麦冬花药绒毡层和乌氏体的细微结构

麦冬(Ophiopogon japonicus)的绒毡层发育为分泌型。在小孢子母细胞时期,绒毡层细胞达到了发育的高峰。此时,绒毡层细胞中细胞器非常丰富,具大量线粒体、高尔基体和质体,尤以肉质网含量最多;原乌氏体出现较早,在小孢子母细胞时期绒毡层细胞中就已出现;四分体时期,大量原乌氏体被排入内切向面的质膜和纤维素壁之间;到了小孢子早期,绒毡层细胞失去细胞壁,原乌氏体分布在质膜的凹陷处,孢粉素物质在其上沉积,发育为乌氏体,乌氏体有单个和复合两种类型;当花粉成熟时,绒毡层细胞完全解体。

FINE STRUCTURE OF TAPETAL CELLS AND UBISCH BODIES IN THE ANTHER OF OPHIOPOGON JAPONICUS

The development of the tapetum in Opkiopogon japonicus is of secretory type.Tapetum develops at their peak during the microspore mother cell stage.There are abundant organelles, consisting of a lot of mitochondria,dictyosom es and plastids,especially endoplasmic reticu- lum.Pro-Ubisch bodies e.merge as early as at the stage of microspore mother cell.At tetrad sta- ge,a large number of pro-Ubisch bodies accumulate between inner tangential face of the plas- malemma and the cell wall.At the early microspore stage,pro-Ubisch bodies are distributed in the small embayments of the plasmalemma.As the sporopollenin begins to deposit on them,pro- Ubisch bodies develop into Ubisch bodies which consist of two types:single and aggregated. Tapetal cells degenerate completely when pollen grains reach maturity.