峨眉尖舌苣苔（苦苣苔科）花部形态发生及其系统学意义

观察了峨眉尖舌苣苔（Ｒｈｙｎｃｈｏｇｌｏｓｓｕｍｏｍｅｉｅｓｅ）花序及花的形态过程，并同近缘类群进行比较和分析，结论如下：１）峨眉大苣台花发生组织通常的总状花序分生组织已发生开变和位偏从而导致不平衡分化，形成花偏向于腹部一侧的变了形的总状花序，２）花冠和能育雄蕊的两侧性可能和退化雄蕊相关。３）该种雄蕊早期的发生和发育过程仍保持着祖先群群中轴肥座的系统发生式样，并在随后的发育过程中转变成侧膜胎座，４

FLORAL MORPHOGENESIS OF RHYNCHOGLOSSUM OMEIENSE (GESNERIACEAE) AND ITS PHYLOGENETIC IMPLICATION

The morphogenesis of the inflorescences and flowers ofRhynchoglossum omeiense has been observed and compared with its allied groups. Several remarkable features and evolutionary trends have been revealed as the followings: 1) The inflorescence meristem of R. omeiense is different from the normal racemose meristem in its fusiform and ellipsoid apices having been shifted to the ventral side of the axis instead of globose apices on the top of axis in the normal raceme, which has developed a unilateral raceme with only two ventral rows of bracts bearing flowers. 2) The bilabiate corolla and didynamous stamens in this species is probably related to the state of suppression of the staminode. 3) The initiation and early development of its gynoecium has maintained the nature of the carpellary closure which may have formed the axile placentation in its ancestral group. The bilocular ovary at the early developmental stage gradually transforms into unilocular one in the following developmental process. 4) As a relict species of the Klugioid plants, it is close to the genus Whytockia. However, it is almost impossible that R. omeiense has been evolved from Whytockia, but that Klugia and Whytockia may have come from a common ancestor and developed in different ways.