agro-ecosystems: (3) the number of weed seeds present on soil surface in autumn and winter to feed field birds, (4) lipid-rich seeds on soil surface in summer to feed insects such as carabids, and (5) weed flowers in spring and summer to feed domestic bees. These indicators were tested in a series of contrasted cropping systems identified in farm surveys and simulated with FlorSys. Analyses of variance showed that the cropping system and the crop sequence presented the highest impact on indicator values. Weather scenario and pedoclimate had little effect. Antagonisms and synergies between weed-related harmfulness and biodiversity were identified with Spearman correlations. Harmfulness indicators were all positively correlated, except for additional disease risk which was at best poorly correlated with other indicators. Most weed-related biodiversity indicators were also positively correlated, except species richness which was negatively correlated with species equitability, bird resource and insect resource. Weed harmfulness generally increased with increasing weed-related biodiversity. These correlations were though weak, and others were negative, showing that increased biodiversity could occur with decreased harmfulness (e.g. trophic resource for insects vs. yield loss or field infestation). Consequently, there are cropping systems that reconcile agricultural production and biodiversity.     

The conservation of arable weeds at crop edges of barley fields in northeast Spain

The effects of weed control practices and fertilisation on weed flora and crop yield were evaluated in crop edges of barley fields in northeastern Spain. The study was carried out in four organic and four conventional barley fields. In each field, four permanent plots were delimited at the crop edge, and fertilisation and weed control treatments in a factorial design were applied over 3 years. Weed composition and the aboveground biomass of weeds and barley were recorded before the crop harvest in the first and the third year. We found relatively low values of species richness per field, as well as low values of weed biomass, especially in the organic crop edges (3.9% of total biomass). Weeds were significantly reduced by herbicide applications on conventional fields and were not affected by weed harrowing on organic fields or fertilisation. These results demonstrate that specific measures are needed to enhance biodiversity at crop edges both in organic and conventional fields. Our results also suggest that under Mediterranean conditions and among impoverished weed communities, limiting the use of herbicides is crucial to enhancing arable diversity and that, contrary to findings found in previous studies in temperate climates, fertilisation and weed harrowing have little effect on weeds.     

保护性耕作对陇东黄土高原轮作田杂草的影响

以陇东黄土高原旱地连续12年保护性耕作为基础,通过田间取样调查,研究了4种耕作措施下冬小麦-箭筈豌豆-玉米轮作田中杂草的组成和群落特征.结果表明： 不同作物生长期的杂草种类、密度和群落特征各异.免耕使冬小麦生长期杂草密度显著增大,轮作箭筈豌豆后免耕处理的杂草密度变小;秸秆覆盖显著降低了玉米生长期的杂草密度,玉米生长期免耕+秸秆覆盖处理的杂草密度最低.冬小麦和箭筈豌豆轮作前后杂草的群落多样性特征完全相反,从冬小麦生长期到箭筈豌豆生长期,免耕处理杂草群落的多样性指数先高于常规耕作处理,后期低于常规耕作处理;玉米生长期免耕处理的多样性指数均大于其他3种处理.因此,免耕和秸秆覆盖在一些作物生长期能抑制杂草的发生和分布,3种作物轮作对杂草防治具有明显作用.     

A phytosociological study of weed communities on the southwestern Coastal Plain of North Carolina

Summary The vegetation of cultivated, fallow and ‘old’ fields of Scotland County, N.C. was both randomly and subjectively sampled throughout the growing season of 1973. Relevés were placed in each of the above management units, and the data collected in this relevé analysis were sorted into phytosociological charts, with the aid of the digital computer. Weed associations delimited in the charts reflected the importance of edaphic factors, time (aspect), and man's agricultural activities as causal factors in the determination of characteristic assemblages of weeds. Several character and differential species were denoted, in a tentative classification of the weed associations. The cultivated and fallow stands in the spring aspect were noted to be dominated by a group of spring annuals. Summer annuals were more prevalent in homologous situations in the summer. It was determined that perennials, appearing mainly as seedlings in cultivated and fallow fields, attained dominance in ‘old’ fields, one to five years in age. Within each of the major spheres of management, and in each season, associations of weeds mainly reflected differences in edaphic factors. Many character species were noted to maintain affinities to an assortment of soil types that ranged from moist loams to xeric, sterile sands. A study of roadsides, hedgerows, field margins and other ecotonal areas inhabited by weeds was carried throughout the growing season of 1973, also. Species presence was documented in each of these habitat types and reported in Sharp (1974). The weed flora of Scotland County was found to be rich in species. A total of 506 species representing 248 genera and 80 families were encountered throughout the study. Of the total number of species listed, approximately 25 percent were of foreign origin, and 28 percent represented additions to the flora of Scotland County. This paper is the short version of a masters thesis for the Department of Botany, University of North Carolina at Chapel Hill. The author would like to extend his appreciation and gratitude to Professors H. Lieth, A. E. Radford and J. Hardin for their generous advice and encouragement extended through the duration of the study.     

不同耕作与施肥方式对有机玉米田杂草群落和作物产量的影响

农田杂草是农业生态系统的重要组成部分。有机农田杂草控制是农业生产的一大难题,通过施肥方式改变杂草的竞争格局有望成为杂草控制的新途径。以暖温带有机玉米田为研究对象,设置了5种不同处理,即不施肥对照处理（CK）、施用牛粪处理（CM）、施用蚯蚓处理过的牛粪处理（EM）、免耕秸秆不覆盖处理（NT）和免耕秸秆覆盖处理（NS）,研究不同耕作和施肥方式对田间杂草密度、生物量、生物多样性及作物产量的影响。结果表明,在有机玉米农田内共发现杂草17种,CK杂草总密度最高,但杂草总生物量和生物多样性指数较低。EM杂草总生物量和牛筋草（Eleusine indica）生物量最高,分别比CK增加了192.6%和224.8%（P〈0.01）,物种丰富度和生物多样性指数较低,优势集中度指数较高。NS杂草总密度、总生物量和生物多样性与NT相比显著降低。此外,EM玉米（Zea mays）产量比CK高40.2%（P〈0.01）,比CM高19.6%（P〈0.01）。施用蚯蚓处理过的牛粪不仅提高了玉米产量,而且可以促进优势种杂草的生长,提高杂草群落的优势集中度。     

黄淮海地区夏玉米田一年生杂草的生态经济防治阈期研究

 本文通过北京和唐山多年多点田间试验,研究了黄淮海地区夏玉米田苗后放任和免除自生一年生杂草生长的相对时间与夏玉米相对产量的函数关系,并据此导出了该区夏玉米田一年生杂草的生态经济防治阈期计算模型。结果发现,该区夏玉米田一年生杂草的生态经济防治阈期约处在夏玉米苗后总生育期的(天)第11.2—50.9%之间。该区夏玉米生长季节中只要此期田间保持无草,便可望以最小的代价和费用,最大限度地使杂草弃害扬利,从而获得较高的生态经济除草效益。     

Introduced weed richness across altitudinal gradients in Hawai’i: humps,humans and water-energy dynamics

Native species richness commonly declines with increasing altitude, but patterns of introduced species richness across altitudinal gradients have been less frequently studied. We surveyed introduced roadside weeds along altitudinal transects ranging from 30 to 4,100 m in Hawai’i, with the objectives of (1) testing the hypothesis that a mass effect due to mixing of tropical and temperate species at mid-elevation promotes a hump-shaped pattern of introduced species richness with altitude, and (2) testing the potential roles of anthropogenic activity, energy (temperature) and water-energy dynamics (productivity-diversity hypothesis) in determining introduced weed richness. A total of 178 introduced weeds were recorded. Introduced weed richness does not decline monotonically with altitude. Rather, mixing of tropical and temperate species helps to maintain high mean richness up to 2,000 m, suggesting a mass effect, but without a distinct richness peak. Patchy occurrence of a transformer species, Pennisetum clandestinum, introduced high variance in richness at mid-elevations. General linear models considering estimated actual evapotranspiration (AET, a measure of energy-water dynamics) together with an index of human activity (distance from urban area or length of major roads) accounted for more variance in introduced weed richness than models with energy alone (temperature) and human activity. Native Hawaiian species richness along roadsides was also weakly correlated with AET but negatively associated with human activity. Our observed association between introduced species richness and AET mirrors patterns reported for native species richness around the world, indicating that AET-richness patterns can develop on a short time scale (on the order of 100 years). To test the generality of introduced weed richness patterns, we tried using the Hawai’i island model to predict weed richness on the neighboring island of Maui. Although weed richness on Maui was under-predicted, the same predictors (human activity and AET) were important on Maui. Scaling for differences in regional human population density or economic activity (both higher on Maui) may allow more accurate and transferable quantitative predictions of introduced weed richness patterns.     

金华地区秋旱作物田杂草生态相似关系研究

对浙江金华地区秋旱作物田１７个样点的杂草优势度等级进行了七级目测调查,计测了２２种主要杂草的生态位宽度和生态位重叠值,作出了反映它们间生态学相似程度的最小生成树。结果表明,马唐Ｄｉｇｉｔａｒｉａ ｓａｎｇｕｉｎａｌｉｓ、狗尾草Ｓｔｅａｒｉａ ｖｉｒｉｄｉｓ、千金子Ｌｅｐｔｏｃｈｌｏａ ｃｈｉｎｅｎｓｉｓ、碎花莎草Ｃｙｐｅｒｕｓ ｉｒｉａ、牛筋草Ｅｌｅｕｓｉｎｅ ｉｎｄｉｃａ、石胡荽Ｃｅｎｔｉｐｅｄ     

An ecological approach to study the physical and chemical effects of rye cover crop residues on Amaranthus retroflexus, Echinochloa crus-galli and maize

Weeds can be suppressed in the field by cover crop residues, extracts of which have been demonstrated to exert chemical inhibition of crop and weed germination and early growth in bioassays. In this study, two complementary bioassays were developed with soil and mulch material originating from a long‐term maize–cover crop experiment to determine the relative physical and chemical effect of rye cover crop residues on weed and maize germination and early growth. This was compared with the effect exerted by residue material from the natural vegetation that developed in the crop stubble during the winter before maize sowing. Germination percentage and early growth of maize and two maize weeds, Amaranthus retroflexus and Echinochloa crus‐galli, were assessed in a seed incubator in tilled (green manured) and nontilled (surface mulched) soil, with and without N fertilisation, at various dates after cover crop destruction. Responses were compared to those of the same species in a standard soil without mulch or with an inert poplar mulch. A second bioassay was set up in a glasshouse to determine the effect of different quantities of fresh residue material and additional N fertilisation on emergence speed and percentage and on plant vigour during the first 22 days after cover crop destruction. These results were compared with no‐mulch controls and poplar mulch controls. Results of these trials were compared with weed density and biomass that developed in the maize crop sown after cover crop destruction. Soil and mulch chemical and biological properties were determined for material collected in the field at different times after cover crop destruction. Chemical properties of the mulch differed only occasionally between the treatments, but variation in cover crop biomass production led to significantly different soil chemical properties. Although soil total phenolic acid content did not always correlate to weed and maize germination and early growth inhibition, soil microbial activity did. In suboptimal conditions, as is often the case in the field, plant residue material exerted both a physical and a chemical effect on maize and weed emergence and early growth. Nitrogen fertilisation and application timing can give the maize crop a competitive advantage with respect to the weeds, but the final response and the practical consequences depended largely on the weed species involved.     

Weed Diversity and Uses: a Case Study from Tea Plantations in Northern Thailand

Based on concerns that useful weed diversity in agroecosystems as well as associated traditional knowledge may be declining due to agricultural intensification and indiscriminate eradication of weeds, we studied weed diversity and use by local ethnic groups in tea plantations in northern Thailand under different agricultural intensities and landscape complexities, namely agroforestry, organic, and Good Agricultural Practice (GAP) based conventional system. In each system, we sampled five tea plantations using belt-transects of 25 m². Use data were collected through interviews with specialist informants. We recorded 214 weed species (gamma-diversity) of which 66 were useful. Agroforestry system had the highest alpha- and beta-weed diversity, suggesting that the low agricultural intensity and high landscape complexity in this system is associated with higher weed diversity in tea plantations. The common weed species were clearly different in the three systems and only a small fraction (22 species) of widespread weeds was shared among them, resulting in low weed similarity between systems. The 66 useful species of weeds were mostly for food and medicine. Uses of some weeds were shared between systems. Many of the useful weeds are invasive species, suggesting that they are prevalent and inexpensive plant resources for rural people and could represent alternative resources in the future. Because the occurrence of invasive weeds may affect the natural habitats of the native flora, integrating the exploitation of weeds into weed management strategies may reduce the weed population while sustaining agrobiodiversity and conserving associated traditional knowledge in the long term.     

河南省封丘农田杂草类型的初步研究

采用聚类分析的方法,研究阐明在气候差异不明显的冲积平原区,农田杂草类型和相应的土壤类型(相当于土属)有明显的相关性。在划分旱地农田杂草类型时,环境条件的差异比栽培作物具有更重要的指示意义。由于杂草多为广布种,采用聚类分析的方法,比单纯用优势种能够更客观地划分杂草类型。 根据实地观测,指出了封丘地区危害严重的主要杂草种类和危害时间,以及不同环境条件下杂草的分布和危害程度,这对于农业生产上杂草的防除工作有一定指导意义。     

Interaction of local and landscape features in the conservation of Hungarian arable weed diversity

Question: How do local and landscape management contribute to weed diversity in Hungarian winter cereal fields? Location: Central Hungary. Methods: Vascular plants were sampled in 18 winter cereal fields along an intensification gradient according to nitrogen fertilization, in the first cereal rows (edge) and in the interior part of the fields. Weed species were divided into groups according to their residence time in Central Europe (native species, archaeophytes, neophytes) and nitrogen preference (low to medium, LMNP, and high, HNP species). The percentage of semi‐natural habitats was calculated in the 500 m radius circle. Effects of fertilizer use, transect position and semi‐natural habitats were estimated by general linear mixed models. Results: We recorded 149 weed species. Fertilizer had a negative impact on the species richness of archaeophytes and LMNP species, and on the cover of native weeds. There was greater species richness and weed cover at the edge of the fields than in the centre. A higher percentage of seminatural habitats around the arable fields resulted in greater total species richness, especially of archaeophytes and LMNP species. We found an interaction between the percentage of semi‐natural habitats and transect position for species richness of archaeophytes and LMNP species. Conclusions: Reduced use of fertilizers and a high percentage of semi‐natural habitats would support native and archaeophyte weed diversity even in winter cereal fields, while large amounts of fertilizer may promote invasion of neophytes. However, the beneficial effect of the semi‐natural habitats and greater species pool on the arable flora may prevail only in the crop edges.     

Crop/weed gene flow:Chenopodium quinoa Willd. andC. berlandieri Moq.

Introduction of the Andean grain chenopod (Chenopodium quinoa) into North America placed this crop within the distributional range of a related wild species,C. berlandieri. This wild species, native to the North American flora, is cross-compatible withC. Quinoa. Isozyme analysis of progeny fromC. berlandieri plants growing within and at the periphery of theC. Quinoa fields, combined with fertility assessment and phenetic comparison among putative hybrids and parental types, indicates that over 30% of progeny from wild plants growing as weeds withC. quinoa in 1987 were crop/weed hybrids. This high incidence of interspecific gene flow from crop to weed appears to be the result of asymmetric pollen flow to free-living plants from high-density cultivated populations. The observed level of crop/weed hybridization, combined with heterosis and partial fertility of F₁ crop/weed hybrids, suggests that repeated annual cycles ofC. quinoa cultivation within the North American range ofC. berlandieri could produce introgressive change among sympatric wild populations. In terms of risk assessment for biotechnology, these results suggest that the breeding system may not provide an accurate indication of the potential for genetic interaction among predominately self-pollinating grain crops and their free-living relatives.     

Weed community responses in a corn-soybean intercrop

Abstract. Weed community responses were examined in a corn-soybean intercrop in southwestern Michigan, USA, with and without nitrogen fertilization. Weed suppression was observed when intercropping was additive, but the depression in above-ground weed biomass was not due to a synergistic effect of crop diversity, but rather to the effects of crop density and identity. Intercropping did not have a greater suppressive effect than monocrops on weed biomass in a replacement intercrop when crop densities were similar to those in monocultures. Similarly, intercropping per se did not alter the diversity, species richness or composition of the weed community. The presence and density of corn was more important in affecting the weed community than was the density of soybeans or than intercropping itself. Fertilization and the interaction between fertilization and crop type had few effects on the weed community.