异株百里香花蜜腺的发育解剖学研究

异株百里香（Thymus marschallianus Willd）花蜜腺分布于子房基部的花托上，结构蜜腺盘状，成熟时膨大，环绕在花托外。蜜腺组织由分泌表皮、产蜜组织和维管束三部分组成；组织化学染色显示淀粉粒的积累是在蜜腺细胞发育的最初和最后，因此将其归为非淀粉型蜜腺。在发育的过程中细胞液泡化动态明显，而淀粉粒和多糖均不具有明显的消长变化；蜜汁是由韧皮部运转到泌蜜组织中的，再由表皮细胞的角质层渗到细胞外。

Developmental Anatomy of Floral Nectary in Thymus marschallianus Willd

The floral nectary of Thymus marschallianus Willd is located on the base of ovary receptacle, disk-like, which inflated after mature and surrounded around the receptacle. The top of the floral nectary is divided into 4 same size slices. The nectary is composed of epidermis, nectariferous tissue and vascular bundles. Histochemistry staining showed that the starch grains were accumulated in the beginning and the later stages of nectary development, so the nectary belonged to non-starch nectary. During the development of nectary, the morphological changes of vacuoles are obvious, but the changes of starch grains and polysaccharide are not obvious. It is suggested that the nectar is transported from phloem to nectariferous tissue and then excreted by permeation through the thin cuticle of epidermis.